identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A987B9D177B71FFF0045C4E8D90CA3.text	03A987B9D177B71FFF0045C4E8D90CA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrmecina andalas Satria & Yamane 2019	<div><p>Myrmecina andalas sp.nov.</p><p>(Figs 1, 2, 3, 4)</p><p>Holotype. Worker, Indonesia, Sumatra, Aceh, Leuser Ecosystem, Putri Betung, Gunung Kemiri, N 3°49'521'', E 97°31'198'', alt. ca. 950−1200 m, 19.IX.2012, R. Satria leg., individual code: SEMU- T20180326A (MZB).</p><p>Paratypes. One worker (SEMUT20180731A), same data as for holotype (RSC); 3 workers (SKYUSI-FOR001–003), Leuser Ecosystem, Gunung Kemiri, alt. 1200 m, good forest, 20.IX.2012, Yamane &amp; Syaukani leg. (SKYC); 1 ergatoid queen (SEMUT20180731B), same data as for holotype (RSC) .</p><p>Worker measurements and indices. Holotype and paratypes (n = 3): HL 0.90 mm, HW 1.00− 1.02 mm, MDL 0.50−0.52 mm, EL 0.24 mm, SL 0.94 mm, WL 1.19−1.20 mm, PNW 0.66−0.68 mm, PSL 0.12−0.13 mm, PTH 0.27−0.28 mm, PTL 0.26 mm, CI 111−113, MDI 55−57, SI 92−94, PTHI 103−107.</p><p>Worker description. Relatively large (HL 0.90 mm, HW 1.00− 1.02 mm, WL 1.19−1.20 mm). Head in frontal view rectangular, slightly shorter than broad, with posterior margin almost straight (Fig. 3A); lateral sides of head shallowly convex. Clypeus consisting of flat dorsal disc and steep anterior face, laterally serrate; its anterior margin almost straight, with small median tooth and pair of lateral denticles. Labrum with pair of apical processes. Eye relatively large (EL 0.24 mm; 72– 73 ommatidia), distinctly breaking lateral margin of head, in profile occupying almost 1/4 length of head. Mandible triangular; masticatory margin with large apical and smaller preapical teeth, followed by 6 denticles. Antenna 12-segmented, with 3-segmented club; apical segment slightly longer than segments 10 and 11 combined; scape relatively long, distinctly surpassing posterolateral corner of head. Mesosoma in lateral view short and stout, with dorsal outline evenly convex; anterior ventrolateral portion of pronotum forming distinct process; mesonotum completely fused with pronotum; lower portion of mesopleuron broadly concave anteriorly (red arrow in Fig. 3B); mesopleuron not differentiated from metapleuron. Propodeum in dorsal view weakly demarcated anteriorly, but metanotal groove indistinct; anterior pair of denticles present on propodeal dorsum, very small and short (difficult to distinguish them from longitudinal rugae on propodeum) (red arrow in Fig. 2A); propodeal spine triangular, much shorter than broad and pointing dorsally; propodeal declivity almost vertical with lateral carinae well developed, with posterodorsal angle about 90°; posteriormost portion of propodeum with high lateral walls (propodeal lobes) (Figs 3B, 3C, 3D). Petiole in dorsal view as broad as long, rectangular; its anterior slope in profile almost straight, dorsal face flat to weakly concave, and ventral outline straight with a small anterior denticle; postpetiole in dorsal view much broader than long, broader than petiole; sternopostpetiolar process absent. Gaster in dorsal view slightly elongate circular; anterior margin of first gastral tergite straight, with anterolateral corner very weakly angled. Mid- and hind tibiae broad and long, 3.5 times as long as broad.</p><p>Head dorsally with coarse longitudinal rugae; those on lateral part of head running posterolaterally toward posterolateral corner of head; interspaces between lateral rugae superficially sculptured and shiny; flat dorsal disc of clypeus smooth and shiny, with irregular punctures around anterolateral corner; ventrolateral area (temple + gena) of head smooth and shiny, with 1–2 longitudinal rugae (see Fig. 1A). Mesosoma, petiole and postpetiole with coarse irregular longitudinal rugae; declivity and posteriormost part of propodeun smooth and shiny. Gaster smooth and shiny.</p><p>Body covered with abundant long erect setae; clypeus with several long setae and a pair of long setae arising from median tooth of its anterior margin; scape with many suberect to erect setae that are longer than scape width (some of them as long as eye); all legs with numerous suberect to erect setae.