taxonID	type	description	language	source
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	description	(Figs. 2 – 5)	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	materials_examined	Material examined. Holotype. CBM-ZC 17868, ovigerous female (cl 4.6 mm), Japan, Honshu, Wakasa Bay, off Ine, 35 ° 45 ’ 05 ” N 135 ° 19 ’ 44 ” E, 96 m, mud, SM grab, coll. Y. Henmi, 17 June 2022, DNA voucher (LC 832870). Paratype. CBM-ZC 15617, ovigerous female (cl 4.1 mm), Japan, Honshu, Sagami Bay, W of Jogashima, TRV Shin’yo-maru, 2015 research cruise, station 1 (T 3), 35 ° 08 ’ 13 ” N, 139 ° 34 ’ 26 ” E, 92 m, sand, dredge, coll. H. Kohtsuka, 31 October 2015, DNA voucher (LC 832871).	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	description	Description. Holotype. Body moderately robust. Carapace (Figs. 2, 3 A, B) glabrous, not setose. Rostrum about half as long as carapace, slightly flattened dorsoventrally, directed forward, reaching distal end of second article of antennular peduncle, narrowly triangular with acute apex in dorsal view; rostral dorsal carina discernible. Frontal margin devoid of supracorneal spines; extracorneal spines acuminate, not reaching distal corneal margins; infracorneal spines small, not reaching extracorneal spines. Anterolateral margin nearly straight. Pterygostomial margin broadly rounded, not protruding anteriorly. Cardiac notch deep. Pleura of first to fourth pleomeres (Fig. 2) rounded, unarmed, while that fifth pleomere (Fig. 2) acutely pointed at posteroventral angle. Sixth pleomere (Figs. 2, 3 E) with articulated triangular flap posteroventrally; posterolateral process pointed but not acuminate. Telson (Fig. 3 C, D) widest at proximal 0.4 of length, narrowing posteriorly, with 2 pairs of dorsolateral spiniform setae, anterior pair near telson mid-length, posterior pair at 0.7 telson length; posterior margin medially slightly convex, with 1 pair of tiny posterolateral spiniform setae and long mesial plumose setae, numbering about 16. Eyes (Fig. 3 A, B) largely exposed in dorsal and lateral views; cornea well pigmented. Antennular peduncle (Fig. 3 A, B) reaching distal margin of antennal scaphocerite. Basal article subequal in length to distal 2 articles combined, second article longer than third article; stylocerite acute, reaching midlength of second article; ventromesial carina with well-developed, acute spine. Lateral flagellum biramous, fused proximal portion composed of 7 segments, aesthetasc-bearing shorter ramus composed of 5 segments. Antennal peduncle (Fig. 3 A, B) with basicerite bearing small ventrolateral distal spine. Scaphocerite oval; lateral margin straight; distal lamella produced, rounded, slightly overreaching distolateral spine. Carpocerite stout, falling slightly short of distal margin of scaphocerite. Mouthparts without specific features, typical for genus. Third maxilliped (Fig. 4 A) with endopod slender, reaching distal margin of scaphocerite by tip of ultimate article. Ultimate article 1.6 times as long as penultimate article, tapering to subacute tip, latter bearing single subterminal seta, without spiniform setae. Antepenultimate article slightly sinuous, widened proximally. Coxal plate distal margin strongly produced into triangular process. Exopod not reaching distal end of antepenultimate article of endopod. First pereopods (chelipeds) (Fig. 4 B – E) subequal in length and similar in shape, elongate, slender, exceeding antennal carpocerite by about half-length of merus; surface of all articles smooth, devoid of teeth, spines or other conspicuous armature or ornamentation. Chela 5.2 times as long as wide; palm 3.2 times longer than wide, subcylindrical; dactylus slender, 0.7 of palm length, occlusal margin entire; pollex gently deflexed, occlusal margin also entire; fingers not gaping. Carpus elongate, gradually widening distally, 1.6 times chela length, 10.8 times longer than distal width. Merus subequal in length to carpus, faintly recurved, 11.7 times longer than central width. Ischium 0.8 times meral length, 9.6 times longer than distal width. Basis small, with rudimentary, bud-like exopod. Coxa robust. Second pereopod (Fig. 4 F) moderately slender, overreaching distal margin of