identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B187B9FFCBFF9B00FEFC56A2E6F89D.text	03B187B9FFCBFF9B00FEFC56A2E6F89D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia (sect. Eugeniopsis) (O. Berg) M. F. Santos & E. Lucas.	<div><p>Myrcia sect. Eugeniopsis (O. Berg) M.F.Santos &amp; E. Lucas. Type: Eugenia laevigata De Candolle (1828: 283) .</p><p>= Eugeniopsis O. Berg (1855 –1856: 80).</p><p>= Marlierea sect. Eugeniopsis (O. Berg) Nied. (1893: 76) .</p><p>= Marlierea subg. Eugeniopsis (O.Berg) Kiaerskou (1893: 50)</p><p>= Marlierea sect. Pseudocalyptra D. Legrand (1975: 7) .</p><p>Shrub to tree. Trichome reddish, dibrachiate or simple; pellucid dots densely present on leaves and reproductive structures. Twig cylindrical when mature; branching usually monopodial (rarely sympodial); cataphylls present or absent at the base of the Seasonal Growth Unit (SGU – sensu Briggs and Johnson 1979). Leaf with midvein sulcate on the adaxial surface and raised on the abaxial surface, secondary veins usually thin and never strongly raised. Inflorescence pyramidal, axillar at terminal or subterminal nodes of the SGU (central bud developing a vegetative branch) or often ramiflorous (i.e., axillar in previous SGU), one inflorescence per axillary bud or often a pair arising in a short internode with abortive central bud, inflorescence pherophyll usually deciduous, opposite branching, three branching per node; bracts and bracteoles usually early deciduous. Flower (3)4–5(6)–merous, flower bud clavate (rarely turbinate or globose), corolla usually barely apparent before anthesis; hypanthium extending into a tube beyond the ovary, not tearing at anthesis or often with a small vertical rip (but usually not tearing the staminal ring), internally glabrous; calyx not fused and deciduous parallel to the hypanthium ring (rarely totally fused and opening irregularly); corolla without distinctive features; staminal ring thin, usually comprising less than 30% of total disc width, usually with trichomes, anther thecae of equal height, reversing curvature on dehiscence, exposing interior of sacs as a convex surface; ovary glabrous at the apex, 2-locular with 2 ovules per locule. Fruit globose, base rounded or attenuate, hypanthium tube persistent but not straight, calyx remains falling or marcescent (rarely accrescent); seeds 1–2(3) per fruit (Figure 1).</p><p>Distribution and Habitat:— Myrcia sect. Eugeniopsis comprises 16 species mainly distributed in rainforest areas (without a dry season) of the Atlantic Forest domain (Figure 2, Table 1), whose southern region was particularly significant for the initial diversification of the group (Santos et al. 2017). Montane rainforests along the Serra do Mar and other mountains ranges hold the highest diversity, with 13 species (four endemic). Diversity decreases towards the southern and northern limits of the Atlantic Forest, though there are endemic subtropical species: Myrcia hatschbachii D.Legrand and Myrcia oblongata DC., with the latter occurring in seasonal forests in Argentina and Paraguay. Records from the northern part of Atlantic Forest (Northeast Brazil, except southern Bahia) are sparse and do not include any endemic species. In seasonal habitats, eight species have been recorded, but only Myrcia advena M.F.Santos is endemic (Table 1). Myrcia clausseniana (O.Berg) A.Maruy. &amp; Gaem is especially common along the Espinhaço range (Cerrado and Caatinga domains), and Myrcia teuscheriana (O.Berg) M.F.Santos is frequently recorded in semideciduous forests along the Rio Doce basin. Other species, such as Myrcia vellozoi Mazine, occur occasionally in seasonal areas. Species of Myrcia sect. Eugeniopsis inhabitat a range of vegetation types, including campo rupestre, restinga, cerrado, semideciduous forest, and rainforest, with the latter being by far the most common habitat.</p><p>Accepted species LowAF MounAF ArauF SeasoAF Cerrado Caatinga CRup CoSt Myrcia advena X CR Myrcia clausseniana X X X LC Myrcia eugenioides X X X VU Myrcia eugeniopsoides X LC Myrcia ferruginosa X LC Myrcia gaudichaudiana X X X DD Myrcia hatschbachii X LC Myrcia maculata X DD Myrcia oblongata X X X LC Myrcia polygama X X X DD Myrcia reitzii X DD Myrcia suberosa X X CR Myrcia tenondeporan X VU Myrcia tenuivenosa X X LC Myrcia teuscheriana X X X DD Myrcia vellozoi X X X LC</p><p>Phenology:— Flowering occurs from spring through the first half of summer (September to January in South Hemisphere). It typically begins in late winter (August) and decreases abruptly in the later part of summer (February to March), with flowering being rare outside this period. Flowering appears to peak around early summer, though this may reflect a sampling bias, as most botanical expeditions in Brazil occur during summer vacation months. Records indicate mature fruits during throughout the year, although fruit ripening appears to be most common from late spring to the end of summer.