identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B06032E118CE0B5ED6F9CAFC49FCE4.text	03B06032E118CE0B5ED6F9CAFC49FCE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triclavitermes Ferreira, Carvalho & Vasconcellos 2025	<div><p>Triclavitermes Ferreira, Carvalho &amp; Vasconcellos, gen. nov.</p><p>Type-species. Triclavitermes catoleensis sp. nov.</p><p>Diagnosis. Enteric valve seating (EVS) forms a well-defined ring at the enteric valve (EV) insertion, followed by a short tubular portion of P3. It also features a mesenteric tongue with a whitish spherical mesenteric nodule in ventral view. EV with three semi-spherical sclerotized plates.</p><p>Imago. Unknown.</p><p>Worker. External morphology (Fig. 1): Monomorphic, with a pale-yellow head covered by sparsely spaced bristles of the same color. Head slightly convex in profile, post-clypeal region inflated, and fontanelle not visible. Antenna composed of 14 segments, with relative lengths 2&gt;3=4&lt;5. The pronotum, mesonotum, and metanotum are adorned with long bristles distributed along the anterior margin of each nota. The abdomen is covered, dorsal and ventrally, by small, posteriorly directed bristles, with only its ventral portion also exhibiting larger and more spaced, anteriorly oriented. The foretibia is slightly inflated and covered with both bristles and hairs. Measurements (in millimeters) were taken from a total of 25 workers (five specimens from each of the five different localities): WH: 0.64–0.71 (mean: 0.67); LfT: 0.45–0.74 (mean: 0.50); WfT: 0.12–0.17 (mean: 0.14); and LhT: 0.50–0.64 (mean: 0.59).</p><p>Mandibles (Fig. 2): Left mandible with an apical tooth larger than M1+2, M1+2 possessing a rounded tip and M3 forming a right angle. The incision between M1+2 and M3 is quite wide. The molar process is small and not hidden by the molar prominence, which is well developed. Right mandible apical tooth three times the size of M1, with its posterior edge forming an acute angle with M1. The edge between M1 and M2 forms an obtuse angle, and the molar plate is concave, lacking ridges.</p><p>Gut (Fig. 3–4): The crop is globular, located immediately posterior to the thorax, visible centrally in dorsal view. The gizzard is inconspicuous and unarmed and follows the pattern found in most soil-termites, being underdeveloped, with four first-order pulvilli, and the pulvilli surface with pectinated scales. The junction between P1 and the mesenteron shows a moderately long mixed segment. The mesenteric tongue is slightly inflated in ventral view. Malpighian tubules are filiform, with no nodules. P1 is slightly sinuous with a uniform width along its length, connecting to P3 through an enteric valve seating (EVS), forming a well-defined ring where the armature of the enteric valve (EV) is everted (Fig. 4 and 5E). P3 is elongated and dilated, connected to P4 by a narrow isthmus located on the left lateral side in dorsal view (Fig. 3B). P4 elongated and exhibiting a pronounced U-turn. The connection P4–P5 forms a slight curve in dorsal view.</p><p>Etimology. Referring to the clava, a solid sphere with spines that was part of the maces used in medieval battles, due to its resemblance to the species’ enteric valve armature.</p></div>	https://treatment.plazi.org/id/03B06032E118CE0B5ED6F9CAFC49FCE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ferreira, Renan Rodrigues;Carvalho, Antonio;Lucena, Emanuelly Félix De;Figueiredo, Rozzanna Esther Cavalcanti Reis De;Vasconcellos, Alexandre	Ferreira, Renan Rodrigues, Carvalho, Antonio, Lucena, Emanuelly Félix De, Figueiredo, Rozzanna Esther Cavalcanti Reis De, Vasconcellos, Alexandre (2025): Triclavitermes catoleensis (Termitidae: Apicotermitinae), a new soldierless termite genus and species from the Caatinga dry forest, northeastern Brazil. Zootaxa 5706 (3): 383-396, DOI: 10.11646/zootaxa.5706.3.4, URL: https://doi.org/10.11646/zootaxa.5706.3.4