scaphocerite by half-length of carpus. Ischium slightly shorter than merus. Carpus composed of 5 segments, first segment longer than four others combined; ratio of carpal segments (from proximal to distal) approximately: 3.4: 0.4: 0.4: 0.5: 1.0. Chela simple, longer than distal carpal segment, fingers slightly longer than palm. Third to fifth pereopods slender, generally similar. Third pereopod (Fig. 4 G, H) overreaching distal margin of scaphocerite by full length of propodus; ischium unarmed at distodorsal end, with 2 spiniform setae on ventrolateral surface; merus, carpus and propodus unarmed, merus 7.0 times as long as wide; dactylus simple, slender, scythe-shaped, half-length of propodus, 10.2 times as long as basal width, gently curved, with minute setae slightly distal to mid-length. Fourth pereopod (Fig. 4 I, J) overreaching distal margin of scaphocerite by half-length of propodus; ischium 2 ventrolateral spiniform setae; dactylus 0.6 times as long as propodus. Fifth pereopod (Fig. 4 K, L) overreaching distal margin of scaphocerite by length of dactylus; ischium unarmed; propodus with grooming apparatus consisting of 5 transverse rows of microserrulate setae in distal 0.3; dactylus 0.5 times as long as propodus. Gill formula typical for genus: pleurobranchs on to eighth thoracomeres (above bases of first to fifth pereopods); podobranch and arthrobranch absent; lobe-like epipods on first and second maxillipeds, strap-like epipods on coxae of third maxilliped and first to third pereopods; setobranchs, each consisting of single seta, present on coxae of first to fourth pereopods; first and second maxillipeds each with well-developed, flagellum-like exopod. Second to fifth pleopods each with appendix interna on endopod. Uropod (Fig. 3 E, F) with protopod ending in 2 lobes, mesial lobe acuminate, lateral lobe terminating in 2 minute teeth. Exopod with lateral margin nearly straight, terminating in small sharp tooth; distolateral spiniform seta distinctly longer than distolateral tooth; diaeresis straight, unarmed. Endopod subovate, subequal in length to exopod. Paratype. Generally similar to holotype. Rostrum reaching mid-length of second article of antennular peduncle. Right first pereopod relatively shorter compared to that of holotype. Chela 5.2 times as long as wide; palm 3.3 times longer than wide, subcylindrical; dactylus 0.7 of palm length. Carpus 1.3 times chela length, 9.1 times longer than distal width. Merus subequal to carpus length, faintly recurved, 10.7 times longer than central width. Ischium 0.8 times meral length, parallel-sided, 8.6 times longer than distal width. Third pereopod with dactylus 0.55 times as long as propodus. Colouration in life. Body generally red; carapace and pleon with whitish median stripe; telson and uropods whitish posteriorly; cornea grey; antennular peduncle and antenna translucent with tinge of red, flagella translucent; third maxilliped and pereopods generally translucent, merus and ischium of first pereopod (cheliped) with tinge of red; eggs dark red (Fig. 5).	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	distribution	Distribution. Presently known from Japan: Wakasa Bay (Sea of Japan) and Sagami Bay (Pacific coast), at depths of 92 – 96 m.	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	etymology	Etymology. From the Latin acus (= needle) and dactylus (= finger), referring to the slender dactyli of the third to fifth pereopods of the new species.	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFCCFFCE0FC0FEE452CFFC03.taxon	discussion	Remarks. Among the 43 named species of Athanas (De Grave & Fransen 2011; Ďuriš & Anker 2014; Ahyong 2015; Marin 2017; Anker & Ďuriš, 2022; Anker 2023; Komai & Henmi 2023), A. acudactylus sp. nov. appears closest to A. hongkongensis Bruce, 1990, in the combination of the following features: (1) frontal margin of carapace with extracorneal and infracorneal teeth but without supracorneal teeth; (2) chelipeds elongate, including in females, more or less symmetrical in shape and subequal in size, with slender ischium and non-inflated merus and markedly elongate carpus, without conspicuous armature on every article; and (3) third to fifth pereopods with