</p><p>Conservation Status:— Our preliminary conservation analysis of the 16 species within Myrcia sect. Eugeniopsis indicate that four species face a level of extinction threat (Table 1). This finding is unsurprising, given that the primary habitat of this group, the Atlantic Forest Domain, retains only 24% of its original area, much of which consists of small, disturbed, and disconnected fragments (Fundação SOS Mata Atlântica &amp; INPE 2022). Additionally, some species are classified as Data Deficient, yet are likely threatened to some extent. Several species were documented within conservation units, an encouraging result about their protection; only Myrcia maculata, Myrcia advena, and Myrcia suberosa were not recorded in these protected areas. However, it is critical to preserve forest fragments outside conservation units and ensure connectivity among populations to safeguard the genetic diversity and long-term survival of these species.</p><p>Morphology:— The description above provides a concise overview of Myrcia sect. Eugeniopsis morphology, emphasizing the typical features of the group and those that distinguish it from other Myrcia sections. Notable traits (though not universal across all species) include numerous and visible pellucid dots, reddish trichomes, clavate floral bud with a barely visible corolla, and small, free calyx lobes. Another typical feature is the discolorous leaf blade seen in dry specimens, where the adaxial surface darkens, contrasting with the lighter abaxial surface. The inflorescence structure is also distinctive: it emerges at subterminal nodes of the season growth unit (SGU), with an early deciduous inflorescence bract (or pherophyll). Variation in inflorescence structure is significant for species circumscription. Five species ( Myrcia advena, Myrcia hatschbachii, Myrcia oblongata, Myrcia tenuivenosa and Myrcia eugeniopsoides) exhibit unique floral bud morphology. The first four display an open floral bud revealing the corolla before anthesis, which led to previous placement in Myrcia rather than Eugeniopsis or Marlierea (De Candolle 1828, Kiaerskou 1893, Legrand 1961). Conversely, Myrcia eugeniopsoides (D.Legrand &amp; Kausel) Mazine has a closed floral bud lacking distinct calyx lobes, that splits irregularly during anthesis, a trait that led to its initial placement in Calyptranthes (Legrand 1962b) (Figure 1).</p><p>Myrcia sect. Eugeniopsis shows morphological similarity to Myrcia sect. Aulomyrcia and Myrcia sect. Sympodiomyrcia . The resemblance to M. sect. Sympodiomyrcia is especially notable in species with reddish trichomes and clavate floral bud, such as Myrcia crassa Sobral (2010: 138) and Myrcia pseudomarlierea Sobral (2010: 147) . While Santos et al. (2019) initially included Myrcia crassa and Myrcia pseudomarlierea in M. sect. Eugeniopsis, they are reassigned here to M. sect. Sympodiomyrcia based on taxonomic reanalysis. Both species exhibit consistent sympodial branching and an inflorescence that arises at the last internode of the previous SGU, features that are not typical of M. sect. Eugeniopsis . Recent phylogenetic analysis further supports the placement of M. pseudomarlierea in M. sect. Sympodiomyrcia (NMWG 2024). Overall, Myrcia sect. Sympodiomyrcia can be differentiated by features such as sympodial branching, a cataphyll at the SGU base, an inflorescence arising at the terminal node of the previous SGU, and a fruit with a straight hypanthium tube (Santos et al. 2018). Considering Myrcia sect. Aulomyrcia, the resemblance to Myrcia sect. Eugeniopsis is particularly strong in species with pyramidal inflorescences bearing deciduous bracts, clavate floral buds, and free and deciduous calyx lobes. However, these species can be distinguished by the trichome colour (white to light brown in M. sect. Aulomyrcia), absence of numerous pellucid dots and leaf blade concolorous when dry.</p><p>Phylogeny and Biogeography:— Lucas et al.(2007) included for the first time a species of Myrcia sect. Eugeniopsis in a phylogenetic study ( Myrcia eugeniopsoides), but the group was recovered later in a phylogenetic hypothesis by Lucas et al. (2011). Santos et al. (2016b) conducted a phylogenetic analysis of Myrcia sect. Sympodiomyrcia, including a broader sampling of Myrcia sect. Eugeniopsis (nine species) to investigate phylogenetic relationship among the sections. Santos et al. (2016b) showed that the groups are not closely related, with morphological similarities being convergences. Myrcia sect. Eugeniopsis consistently appears (posterior probability usually 1.00) as a sister group of Myrcia sect. Tomentosae E.Lucas &amp; D.F.Lima (2018: 8–9), despite their distinct morphologies (Santos et al. 2016b, Lucas et al. 2018). Santos et al. (2016b) included a representative sample of Myrcia sect. Eugeniopsis morphology, providing a basis for the section circumscription and the addition of species not included in the phylogenetic analysis. The phylogenetic analysis identified a clade comprising the three species with open floral bud ( M. hatschbachii, M. oblongata and M. tenuivenosa), suggesting a single evolutionary origin for this feature (Santos et al. 2016b, 2017). Santos et al. (2017) estimated that the section originated in the Miocene (17.1–8.2 Ma, 95% HPD) within the forests of the Serra do Mar Mountains. Despite these insights, phylogenetic and biogeographical studies specifically focused on Myrcia sect. Eugeniopsis are still missing.