</p><p>For color pattern see Figs 1−3; mesosoma, petiole and postpetiole dark reddish brown; head and gaster blackish brown, with apex of gaster orange. Mandible, clypeus, antenna and all legs reddish brown to brown, with antennal flagellum, tibiae and tarsi paler.</p><p>Ergatoid queen measurements and indices. Paratype (n = 1): HL 0.92 mm, HW 1.09 mm, MDL 0.54 mm, EL 0.27 mm, OED 0.39 mm, SL 0.97 mm, WL 1.30 mm, PNW 0.72 mm, PSL 0.16 mm, PTH 0.29 mm, PTL 0.26 mm, CI 118, MDI 58, SI 88, PTHI 111.</p><p>Ergatoid queen description. Very similar to worker including size, but differentiated from worker as follows: size of eye and number of ommatidia larger than in worker (EL 0.27 mm; 85 ommatidia); three ocelli present (Fig. 4A); dorsal outline of mesosoma in lateral view more convex than in worker; small hole present at forewing position on mesosoma (blue arrow in Fig. 4B) near small round pale area (red arrow in Fig. 4B).</p><p>Comparative remarks. Myrmecina andalas sp. nov. is morphologically most similar to M. magnificens Wong et Guénard, 2016 among the species known from Sundaland. However, it is distinguishable from the latter by the following characteristics of worker: ventrolateral area of head (temple + gena) smooth and shiny; anterior pair of denticles on propodeal dorsum very small and short (distinct in M. magnificens); propodeal spine triangular, much shorter than broad at base and pointing dorsad; propodeal declivity vertical, with posterodorsal angle almost 90°; sternopostpetiolar process absent. Worker of M. andalas sp. nov. also resembles that of M. nipponica Wheeler, 1906 . However, it is easily separated from the latter by the much larger eye that is more than twice as long as the antennal pedicel (vs. eye small, as long as pedicel), long erect setae on scape, some of which are 1.5 times as long as scape width (vs. most erect setae very short, not longer than scape length), and the propodeal lobe dorsally roundly convex (vs. in shape of a narrow strip).</p><p>The ergatoid queen in Asian Myrmecina has also been found in M. nipponica (Ohkawara et al., 1993; Miyazaki et al., 2005; Peeters, 2012) and in undescribed species from Java (Ito, 1996). Despite the difficulty in recognizing the ergatoid queen among dead specimens, some morphological characters are used for the recognition of ergatoid queen, such as the presence of ocelli and vestiges of wing articulation on mesosoma (Fig. 4). The vestige of wing articulation was observed in the ergatoid queen of M. andalas as a small hole in place of the forewing near small round pale area on the mesosoma, and the same feature was also reported in ergatoid queens of M. nipponica which were determined by a direct observation of the ovaries and spermatheca (Ohkawara et al., 1993). The ergatoid queen has developed eyes as in the dealate queen, but smaller in body size with a smaller number of ommatidia than in the dealate queen. On the other hand, the worker has smallest eyes and number of ommatidia among the three castes, but ommatidia each being larger than those in both the dealate queen and ergatoid queen in M. nipponica (Miyazaki et al., 2015). Although in M. andalas the ergatoid queen has larger eyes and number of ommatidia than in the worker, the size of each ommatidium was similar in both castes.</p><p>Etymology. The specific epithet is a previous name of Sumatra Island.</p><p>Distribution. So far known only from Sumatra.</p><p>Bionomics. No information about the biology of this species is available, except that specimens were collected from leaf litter in secondary forests.</p></div>	https://treatment.plazi.org/id/03A987B9D177B71FFF0045C4E8D90CA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Satria, R.;Yamane, Sk.	Satria, R., Yamane, Sk. (2019): Two new species of the ant genus Myrmecina (Hymenoptera: Formicidae: Myrmicinae) from Sumatra. Zoosystematica Rossica (China) 28 (1): 183-193, DOI: 10.31610/zsr/2019.28.1.183, URL: https://doi.org/10.31610/zsr/2019.28.1.183