simple, slender, scythe-shaped dactyli, attaining half or more than half-length of propodi. Athanas hongkongensis is known only by the type specimens from Hong Kong, South China Sea (Bruce 1990; Sha et al. 2019). A comparison with the original description of A. hongkongensis by Bruce (1990) revealed the following differences of possible diagnostic significance: (1) the posterolateral angle of the telson bears only one pair of spiniform setae in A. acudactylus sp. nov., whereas there are two pairs of spiniform setae in A. hongkongensis, of which the mesial pair is much longer (cf. Fig. 3 C, D; Bruce 1990: fig. 7 I); (2) the antennular peduncle reaches the distal margin of the scaphocerite in A. acudactylus sp. nov., rather than falling distinctly short of it in A. hongkongensis (Fig. 3 B; Bruce 1990: fig. 7 A, B); (3) the dactylus of the third to fifth pereopods is proportionally slightly shorter in A. acudactylus sp. nov. than in A. hongkongensis (0.5 – 0.6 times as long as propodus versus about 0.7 times as long); (4) the flexor margins of the propodi of the third and fourth pereopods are devoid of minute spiniform setae in A. acudactylus sp. nov., whereas some minute spiniform setae are present in A. hongkongensis (cf. Fig. 4 G, I; Bruce 1990: fig. 8 G, I); (5) the ischia of the third and fourth pereopods bear two spiniform setae in A. acudactylus, instead of one spiniform seta in A. hongkongensis (cf. Fig. 4 G, I; Bruce 1990: fig. 8 G, I); and (6) the lateral distal lobe of the uropodal protopod is ending in two acute teeth in A. acudactylus sp. nov., while being simple in A. hongkongensis (cf. Fig. 3 E; Bruce 1990: fig. 7 F). No genetic data are presently available for A. hongkongensis. Bruce (1990) compared A. hongkongensis with A. polymorphus Kemp, 1915, A. gracilipes Banner & Banner, 1978 and A. tenuipes De Man, 1910. Among them, A. polymorphus is immediately distinguished from the aforementioned congeners by the presence of a pterygostomial tooth on the carapace (Kemp 1915: fig. 32 a; Anker 2003: fig. 9 a). The female chelipeds and the colour pattern of A. polymorphus also differs markedly from that of the new species (cf. Fig. 5 versus Anker 2003: figs. 12, 20 e, f). Athanas gracilipes was described on the basis of two fragmentary specimens, both without chelipeds (Banner & Banner 1978; Anker & Ahyong 2007). Athanas acudactylus sp. nov. differs from A. gracilipes in the longer rostrum (in the latter species, the rostrum fails to reach the distal margin of the basal article of the antennular peduncle; the distolateral tooth of the antennal scaphocerite distinctly exceeded by the anteriorly produced lamella; and in the absence of a mesial pair of spiniform setae on the posterolateral angles of the telson (cf. Banner & Banner 1978: fig. 3 a – c). Athanas tenuipes is known with certainty only from the possibly juvenile holotype, which, like A. gracilipes, lacks both chelipeds, although Anker & Ahyong (2007) mentioned a specimen from southern Vietnam possibly representing A. tenuipes. This species can be diagnosed by the narrow telson with relatively large dorsolateral spiniform setae and a truncate posterior margin, the latter also with a shallow median notch and narrowly spaced posterior spiniform setae (De Man 1911: pl. III, fig. 8 b). In the new species, the telson has small spiniform setae and a broadly rounded posterior margin, with only two spiniform setae, one at each angle (Fig. 3 C). The sympatrically occurring A. exilis (see below) is readily distinguished from A. acudactylus sp. nov. by the shorter rostrum, which does not reach the distal margin of the basal article of the antennular peduncle versus overreaching it in the new species (cf. Fig. 3 A, B; Komai & Henmi 2023: fig. 2 B, C); the more robust female chelipeds, with row of small tubercles on the ventral margin of the merus and carpus, versus female chelipeds being slenderer and without conspicuous armature in the new species (cf. Fig. 4 B, C; Komai & Henmi 2023: fig. 3 D, E); the presence of long spiniform setae on the dorsal margin of the cheliped ischium, which are absent in the new species (cf. Fig. 4 B, C; Komai & Henmi 2023: fig. 3 D, E); and the more strongly produced distal lamella of the antennal scaphocerite (cf. Figs. 2 G, 6 B; Komai & Henmi 2023: fig. 3 B).	