</p></div>	https://treatment.plazi.org/id/03B187B9FFCBFF9B00FEFC56A2E6F89D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFC7FF9B00FEFE8EA0CCFFB2.text	03B187B9FFC7FF9B00FEFE8EA0CCFFB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia (sect. Eugeniopsis)	<div><p>Key to the species of Myrcia sect. Eugeniopsis</p><p>1.</p><p>1’.</p><p>2.</p><p>2’.</p><p>3.</p><p>3’.</p><p>4.</p><p>4’.</p><p>5.</p><p>5’.</p><p>6.</p><p>6’.</p><p>7.</p><p>7’.</p><p>8.</p><p>8’.</p><p>9.</p><p>9’.</p><p>10.</p><p>10’.</p><p>11.</p><p>11’.</p><p>12.</p><p>12’.</p><p>13.</p><p>13’.</p><p>14.</p><p>14’.</p><p>15.</p><p>15’.</p><p>16.</p><p>16’.</p></div>	https://treatment.plazi.org/id/03B187B9FFC7FF9B00FEFE8EA0CCFFB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFC6FF9C00FEFF5CA162FD66.text	03B187B9FFC6FF9C00FEFF5CA162FD66.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia advena M. F. Santos 2025	<div><p>1. Myrcia advena M.F.Santos, sp. nov. Type: — Brazil. Distrito Federal: Brasília, University campus, cerrado along road parallel to lago Paranoá, 6 October 1975 (fl., fr.), Oldenburger 1685 (holotype UB!; isotypes K!, NY!, SP!) (Figures 3, 4).</p><p>Diagnosis: — Myrcia advena is related to Myrcia oblongata but differs by the swollen node (vs. slender), cataphylls persistent or deciduous (vs. deciduous), leaf blade with base obtuse to rounded or truncate to emarginate (vs. narrowly cuneate to rounded), abaxial surface glabrous or subglabrate (vs. pubescent to puberulent), and inflorescence with opposite apical branching (vs. subopposite).</p><p>Description:— Shrub to tree, 2.5 to 5.0 m high. Trichome white, ochraceous or often reddish, simple or dibrachiate, 0.08–0.48 mm. Twig when young flattened, puberulent, when mature greyish (when dry), cylindrical, cortex slightly cracked; branching monopodial, internode 1.5–5.2 cm, node swollen; cataphylls present (sometimes more than one pair per SGU), deciduous (scale-like) or persistent (leafy); buds sericeous. Leaf concolorous, chartaceous, blade 1.9– 8.9 × 1.4–4.2 cm, widely elliptic, ovate to widely ovate, obovate to widely obovate, sometimes circular, apex obtuse to rounded, base obtuse to rounded, truncate or emarginate, margin plane, secondary veins 3–8 mm apart, held at an angle of 65–80° relative to the midvein, marginal vein 1.0– 1.5 mm from the margin, tertiary veins conspicuous to inconspicuous in both surfaces, areoles 0.5–1.0 mm, pellucid dots conspicuous in the abaxial surface, ca. 7 per mm 2; adaxial surface glabrous, secondary veins plane to raised; abaxial surface subglabrate to glabrous, secondary veins plane to raised; petiole 2–4 × 2–4 mm, canaliculate, puberulent to glabrous. Inflorescence 2.5–9.0 × 2.0–4.0 cm, 10– 21 flowers, axillar at the subterminal nodes of the SGU (central bud developing a vegetative branch), one inflorescence per axillary bud, inflorescence pherophyll persistent, opposite branching (rarely subopposite), three branching per node (rarely four), rachis puberulent to subglabrate, first internode of central rachis ca. 1 mm wide; bracts deciduous, not seen; bracteoles ca. 0.8 × 0.6 mm, deciduous, lanceolate to widely ovate, concave or keeled, apex acuminate or rounded, base truncate, externally subglabrate to glabrous, internally glabrous (often ciliate). Flower bud 3.5–4.0 × 3.0– 3.5 mm, obovate, corolla apparent before anthesis; hypanthium ca. 0.8 mm extending above the summit of the ovary, not tearing at anthesis, externally puberulent to glabrous, pellucid dots covering the whole surface; calyx 5–merous, lobes 0.4–1.2 × 0.6–2.0 mm, not fused, apparently persistent, depressed ovate to widely depressed ovate, concave, apex rounded, both surfaces puberulent to glabrous (often ciliate); corolla 5–merous, petals white, 0.8–2.4 × 1.4–2.8 mm, widely depressed ovate or very widely ovate, concave, apex rounded, both surfaces puberulent to glabrous (often ciliate); staminal ring 0.2–0.4 mm wide, subglabrate to glabrous, stamens 43–65, filament 2.0– 3.8 mm long, anther 0.24–0.56 × 0.32–0.48 mm, gland at the apex; ovary 0.8–1.2 × 0.8–1.2 mm, obconic, style ca. 4.8 mm long, glabrous, stigma punctiform. Fruit immature green, globose, base rounded, glabrous, totally covered by pellucid dots, hypanthium and calyx present; seeds not seen.</p><p>Distribution and Habitat: —The new species is the only member of Myrcia sect. Eugeniopsis endemic to the Cerrado domain, currently recorded exclusively in the Brazilian Federal District (Figure 4). It inhabits the savanna vegetation (cerrado sensu stricto) and swamp areas (“brejo”) at elevation around 1,000 meters.</p><p>Phenology: —Flowering and fruiting specimens have been collected in August to November.