identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03AB87D9FFD4A412FF2CF8B9F7873C14.text	03AB87D9FFD4A412FF2CF8B9F7873C14.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyssemus inscitus (Walker 1858)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Rhyssemus inscitus (Walker, 1858) (Fig. 2) </p>
            <p>ḝṣạỄAE래ǭ뎅ḏ (ṉÿ)</p>
            <p> Psammodius inscitus Walker, 1858: 207 . Type locality. Ceylon. </p>
            <p> Rhyssemus tarsalis Waterhouse, 1876: 115 . </p>
            <p> Rhyssemus malasiacus Lansberge, 1886: 133 .  Rhyssemus insignicollis Lea, 1923: 12 . </p>
            <p> Rhyssemus australis Petrovitz, 1963: 40 . </p>
            <p> Rhyssemus papuanus Petrovitz, 1965: 169 . </p>
            <p> Rhyssemus mussardi Petrovitz, 1975: 618 . </p>
            <p> Rhyssemus philippineus Masumoto, 1980: 21 . </p>
            <p>Diagnosis. Body size. ♂ (n = 1): BL, 2.95; HL, 0.35; HW, 0.75; PL, 0.78; PW, 1.15; EL, 1.74; EW, 1.3. ♀ (n = 1): BL, 3.02; HL, 0.28; HW, 0.76; PL, 0.81; PW, 1.17; EL, 1.89; EW, 1.22. Habitus. body subparallel, slightly dilated behind the middle of elytra, dark brown. Head. convex, granulate; clypeus obtusely dentate on each side of anterior emargination; genae obtusely rounded, protruding more than eyes with few fine setae; head granules moderately coarse, fairly uniform in size and distribution, a pair of vestigial oblique ridges presented posteriorly. Pronotum. convex, nearly 1.4 times as wide as long; six transversal ridges moderately convex, the 1 st ridge distinct, the 2 nd to 4 th ones joining together to form lateral callus with shallow granules; furrows shallowly impressed, minutely granulate or wrinkled, posterior longitudinal furrow interrupt 4 th to 5 th ridges; lateral-posterior margins of pronotum crenate, with slightly dilated but apically blunt thick setae. Elytra. convex, subparallel, slightly dilated behind the middle; humeral tooth distinct; stria narrow, shallowly impressed with longitudinal punctures; interstria wide, with two rows of indistinct pearl-like granules, randomly fused between two neighboring granules. Pygidium. two apical pygidial setae present. Legs. metafemora widest at middle, margined posteriorly, setigerous punctate anteriorly and near apically on ventral surface; metatibial upper spur as long as basal metatarsus; basal metatarsus slightly shorter than 2 - 4 th joints together. Aedeagus as in Fig. 2.</p>
            <p> Specimens examined.   1♂ (CSL-21-0386): South Korea, Chungcheongnam-do, Taean-gun, Wonbuk-myeon,  Guryepo Campsite , 19.VIII.2021, legs. C. Lim and W.G. Kim [NIBR]  ;   1♂ (CSL-21-0264): South Korea, Chungcheongnam-do, Taean-gun, Wonbuk-myeon,  Guryepo Campsite , 19.VIII.2021, legs. C. Lim and W.G. Lee [KUEM]  ;   1♀: South Korea, Jeju Island, Jeju-si,  Jocheoneup ,  Hamdeok Beach , 3.VIII.2021, leg. C. Lim [KUEM]. Other specimens examined  .   1♂ 2♀ (CSL-21-0317, 0318): Thailand, Udon Thani Province,  Nong Saeng District , Than Ngam Waterfall, 13. VI  .2018. leg. C.J. Uy [KU];   1♀ (CSL-21-0315): Cambodia, Kampong Spueu Province, Dechou Akphivoadth, 3.VIII.2012, leg. Y.J. Bae [KU]. 1ex: Australia,  Katherine , 6 - 10.II.1968. leg. E. Matthews [USNM]  . </p>
            <p>Distribution. South Korea (new country record), Australia, Egypt, India, Indonesia, Nepal, Pakistan, Papua New Guinea, The Philippines, Taiwan, Oriental Region, Afrotropical Region.</p>
            <p> Habitat. All Korean specimens were collected from sand or loamy sand soils near the roots of herbaceous plants (5 - 15 cm in depth) within the costal sand dunes. Trichorhyssemus asperlus,  Psammodius hangangensis , and  Leiopsammodius japonicus were collected together with  R. inscitus . </p>
            <p> DNA barcode. Partial COI sequences of two specimens from South Korea and three specimens from the Oriental regions were obtained (accession numbers: PP431264 - PP431268). The estimates of genetic divergence (p -distance) between the Korean and Oriental group was 1.5% (standard error = 0.4). The estimate of genetic divergence between  R. inscitus and  R. germanus , the only available  Rhyssemus species in GenBank and BOLD Systems (accession numbers: KM443814, KM448164, KM448190, KM450328, KM450806, HQ954340, and HQ954341), was 11.2% (standard error = 1.2). </p>
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	https://treatment.plazi.org/id/03AB87D9FFD4A412FF2CF8B9F7873C14	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Bae, Changseob Lim and Yeon Jae	Bae, Changseob Lim and Yeon Jae (2024): New distribution record of genus Rhyssemus Mulsant (Coleoptera, Scarabaeidae, Aphodiinae) from South Korea. Journal of Species Research 13 (3): 306-309, DOI: 10.12651/JSR.2024.13.3.306