03B06032E11ECE085ED6FB5EFEE9FE4C.text	03B06032E11ECE085ED6FB5EFEE9FE4C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triclavitermes catoleensis Ferreira, Carvalho & Vasconcellos 2025	<div><p>Triclavitermes catoleensis Ferreira, Carvalho &amp; Vasconcellos sp. nov.</p><p>Holotype. CTUFPB-11404 08.VI.2022. A. Vasconcellos col.</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.738613&amp;materialsCitation.latitude=-6.3394446" title="Search Plazi for locations around (long -37.738613/lat -6.3394446)">Municipality of Catolé do Rocha</a>, Monte Tabor (6º20'22"S, 37º44'19"W), Paraíba, Brazil .</p><p>Type repositor. CTUFPB</p><p>Paratypes. BRAZIL. Pernambuco: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-37.233334&amp;materialsCitation.latitude=-8.533334" title="Search Plazi for locations around (long -37.233334/lat -8.533334)">Buíque</a> (8°32'00.0"S 37°14'00.0"W): 16.IV.2009, A. Vasconcellos col. (CTUFPB-3241) ; Piauí: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-43.488945&amp;materialsCitation.latitude=-9.222666" title="Search Plazi for locations around (long -43.488945/lat -9.222666)">Caracol</a> (9°13'21.6"S 43°29'20.2"W): 11.XII.2010, A. Vasconcellos col. (CTUFPB-4090) ; Bahia: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-41.11697&amp;materialsCitation.latitude=-10.231083" title="Search Plazi for locations around (long -41.11697/lat -10.231083)">Campo Formoso</a> (10°13'51.9"S 41°07'01.1"W): 12.VII.2022, R. R. Ferreira col. (CTUFPB-11865) .</p><p>Diagnosis: As described for the genus. EV with three semi-spherical sclerotized plates of different sizes, with the middle plate larger than the other two, all with spines.</p><p>Etymology: In honor of the city of Catolé do Rocha, the type locality, a peaceful and welcoming town in the sertão of Paraíba, located in northeastern Brazil.</p><p>Imago: Unknown.</p><p>Worker: As described for the genus. EV asymmetric, with an armature composed of three semispherical sclerotized plates that bear several spines: one larger and central, and two smaller, adjacent, non-identical (Fig. 5A and 5D). An occasional single spine, the same size as the spines on the plates, is isolated adjacent to the smaller plate (Fig. 5B). The cushions are subrectangular and elongated, with 30 to 40 elongated scales in the proximal region (some of which may be fringed), and 20 to 30 smaller, entirely fringed scales in the basal portion (Fig. 5C).</p><p>Comparison. The left mandible incision between M1+2 and M3 of Triclavitermes catoleensis sp. nov. is wider than in most genera, e.g., Anoplotermes, Krecekitermes, Ourissotermes, but narrower than in Mangolditermes . T. catoleensis sp. nov. differs from Ourissotermes giblinorum Scheffrahn, Carrijo &amp; Castro, 2023 and Echinotermes biriba Castro &amp; Scheffrahn, 2018 by having a more prominent mesenteric tongue. Additionally, E. biriba has a highly enlarged crop, a feature not found in T. catoleensis sp. nov. In O. giblinorum, the EV is inserted directly into the P3, without a distinct EVS, while in T. catoleensis sp. nov., the EVS is conspicuous. Anoplotermes spp. and Humutermes spp. have a mesenteric tongue with a whitish spherical mesenteric nodule in ventral view, but can be distinguished from T. catoleensis sp. nov. by the shape of the enteric valve seating, which forms a ring in T. catoleensis sp. nov., absent in Anoplotermes spp. and Humutermes spp. The asymmetric pattern of the EV armature in T. catoleensis sp. nov. closely resembles that observed in Anoplotermes susanae Scheffrahn, Carrijo &amp; Castro, 