</p><p>Etymology: —The specific epithet advena (meaning “foreigner”) honours the immigrant workers who, through their labour, built the Brazilian Federal District, the only known location of this new species. Additionally, Myrcia advena represents a “migrant” within Myrcia sect. Eugeniopsis, providing a westward expansion of the group distribution.</p><p>Conservation Status:— According to the Geocat analysis, the species has an Extent of Occurrence of 23 km 2 (threshold of Critically Endangered), and the Area of Occupancy is 12 km 2 (threshold of Endangered) (sensu IUCN 2022). The species is known from only four records in the Federal District, a well-documented region (sensu Campbell 1989) with more than one record per km 2 (estimation based on records of speciesLink; CRIA 2024). The Cerrado vegetation is frequently affected by anthropogenic fire, a situation intensified in recent years by climate changes (Tomas et al. 2024). Due to these factors, we preliminarily classify this species as Critically Endangered (CR, criteria B2a, biii; IUCN 2022).</p><p>Taxonomy:— Myrcia advena is closely related to Myrcia oblongata, sharing several distinctive features: a shrub to tree habit, trichomes usually white to ochraceous, monopodial branching, leaf blade with an apex usually obtuse to rounded (though less angular in M. advena), axillary inflorescence at subterminal nodes of the SGU, pherophyll persistent (sometimes deciduous in M. oblongata), obovate flower bud with a visible corolla, and pentamerous flower with persistent calyx. These species exhibit a disjunct distribution, and further phylogenetic studies including M. advena would be valuable to explore their shared biogeographic history.</p><p>Illustrations and images in previous works:— None.</p><p>Additional specimens examined:— BRAZIL. Distrito Federal: Brasília, 12 August 1980 (fl.), R . C . Mendonça 144 (K); ibidem, 22 September 1975 (fl.), J . E . Paula 1072 (IBGE, SPF); ibidem, 25 September 1961 (fl.), E . P . Heringer 8695 (SP) .</p></div>	https://treatment.plazi.org/id/03B187B9FFC6FF9C00FEFF5CA162FD66	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFDAFF8B00FEFDA4A231F941.text	03B187B9FFDAFF8B00FEFDA4A231F941.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia eugenioides Cambessedes 1832	<div><p>3. Myrcia eugenioides Cambessèdes (1832 –1833: 302) (Figures 11, 12).</p><p>≡ Calyptromyrcia eugenioides (Cambess.) O. Berg (1855 –1856: 34). Marlierea eugenioides (Cambess.) D. Legrand (1958: 265) . Type: — Brazil, Rio de Janeiro, 1816–1821 (fl.), Saint-Hilaire A1-389 (lectotype P! [P00161444] designated by Santos et al. 2019; isolectotypes MPU! [MPU011021] [image], P! [P00161445], P! [P00161446]).</p><p>= Calyptromyrcia eugenioides var. glabra O. Berg (1857 –1859: 56). Type: — Brazil, Rio de Janeiro, Jurujuba bay, 1837 (fl.), Gardner 758 (neotype K! [K000330592] designated by Santos et al. 2019; isoneotypes E! [E00504684] [image], E! [E00504685] [image], G!, K! [K000330593], K! [K000342676], NY! [NY00405395] [image], P! [P00547506]).</p><p>= Aulomyrcia lineata O. Berg (1857 –1859: 68). Myrcia lineata (O.Berg) Niedenzu (1893: 76) . Type: — Brazil, Rio de Janeiro, mun. Rio de Janeiro, “Habitat in silvis montis prope Praya dos Flamingos, Sebastianopoli, prov. Rio de Janeiro ” (in the protologue), September 18?? (fl.), Beyrich s.n. (syntype B†). Neotype: — Brazil, Rio de Janeiro, 1844 (fl.), Widgren 787 (BR!, designated by Santos et al. 2019).</p><p>Description:— Tree, 3 to 9 m high. Trichome reddish, simple or dibrachiate, 0.05–0.3 mm. Twig when young flattened and distally sulcate, tomentose, when mature reddish to greyish (when dry), cylindrical, glabrous, cork slightly cracked; branching monopodial, internode 2.0– 6.5 cm, node slender; cataphylls absent; buds pubescent. Leaf concolorous, chartaceous, blade 5.5–16.8 × 2.3–7.5 cm, elliptic or oblong, apex acuminate to obtuse, base cuneate, sometimes attenuate, margin plane, secondary veins 4–9 mm apart, held at an angle of 70–80° relative to the midvein, marginal vein 1.0–3.0 mm from the margin, tertiary veins conspicuous to barely conspicuous in both surfaces, areoles 0.5–1.0 mm, pellucid dots conspicuous in the abaxial surface, 6–8 per mm 2; adaxial surface glabrous, secondary veins flat to raised, usually thin; abaxial surface glabrous (rarely subglabrate), secondary veins flat to raised; petiole 6–9 × 2-3 mm, canaliculate, glabrous. Inflorescence 5.0–20.5 × 2.5–17.0 cm, 30–90 flowers, axillar at the terminal or subterminal nodes of the SGU (central bud usually not developed), one or two inflorescence per axillary bud (rarely a short internode with a pair of lateral inflorescences and an abortive central bud), inflorescence pherophyll persistent or deciduous, opposite or subopposite branching, three to five branching per node, rachis puberulent to glabrous, first internode of central rachis 1–3 mm wide; bracts 2.6–3.2 × 1.2–1.6 mm, deciduous, elliptic or ovate, plane or concave, apex acute to obtuse, base cuneate, externally puberulent to glabrous, internally glabrous; bracteoles 2.0–5.0 × 2.0–5.0 mm, deciduous, ovate to widely ovate, concave, apex acuminate to acute, base truncate, both