03A987B9D170B712FF0F40D1EF860B91.text	03A987B9D170B712FF0F40D1EF860B91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrmecina nitidiuscula Satria & Yamane 2019	<div><p>Myrmecina nitidiuscula sp. nov.</p><p>(Figs 2C, 5, 6)</p><p>Holotype. Worker, Indonesia, Sumatra, Aceh,</p><p>Leuser Ecosystem, Putri Betung, 21.IX.2012,</p><p>R. Satria Leg., individual code: SEMUT20180326B (MZB).</p><p>Paratypes. Two workers (SEMUT20180808A, SEMUT20180808B), same data as for holotype (RSC); 2 workers (SKYUSI-FOR004–005), same locality, but alt.&gt; 1100 m, good forest, 20.IX.2012, Yamane &amp; Syaukani leg. (SKYC); 1 dealate queen (SEMUT20181112), same locality, but alt. 850 m, plantation, 19.IX.2012, Yamane &amp; Syaukani (RSC) .</p><p>Worker measurements and indices. Holotype and paratypes (n = 4): HL 0.65−0.67 mm, HW 0.72−0.74 mm, MDL 0.35−0.38 mm, EL 0.15−0.16 mm, SL 0.58−0.59 mm, WL 0.87−0.93 mm, PNW 0.50−0.52 mm, PSL 0.14−0.15 mm, PTH 0.21 mm, PTL 0.18−0.22 mm, CI 110, MDI 53−56, SI 78−80, PTHI 116.</p><p>Worker description. Relatively small (HL 0.65−0.68 mm, HW 0.72−0.75 mm, WL 0.87−0.93 mm). Head in frontal view rectangular, slightly shorter than broad, with posterior margin</p><p>R. Satria &amp; Sk. Yamane. Two new species of Myrmecina from Sumatra slightly concave (Fig. 5A); lateral sides of head evenly convex. Clypeus consisting of short dorsal disc and steep anterior slope; its anterior margin almost straight, with small median tooth and a pair of weak lateral processes. Eye relatively large (EL 0.15−0.16 mm; 34−35 ommatidia), breaking outer margin of head by 1/5 length of eye in profile. Mandible triangular; masticatory margin with large apical tooth, followed by two relatively large teeth, 3–4 small denticles, diastema and low but broad basal tooth. Antenna 12-segment- ed, with 3-segmented club; apical segment slightly longer than segments 10 and 11 combined; scape relatively short, not surpassing posterolateral corner of head. Mesosoma in lateral view short and stout, with dorsal outline roundly convex; anterior ventrolateral portion of pronotum weakly angulate, without anterior process; mesonotum completely fused with pronotum; anteroventral corner of mesopleuron angulate; mesopleuron not differentiated from metapleuron. Propodeum in dorsal view weakly separated from mesonotum but without distinct metanotal groove; propodeal spine in profile triangular, apically blunt but pointing posteriad, with very broad base; dorsum with a pair of barely recognized denticles anteriorly (Fig. 2C); declivity steep, with transverse carina connecting propodeal spines; flat posteriormost portion of propodeum laterally margined by wall (part of propodeal lobe) (Figs 5B, 5C, 5D). Petiole in dorsal view as broad as long, in lateral view consisting of flat to weakly concave dorsal face and gently sloping anterior face; ventral outline of petiole straight, with longitudinal median keel that is posteriorly angulate; postpetiole in dorsal view much broader than long, shorter but broader than petiole, in profile dorsally shallowly convex, with ventral margin almost straight; sternopostpetiolar process absent. Gaster in dorsal view almost circular; first gastral tergite without distinct antero-</p><p>R. Satria &amp; Sk. Yamane. Two new species of Myrmecina from Sumatra lateral angle. Mid- and hind tibiae very broad and short, 2.5 as long as broad.</p><p>Head superficially sculptured with scattered shallow punctures but generally smooth and shiny; frontal lobe, area between eye and frontal lobe, and area close to mandible more strongly sculptured; ventrolateral surface of head (temple + gena) smooth and shiny; mandible smooth, with dorsobasal portion irregularly sculptured; antennal scape superficially sculptured and shiny. Mesosoma, petiole and postpetiole with coarse irregular longitudinal sculpture; anterior protruding portion (neck) of pronotum transversely rugose. Gaster smooth and shiny.</p><p>Body including legs covered with abundant long erect setae; clypeus with a pair of very long setae and a few short setae on disc and a pair of long setae arising around base of anterior tooth; antennal scape with suberect to erect setae, some of which are longer than scape width.