2023, Anoplotermes janus Bourguignon &amp; Roisin, 2010 (Fig. 6F), and Anoplotermes parvus Snyder, 1923 (Fig. 6G–H), which also feature an armature extending along only one or three of the six cushions, or even along all cushions, yet still asymmetric. Dissimulitermes invisibilis also exhibits an asymmetric enteric valve armature, with similar plates extending across five cushions, and only one cushion bearing a plate of a distinct shape. However, the shape of the armature plates is clearly different among these species. The ornamentation and armature of the enteric valve in T. catoleensis sp. nov. also set it apart from all other genera of Apicotermitinae . The EVs of E. biriba, O. giblinorum, Patawatermes spp. (Fig. 6C–D), and Humutermes spp. have six symmetrical and sclerotized plates with spines, while the EV of T. catoleensis sp. nov. is asymmetrical, with only three distinct sclerotized plates. Although Rustitermes boteroi was recovered as close related to T. catoleensis sp. nov. in the Bayesian inference (BI) tree (Fig. 8, see results below), its morphological traits are markedly different. It lacks an armed enteric valve, exhibits a distinct coiling pattern of the digestive tube, and has a row of spines on the foretibia—a feature absent in T. catoleensis sp. nov. Triclavitermes gen. nov. resembles Chasitermes by the ring-shaped arrangement of the EVS, but differs by the EV, as Chasitermes has an unarmed EV with rectangular, non-sclerotized cushions (Scheffrahn &amp; Carrijo 2023).</p></div>	https://treatment.plazi.org/id/03B06032E11ECE085ED6FB5EFEE9FE4C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ferreira, Renan Rodrigues;Carvalho, Antonio;Lucena, Emanuelly Félix De;Figueiredo, Rozzanna Esther Cavalcanti Reis De;Vasconcellos, Alexandre	Ferreira, Renan Rodrigues, Carvalho, Antonio, Lucena, Emanuelly Félix De, Figueiredo, Rozzanna Esther Cavalcanti Reis De, Vasconcellos, Alexandre (2025): Triclavitermes catoleensis (Termitidae: Apicotermitinae), a new soldierless termite genus and species from the Caatinga dry forest, northeastern Brazil. Zootaxa 5706 (3): 383-396, DOI: 10.11646/zootaxa.5706.3.4, URL: https://doi.org/10.11646/zootaxa.5706.3.4
03B06032E111CE055ED6FF65FA43FEDE.text	03B06032E111CE055ED6FF65FA43FEDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Apicotermitinae	<div><p>Updates to the generic key for workers of Apicotermitinae (Carrijo et al. 2023, 338)</p><p>7 Enteric valve armature consists of six palmate pads................................................ Humutermes</p><p>- Enteric valve armature with one to three sclerotized pads or enteric valve unarmed................................ 7 ’</p><p>7’ Enteric valve seating tubular of variable size, or trilobed (only in Anoplotermes meridianus), and enteric valve armature with one to three sclerotized pads/single spines, or enteric valve unarmed.................................. Anoplotermes</p><p>- Enteric valve seating ring-shaped and armature with three spherical, sclerotized spiny pads....... Triclavitermes gen. nov.</p></div>	https://treatment.plazi.org/id/03B06032E111CE055ED6FF65FA43FEDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ferreira, Renan Rodrigues;Carvalho, Antonio;Lucena, Emanuelly Félix De;Figueiredo, Rozzanna Esther Cavalcanti Reis De;Vasconcellos, Alexandre	Ferreira, Renan Rodrigues, Carvalho, Antonio, Lucena, Emanuelly Félix De, Figueiredo, Rozzanna Esther Cavalcanti Reis De, Vasconcellos, Alexandre (2025): Triclavitermes catoleensis (Termitidae: Apicotermitinae), a new soldierless termite genus and species from the Caatinga dry forest, northeastern Brazil. Zootaxa 5706 (3): 383-396, DOI: 10.11646/zootaxa.5706.3.4, URL: https://doi.org/10.11646/zootaxa.5706.3.4