surfaces puberulent. Flower bud ca. 4.0 × 3.0 mm, clavate with obconic base, corolla hidden or barely apparent before anthesis; hypanthium 1.2–1.8 mm extending above the summit of the ovary, not tearing at anthesis or with a small vertical rip (but not tearing staminal ring), externally puberulent to glabrous, pellucid dots conspicuous; calyx 4–5–merous, lobes 0.4–2.5 × 1.0– 2.5 mm, not fused, deciduous parallel to the hypanthium ring, depressed ovate to widely depressed ovate, concave, apex rounded, externally puberulent to glabrous, internally pubescent to puberulent; corolla 4–5–merous, petals cream to white, 1.2–2.0 × 1.6–2.4 mm, widely depressed ovate, very widely ovate or very widely obovate, plane or concave, apex rounded, both surfaces puberulent to glabrous; staminal ring 0.2–0.4 mm wide, puberulent, stamens 35–80, filament 2.4–4.8 mm long, anther 0.24–0.40 × 0.24–0.48 mm, gland at the apex; ovary 0.8–1.0 × 0.8–1.0 mm, obconic, style ca. 7.2 mm long, subglabrate to glabrous, stigma punctiform. Fruit not seen.</p><p>Distribution and Habitat: — Myrcia eugenioides is recorded within the Atlantic Forest domain, with an atypical record in a semideciduous forest in the Cerrado domain (Felício dos Santos municipality) (Figure 12). Mostly records come from the central region of Rio de Janeiro state, with two additional records from Bahia and Alagoas state. The species inhabits rain forest or restinga vegetation (either open or forested), and occasionally occurs in flooded or semideciduous forests. Elevations range from 300 to 1,320 meters.</p><p>Phenology: —The species flowers from June to December. Fruits have been recorded (immature) in February and October.</p><p>Conservation Status: —The species has an Extent of Occurrence of ca. 276,284 km 2 (threshold of Least Concern) and a Small Area of occupancy (40 km 2, threshold of Endangered) (sensu IUCN 2022). Myrcia eugenioides is known from less than 10 localities, with only two of these occurring within Conservation Units. Available data does not allow the estimation of population size and fluctuation, but the Atlantic Forest currently has only 24% of its original extension, and many areas are small, disturbed, and disconnected fragments (Fundação SOS Mata Atlântica &amp; INPE 2022). Based on this scenario, we preliminarily assess the species as Vulnerable (VU, criteria B2a, bii, biii; IUCN 2022).</p><p>Taxonomy: — Myrcia eugenioides is characterized by thick and reddish to greyish young branches, monopodial branching, large elliptic to oblong leaves with acuminate to obtuse apex and usually glabrous with thin secondary veins on the adaxial surface, inflorescences arising from multiple nodes along the SGU, and flower bud with obconic base. The species resembles Myrcia gaudichaudiana (for instance Braga 4554, Kuhlmann s.n. [RB 111588]), but can be distinguished by its branch, the large leaves with thin secondary veins and the shape of the flower bud. Specimens from Felício dos Santos municipality exhibit the typical morphological characteristics of the species, despite the unique occurrence in the Cerrado domain.</p><p>Illustrations and images in previous works: — Fernandes et al. (2020).</p><p>Additional specimens examined: — BRAZIL. Unknown province /state, no date(fl.), F . Sellow s.n. (LE). Alagoas: Mun. Boca da Mata, 29 October 1980 (fl.), G. L . Esteves 611 (HUEFS, RB). Bahia: Mun. Cairú, 13 November 2003 (fl.), J. L . Paixão 309 (BHCB, CEPEC). Minas Gerais: unknown municipality, 13 February 1972 (fr.), W. R . Anderson 35932 (NY, RB). Mun. Felício dos Santos, 8 October 2004 (fr.), P. L . Viana 1890 (BHCB); ibidem, 29 August 2008 (fl.), P. L . Viana 3705 (BHCB). Rio de Janeiro: unknown municipality, no date (fl.), J . Miers s.n. (P00547504); no date (fl.), J . Miers s.n. (P00547505). Mun. Niterói, no date (fl.), J . Miers 3697 (K); ibidem, 25 November 2018 (fl.), T . Fernandes 422 (RB). Mun. Rio de Janeiro, 2 September 1943 (fl.), J. G . Kuhlmann s.n. (NY, RB111588); ibidem, 18 December 1997 (fl.), J. M. A . Braga 4554 (RB); ibidem, 1945 (st.), O. Machado s.n. (RB75807); ibidem, 5 May 1947 (st.), O . Machado s.n. (RB75144). Mun. Saquarema, 23 April 1991 (st.), C . Farney 3214 (ESA, RB). Mun. Teresópolis, 26 June 2007 (fl.), M . Nadruz 1832 (RB) .</p></div>	https://treatment.plazi.org/id/03B187B9FFDAFF8B00FEFDA4A231F941	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFE9FFB400FEFB38A5FBFFDD.text	03B187B9FFE9FFB400FEFB38A5FBFFDD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia gaudichaudiana (O. Berg) M. F. Santos	<div><p>6. Myrcia gaudichaudiana (O.Berg) M.F.Santos (in Santos et al. 2019: 3) (Figures 19, 20).</p><p>≡ Eugeniopsis gaudichaudiana O. Berg (1857 –1859: 147). Marlierea gaudichaudiana (O.Berg) Loefgren &amp; Everett (1919: 219) . Type: — Brazil, Rio de Janeiro, no date (fl.), Schüch 1035 (number 5878 is also presented in original labels) (lectotype W! [W0047995] designated by Santos et al. 2019; isolectotypes F! [876125], K! [K000330596], W! [W0047994]). Remaining syntype: — Brazil, Rio de Janeiro, 1830 (fl.), Gaudichaud-Beaupré s.n. (G! [G00301890]).