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFC7FFC80FC0FC2E5235FC4B.taxon	description	(Figs. 6 – 10)	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFC7FFC80FC0FC2E5235FC4B.taxon	materials_examined	Material examined. CBM-ZC 17869, 1 male (cl 5.4 mm), Japan, Honshu, Wakasa Bay, off Ine, 2018, dredge; CBM-ZC 17870, 1 male (cl 6.4 mm,), Japan, Honshu, Wakasa Bay, off Ine, 35 ° 46 ’ 03 ” N 135 ° 20 ’ 21 ” E, 95 m, dredge, coll. Y. Henmi, 13 July 2021, DNA voucher (LC 832869); CBM-ZC 17871, 2 males (cl 5.7 mm, one damaged), Japan, Honshu, Wakasa Bay, off Ine, 35 ° 44 ’ 39 ” N 135 ° 20 ’ 00 ” E, 95 m, dredge, coll. Y. Henmi, 12 May 2022; CBM-ZC 17872, 1 ovigerous female (cl 4.2 mm), Japan, Honshu, Wakasa Bay, off Ine, 35 ° 45 ’ 10 ” N 135 ° 19 ’ 55 ” E, 92 m, dredge, coll. Y. Henmi, 21 June 2023; CBM-ZC 17873, 1 male (cl 5.7 mm), Japan, Honshu, Wakasa Bay, off Ine, 35 ° 45 ’ 19 ” N 135 ° 19 ’ 60 ” E, 92 m, dredge, coll. Y. Henmi, 19 July 2023. Supplemental description. New material generally similar to female holotype except for antennae, male first pereopods (chelipeds) and male second pleopods. Rostrum (Fig. 6 A – C) 0.2 – 0.25 times as long as carapace, falling short of distal margin of basal article of antennular peduncle. Antennular peduncle (Fig. 6 A – C) proportionally longer in males than in females. First pereopods (Figs. 7 – 9) carried flexed back at mero-carpal articulation, subequal or slightly unequal, somewhat dissimilar in shape of dactylus and carpus; when flexed, fingertips reaching to base of ischium. Chela strongly compressed and expanded, leaf-shaped, 1.5 – 1.7 times as long as wide. Dactylus 0.2 – 0.3 length of palm, obliquely articulated against strongly deflexed pollex, tapering to acute or subacute tip, crossing with pollex, curved, more strongly curved in major chela; extensor margin with 1 – 2 small subdistal tubercles and sometimes blunt protuberance proximally; occlusal margin with subrectangular or subtrapezoidal proximal tooth and excavation, better developed in major chela. Pollex short, triangular, terminating in subacute tip, with subrectangular or subtrapezoidal tooth medially on occlusal margin, better developed in major chela, fitting into excavation on occlusal margin of dactylus; flexor margin with row of small tubercles. Palm slightly thickened along midline, deeply constricted proximally; extensor margin convex, proximal margin expanded to reach distal margin of carpus; flexor margin more convex than dorsal margin, bearing row of low irregular, usually obtuse teeth or tubercles; proximal margin produced to reach distal margin of carpus; mesial surface shallowly concave on ventral part. Carpus somewhat dissimilar between sides, about half-length of chela, 2.9 – 3.2 (major chela) or 3.1 – 4.6 times (minor chela) as long as wide at its widest point at distal one-third, tapering proximally, with shallow constriction subdistally; extensor and flexor margins each with row of small tubercles, mesial face sometimes with additional row of small tubercles adjacent to dorsal margin; lateral face smooth. Merus 1.6 – 1.7 times length of carpus, 4.4 times as long as wide; widest point proximal to mid-length, slightly broader than ischium; dorsal margin faintly sinuous, smooth or with row of minute tubercles; lateral surface almost flat; mesial face bisected by blunt median ridge bearing row of minute tubercles, dorsal half smooth or with scattered minute tubercles, ventral half shallowly excavated; lower margin faintly sinuous, with row of minute tubercles at least in distal half. Ischium 0.8 times as long as merus; dorsal margin slightly convex, armed with 8 – 10 long spiniform setae in proximal half; mesial face with longitudinal row of 3 or 4 long spiniform setae in proximal one-third and longitudinal row of minute tubercles along midline, ventral half slightly concave; lateral face nearly flat or faintly sulcate; ventral margin nearly straight of faintly sinuous, with row of minute tubercles over entire length and sometimes with 1 long spiniform seta proximally. Basis with minute, bud-like exopod. Coxa large, rounded, similar from left to right. First pleopod endopod (Fig. 6 D) about half length and width of exopod, gradually tapering to pointed apex, slightly curved mesially. Second pleopod with appendix masculina rod-like, 1.3 times longer than appendix interna (Fig. 6 E), with about 10 stiff setae terminally or subterminally.	