</p><p>For color pattern see Fig. 5; mesosoma reddish brown; head and gaster blackish brown with apex of gaster orange. Clypeus, mandible, and antenna pale reddish brown. All legs yellowish brown.</p><p>Queen measurements and indices. Paratype (n = 1):HL 0.68 mm,HW 0.75 mm,MDL 0.37 mm, EL 0.19 mm, OED 0.28 mm, SL 0.58 mm, WL 0.97 mm, PNW 0.60 mm, PSL 0.16 mm, PTH 0.21 mm, PTL 0.20 mm, CI 110, MDI 54, SI 77, PTHI 105.</p><p>Queen description. Differing from worker only in structure of eye and in structure and surface texture of mesosoma as follows: size of eye and number of ommatidia greater than in worker (EL 0.19 mm; 67 ommatidia), but each ommatidium similar in size in both; mesosoma in dorsal view with mesoscutum large, broad and roundly convex anteriorly; notaular lines absent; parapsidal lines weakly impressed, shorter than half of mesoscutum length; metanotum medially produced anteriad, dividing mesoscutellum into two triangular sections; propodeum separated from metanotum by deep groove; mesosoma in profile forming single dome; mesonotum and metanotum higher than pronotum and propodeum; mesopleuron demarcated from pronotum and metapleuron, divided into dorsal and ventral sections by deep groove; separation of metapleuron from lateral face of propodeum incomplete; dorsum of pronotum transversely striate; mesoscutelum smooth and shiny, with posteromedian portion coarsely longitudinally rugose; ventral section of mesopleuron smooth and shiny; other parts rugose to irregularly sculptured.</p><p>Comparative remarks. Myrmecina nitidiuscula sp. nov. is easily distinguished from other named species of Myrmecina known from Sundaland by the combination of the following characteristics: rather smooth and shiny head with ventrolateral area (temple + gena) of head smooth, triangular propodeal spine pointing posteriad and with very broad base, and short and broad mid- and hind-tibiae.</p><p>Etymology. The species epithet relates to the extensively shiny dorsum of the head.</p><p>Distribution. So far known only from Sumatra.</p><p>Bionomics. This species was collected from leaf litter in the secondary forest.</p><p>Key to Sumatran species of Myrmecina based on the worker caste</p><p>1. Large size (HW ≥1.00 mm); mesosoma in lateral view with propodeal spines pointing anteriorly or dorsally (Figs 2A, 2B)...... M. andalas sp. nov.</p><p>– Small size (HW &lt;1.00 mm); mesosoma in lateral view with propodeal spines pointing posteriorly (Figs 5D).................................... 2</p><p>2. Mesosoma in profile having propodeal spine small, shorter than basal width or equal to it.......... 3</p><p>– Mesosoma in profile having propodeal spine distinct, with length exceeding basal width....... 5</p><p>3. Dorsum of head only superficially sculptured, generally smooth and shiny; frontal lobe, area between eye and frontal lobe, and area close to mandibular base strongly sculptured.............................................. M. nitidiuscula sp. nov.</p><p>– Head in frontal view entirely distinctly sculptured and mat...................................... 4</p><p>4. In profile, pro-mesonotum weakly to moderately convex. In frontal view, anterior margin of clypeus straight. Eye large; its length approximately 1/4 of HL........................... M. bandarensis</p><p>– In profile, pro-mesonotum strongly convex. In frontal view, anterior margin of clypeus slightly concave. Eye smaller; its length less than 1/4 of HL...................................... M. butteli</p><p>5. In frontal view, clypeus with distinctly concave anterior margin..................... M. undulata</p><p>– In frontal view, clypeus with straight anterior margin................................. M. nesaea</p></div>	https://treatment.plazi.org/id/03A987B9D170B712FF0F40D1EF860B91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Satria, R.;Yamane, Sk.	Satria, R., Yamane, Sk. (2019): Two new species of the ant genus Myrmecina (Hymenoptera: Formicidae: Myrmicinae) from Sumatra. Zoosystematica Rossica (China) 28 (1): 183-193, DOI: 10.31610/zsr/2019.28.1.183, URL: https://doi.org/10.31610/zsr/2019.28.1.183