</p><p>= Eugeniopsis schottiana O. Berg (1857 –1859: 148), syn. nov. Marlierea schottii D. Legrand (1962a: 33) . Myrcia schottii (D.Legrand) M.F.Santos (in Santos et al. 2019: 6). Type:— Brazil, Rio de Janeiro, 1837(?) (fl.), Schott 1083 (number 5866 is also cited in the original label) (lectotype W! [W0037061] designated by Santos et al. 2019; isolectotypes B†, K! [K000330595]).</p><p>Description:— Shrub to tree, 4 to 12 m high. Trichome reddish, simple or dibrachiate, 0.08–0.16 mm. Twig when young flattened and distally sulcate, puberulent, when mature greyish to stramineous (when dry), cylindrical, cortex slightly cracked; branching monopodial, internode 1.9–5.8 cm, node slender; cataphylls rarely present, deciduous; buds pubescent. Leaf concolorous, chartaceous, blade 5.0–11.9 × 2.0– 5.9 cm, elliptic to widely elliptic, oblong or ovate, apex acuminate to obtuse, base cuneate to obtuse (rarely rounded) or slightly attenuate, margin plane, secondary veins 2–8 mm apart, held at an angle of 65–70° relative to the midvein, marginal vein 0.5–3.0 mm from the margin, tertiary veins inconspicuous or sometimes barely conspicuous in both surfaces, areoles ca. 0.5–1.0 mm, pellucid dots conspicuous in the abaxial surface, 1–12 per mm 2; adaxial surface glabrous (except midvein), secondary veins plane to raised; abaxial surface glabrous (often puberulent), secondary veins plane to raised; petiole 4–8 × 1–2 mm, canaliculate, puberulent to glabrous. Inflorescence 4.5–10.0 × 2.0–6.0 cm, 20–45 flowers, axillar at subterminal (rarely terminal) nodes of the SGU (central bud developing a vegetative branch), one inflorescence per axillary bud, inflorescence pherophyll deciduous, opposite branching, three branching per node, rachis puberulent, first internode of central rachis 1–2 mm wide; bracts deciduous, not seen; bracteoles 1.2 × 0.2–0.4 mm, deciduous, lanceolate to ovate, concave or keeled, apex acuminate, base truncate, both surfaces puberulent. Flower bud 2.2–3.2 × 1.4–2.0 mm, clavate, corolla hidden or barely apparent before anthesis; hypanthium 0.8–1.7 mm extending above the summit of the ovary, not tearing at anthesis or with a small vertical rip (but not tearing the staminal ring), externally pubescent to puberulent, pellucid dots covering the whole surface; calyx 4–merous, lobes 0.6–1.4 × 0.8–1.8 mm, not fused, deciduous parallel to the hypanthium ring, depressed ovate to widely depressed ovate or ovate to very widely ovate, concave, apex rounded, both surfaces puberulent to glabrous; corolla 4–merous, petals white to cream, 1.2–2.4 × 0.8–2.0 mm, widely depressed ovate, very widely ovate or widely obovate to very widely obovate, plane, apex rounded, both surfaces puberulent to glabrous; staminal ring 0.2–0.4 mm wide, puberulent to glabrous, stamens 44–71(100), filament 1.6–4.4 mm long, anther 0.16–0.32 × 0.24–0.40 mm, gland at the apex; ovary 0.6–1.0 × 0.6–1.0 mm, obconic, style 3.6–7.2 mm long, subglabrate to glabrous, stigma punctiform. Fruit mature black, ca. 1.0 × 1.0 cm, globose, base rounded, glabrous, totally covered by pellucid dots, remnants of hypanthium and calyx present; seeds not seen.</p><p>Distribution and Habitat: — Myrcia gaudichaudiana is found from the states of Rio de Janeiro to Pernambuco, primarily within the Atlantic Forest domain from 17 to 750 m of elevation (Figure 20). It typically inhabits rain forest, with occasional occurrences in flooded forests, seasonal forest (such as “mata de cipó”), and montane forests within Caatinga domain (“brejo”). It can be found in the understory or along forest edges.</p><p>Phenology: —Flowers occurs from October (peak) to February, with immature fruits observed in March.</p><p>Conservation Status: —The species has an Extent of Occurrence of ca. 333,101 km 2 (threshold of Least Concern) and a small Area of Occupancy (52 km 2, threshold of Endangered) (sensu IUCN 2022). Myrcia gaudichaudiana is documented in over 10 localities (22 records) throughout its range, though only two of these are within Conservation Units. Many populations are in the northern Atlantic Forest, which faces significant environmental pressures (Fundação SOS Mata Atlântica &amp; INPE 2022). Giving the conflicting data and limited information available, the species is preliminarily classified as Data Deficient (DD; IUCN 2022).</p><p>Taxonomy: — Myrcia gaudichaudiana is morphologically similar to Myrcia tenondeporan and Myrcia vellozoi, with overlaps in their circumscriptions. All three species share monopodial branching, typically elliptic leaf blades, and axillar inflorescence at terminal or subterminal nodes of the SGU. Myrcia gaudichaudiana is distinguished by its smaller leaf blade (compared to M. vellozoi), its leaf apex (less prolonged than in M. tenondeporan and M. vellozoi), glabrous or often puberulent leaves (versus glabrous to tomentose in M. vellozoi), and flower bud size (larger than M. tenondeporan and generally smaller than M. vellozoi). The distributions of M. gaudichaudiana and M. vellozoi overlap from Rio de Janeiro to Bahia states, but M. gaudichaudiana is found further norther, whereas M. gaudichaudiana and M. tenondeporan do not co-occur.