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFC7FFC80FC0FC2E5235FC4B.taxon	description	Variation in cheliped. The first pereopods (chelipeds) are subequal to slightly unequal in length with dissimilar fingers and carpi. In the major chela, the dactylus is more strongly curved; the occlusal teeth on the dactylus and pollex are stronger; and the carpus is stouter. In three specimens (CBM-ZC 17869, 17871 and 17873), the right cheliped is major, while the left cheliped is major in one specimen (CBM-ZC 17870); thus, the handedness does not seem to be fixed on one side. In the ovigerous female (CBM-ZC 17872), the right cheliped is normally developed, while the left cheliped is much shorter, slenderer and with a soft integument, indicating a regeneration process. Colouration in life. Generally similar to holotype (Komai & Henmi 2023: 68, fig. 1 A, B). First pereopod with chela generally translucent, thickened median part whitish; fingers yellowish white; carpus with tinge of pale red; merus and ischium with scattered pale red spots (Fig. 10).	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFC7FFC80FC0FC2E5235FC4B.taxon	distribution	Distribution. Presently known only from Wakasa Bay, Sea of Japan, at depths of 90 – 95 m.	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
03AD037FFFC7FFC80FC0FC2E5235FC4B.taxon	discussion	Remarks. Athanas exilis was originally described based on a single ovigerous specimen from Wakasa Bay collected at a depth of 90 m. In the original description, Komai & Henmi (2023) compared A. exilis with A. sydneyensis Anker & Ahyong, 2007 from eastern Australia and A. amazone Holthuis, 1951 from the eastern Atlantic. With the present male specimens, it is now clear that A. exilis is closest to A. phyllocheles Banner & Banner, 1983, presently known only from the western Indian Ocean off Réunion, at depths of 345 – 450 m, with more distant affinities to A. amazone. All three species have unusually expanded, leaf-shaped chelae on the first pair of pereopods in males (Fig. 7; Banner & Banner 1983: fig. 13 d – f; Anker & Ahyong 2007: fig. 5). The morphological differences between A. exilis and A. phyllocheles are as follows: (1) the rostrum falls short of the distal margin of the basal article of the antennular peduncle in A. exilis (Fig. 6 A – C; Komai & Henmi 2023: fig. 2 B, C), rather than reaching it in A. phyllocheles (Banner & Banner 1983: fig. 13 a, b); (2) the rostrum is devoid of a dorsal carina in A. exilis (Fig. 6 C; Komai & Henmi 2023: fig. 2 C), which is present in A. phyllocheles (Banner & Banner 1983: fig. 13 a); (3) the extracorneal tooth falls far short of the distal corneal margin in A. exilis (Fig. 6 B, C; Komai & Henmi 2023: fig. 2 B, C), whilst reaching to the distal corneal margin in A. phyllocheles (Banner & Banner 1983: fig. 13 a, b); (4) the male chelae are much more expanded in A. exilis than in A. phyllocheles (cf. Figs. 7 A, C, 8 A; Banner & Banner 1983: fig. 13 d – f); (5) the dorsal and ventral proximal margins of the male chelae are expanded into rounded lobes in A. exilis (Figs. 7 A, C, 8 A), but not in A. phyllocheles (Banner & Banner 1983: fig. 13 d – f); (6) the meri and ischia of the male chelipeds are ridged along the midline and bear a row of minute tubercles on their mesial face in A. exilis (Fig. 7 A, C), whereas in A. phyllocheles, the mesial faces of the meri and ischium are smooth (Banner & Banner 1983: fig. 13 d – f); and finally, (7) the dactylus of the third pereopod is proportionally longer in A. exilis than in A. phyllocheles (0.6 times as long as the propodus versus 0.4 times as long).	en	Komai, Tomoyuki, Henmi, Yumi (2025): A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from Japan, and additional record of Athanas exilis Komai & Henmi, 2023, clarifying male characteristics. Zootaxa 5583 (2): 353-370, DOI: 10.11646/zootaxa.5583.2.7, URL: https://doi.org/10.11646/zootaxa.5583.2.7