</p><p>Myrcia schottii is here considered a synonym of M. gaudichaudiana due to absence of significant morphological differences between the type specimen of M. schottii (the only known collection) and the leaf and flower bud variations in M. gaudichaudiana . The primary distinguishing feature of M. schottii, a long internode and long inflorescence peduncle, is regarded as an atypical variation. The vernacular names “murta” or “murta-branca” have been recorded for the species in Pernambuco state.</p><p>Illustrations and images in previous works: — Berg (1857 –1859, as Eugeniopsis gaudichaudiana), Fernandes et al. (2020, as Myrcia gaudichaudiana and M. schottii).</p><p>Additional specimens examined: — BRAZIL. Unknown province /state, no date (fl.), J. B. E . Pohl s.n. (K000330502). Alagoas: unknown municipality, 28 October 1980 (fl.), Andrade-Lima 80-9738 (IPA); 28 October 1980 (fl.), Andrade-Lima 80-9755 (IPA); 28 October 1980 (fl.), Andrade-Lima 80-9778 (IPA). Bahia: Mun. Belmonte, 9 November 1967 (fl.), R. S . Pinheiro 401 (CEPEC, MBM, MBML, SPF). Mun. Jequié, 20 October 1970 (fl.), T. S . Santos 1228 (CEPEC, SPF); ibidem, 21 June 2004 (st.), G. E. L . Macedo 968 (PEUFR). Espírito Santo: Mun. Linhares, 30 October 1991 (fl.), D. A . Folli 1466 (K, RB, SPF, UEC). Pernambuco: unknown municipality, no date (fr.), G . Gardner s.n. (K000018779). Mun.Bonito, no date (fl.), A. G . Silva 545 (PEUFR). Mun.Cabo de Santo Agostinho, 29 January 1996 (st.), D. R . Siqueira 140 (PEUFR). Mun. São Vicente Férrer, 30 October 1998 (fl.), E. M. N . Ferraz 494 (PEUFR, RB). Mun. Sirinhaém, 1 November 1968 (fl.), D. P . Lima 12620 (IPA). Rio de Janeiro: unknown municipality, 25 February 1826 (fl.), W. J . Burchell 2605 (K). Mun. Duque de Caxias, 11 February 1939 (fr.), B . Lutz? (R). Mun. Guapimirim, 25 March 1992 (fr.), S. V. A . Pessoa 613 (BHCB, RB). Mun. Magé, 1883 (fl.), M . Palma s.n. (R139896). Mun. Petrópolis, February 1938 (fl.), B . Lutz 1070 (R); ibidem, February 1938 (fl.), B . Lutz 1054 (R); ibidem, no date (fr.), D.Sucre 10760 (RB); ibidem, 5 January 1950 (fl.), L. E . Mello-Filho 989 (R); ibidem, 5 January 1950 (fl.), L. E . Mello-Filho 994 (R) .</p></div>	https://treatment.plazi.org/id/03B187B9FFE9FFB400FEFB38A5FBFFDD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFE6FFBF00FEF89CA17DFAF0.text	03B187B9FFE6FFBF00FEF89CA17DFAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia maculata M. F. Santos	<div><p>8. Myrcia maculata M.F.Santos (in Santos et al. 2019: 4) (Figure 24, 25).</p><p>≡ Eugeniopsis luschnathiana O. Berg (1857 –1859: 147). Type: — Brazil, Bahia, mun. Ilhéus, 1839 (fl.), Luschnath s.n. (but cited as Luschnath 7 in the protologue) (lectotype BR! [845562] designated by Santos et al. 2019; isolectotypes B†, BR! [845491], BR! [845593], BR! [845494], BR! [845524], BR! [845527], BR! [845529], BR! [845560], BR! [845595], LE! [LE00004007]).</p><p>Description:— Tree (?). Trichome reddish, simple or dibrachiate, ca. 0.1 mm. Twig when mature stramineous (when dry), cylindrical, glabrous, cortex slightly cracked with numerous pellucid dots; branching monopodial, internode 1.6– 3.4 cm, node slender; cataphylls rarely present, deciduous; buds pubescent. Leaf concolorous, chartaceous, apparently slightly corrugate, blade 5.2–9.6 × 1.9–4.0 cm, elliptic, apex acute, base cuneate, margin plane or slightly revolute, secondary veins 3–5 mm apart, held at an angle of 55° relative to the midvein, marginal vein 2.0 mm from the margin, tertiary veins inconspicuous in both surfaces, areoles ca. 2 mm, pellucid dots conspicuous in the abaxial surface, ca. 10 per mm 2; adaxial surface glabrous, secondary veins plane; abaxial surface glabrous, secondary veins raised; petiole 2–6 × 2 mm, semicylindrical, glabrous. Inflorescence 7.0–8.0 × 4.5–6.0 cm, ca. 40 flowers, axillar at the terminal or subterminal nodes of the SGU (central bud developing a vegetative branch), one inflorescence per axillary bud, inflorescence pherophyll persistent or deciduous, opposite branching, three branching per node, rachis puberulent, first internode of central rachis 2 mm wide; bracts deciduous, not seen; bracteoles deciduous, not seen. Flower bud ca. 2 × 1.2 mm, clavate (but slightly flat at the apex), corolla hidden or barely apparent before anthesis; hypanthium 1.2 mm extending above the summit of the ovary, not tearing at anthesis or with a small vertical rip (but not tearing the staminal ring), externally pubescent to puberulent, pellucid dots covering the whole surface; calyx 4–merous, lobes 0.8–1.2 × 1.0–2.0 mm, not fused, deciduous, depressed ovate to widely depressed ovate, concave, apex rounded, externally puberulent to glabrous, internally puberulent; corolla 4–merous, 1.2–1.7 × 1.2–1.7 mm, widely depressed ovate, widely ovate to very widely ovate or very widely obovate, plane, apex rounded, externally puberulent, internally puberulent to glabrous; staminal ring 0.2–0.5 mm wide, subglabrate to glabrous, stamens ca. 75, filament 2.0– 2.8 mm long, anther 0.24 × 0.24–0.32 mm, gland at the apex; ovary 1.0 × 1.0– 1.5 cm, obconic, style ca. 2.6 mm long, glabrous, stigma punctiform. Fruit and seed not seen.</p><p>Distribution and Habitat: — Myrcia maculata is known from only a few collections in the Ilhéus region of Bahia state, Brazil (Figure 25). Although habitat is not explicitly recorded, it is likely to be either lowland rainforest or restinga vegetation.</p><p>Phenology: —The flowering period remains unknown due to lack of data from the type collection, and no fruiting specimen have been documented.</p><p>Conservation Status: — Myrcia maculata has only three records, one of which lack a precise location. Due to limited data available, the species is classified as Data Deficient (DD; IUCN 2022).</p><p>Taxonomy: — Myrcia maculata is a rare and poorly understood species, making its precise circumscription challenging. The type specimen displays distinctive characteristics, including stramineous branches with numerous pellucid dots and slightly corrugated leaf with occasionally revolute margins. However, it is unclear to what extent these traits may have been affected by the herborization process. Additional features include monopodial branching, elliptic leaf blades, axillar inflorescence at terminal or subterminal nodes of the SGU, and clavate flower bud with flattened apex.</p><p>Illustrations and images in previous works: —None.</p><p>Additional specimens examined: — BRAZIL. Unknown province/state, no date (fl.), unknown collector s.n. (M-0171096); no date (fl.), unknown collector s.n. (M-0171097); no date (fl.), unknown collector s.n. (M-0171098).</p><p>Bahia: unknown municipality, no date (fl.), C. F. P.Martius s.n. (P 05131554); ibidem,1841 (fl.), B.Luschnath s.n. (NY 615572).</p></div>	https://treatment.plazi.org/id/03B187B9FFE6FFBF00FEF89CA17DFAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
03B187B9FFF2FFD200FEFF5CA53CF92D.text	03B187B9FFF2FFD200FEFF5CA53CF92D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myrcia tenondeporan M. F. Santos & E. Barretto 2023	<div><p>13. Myrcia tenondeporan M.F.Santos &amp; E.Barretto (2023: 70) . Type: — BRAZIL. São Paulo: Mun. São Paulo, Marsilac, Sítio Maravilha, Mata secundária, Rio Capivari, 23 K 324844 7350922, 12 November 2014 (fl.), Barretto 338 (holotype PMSP!; isotypes HUFABC!, RB!) (Figures 20, 32E–H, 36 and 37).</p><p>Description:— Shrub to tree, 1 to 9 m high. Trichome reddish, simple or dibrachiate, 0.08–0.32 mm. Twig when young flattened, pubescent to puberulent, when mature greyish (when dry), cylindrical, cortex slightly cracked; branching monopodial (often sympodial), internode 1.0–8.0 cm, node slender; cataphylls absent or present at the base of the SGU, foliaceous, deciduous; buds pubescent. Leaf concolorous, chartaceous, blade 4.8–15.8 × 1.5–6.0 cm, narrowly to widely elliptic, oblong, ovate to widely ovate, obovate to widely obovate, apex acuminate or caudate, base narrowly cuneate to rounded or attenuate, margin plane, secondary veins 1–8 mm apart, held at an angle of 50–75° relative to the midvein, marginal vein 0.5–2.0 mm from the margin, tertiary veins usually inconspicuous in the adaxial surface, conspicuous to inconspicuous in the abaxial surface, areoles 0.3–2.0 mm, pellucid dots usually conspicuous in the abaxial surface, 1–5 per mm 2; adaxial surface glabrous, secondary veins plane to raised; abaxial surface puberulent to subglabrate, secondary veins plane to raised; petiole 9–21 × 1–2 mm, canaliculate, pubescent to puberulent. Inflorescence 2.5–8.5 × 1.5–5.5 cm, 9–40 flowers, axillar at the subterminal nodes of the SGU (central bud developing a vegetative branch), usually only one pair per SGU, one inflorescence per axillary bud, inflorescence pherophyll deciduous, opposite branching, three branching per node, rachis pubescent to puberulent, first internode of central rachis 1–2 mm wide; bracts 1.0–4.0 × 0.4–1.0 mm, deciduous, lanceolate to ovate, concave, apex acuminate to acute or caudate, base truncate, externally puberulent, internally puberulent to glabrous; bracteoles 0.8–3.0 × 0.2–2.0 mm, deciduous, narrowly elliptic or lanceolate to ovate, concave or keeled, apex acuminate or caudate, base truncate, both surfaces puberulent. Flower bud 1.6–1.8(3.0) × 1.2–2.5 mm, clavate, corolla hidden or barely apparent before anthesis; hypanthium 1–1.5 mm extending above the summit of the ovary, not tearing at anthesis or often with a small vertical rip (but usually not tearing the staminal ring), externally pubescent to puberulent, pellucid dots covering the whole surface (sometimes covered by the indument); calyx 4–merous, lobes 0.4–1.5 × 1.0–1.2(2.0) mm, not fused, deciduous parallel to the hypanthium ring, depressed ovate to widely depressed ovate, concave, apex rounded, externally pubescent to puberulent, internally puberulent; corolla 4–merous, petals white, 1.1–1.2(2.0) × 1.2–1.6(2.0) mm, obovate to very widely obovate, plane to concave, apex rounded, both surfaces puberulent to glabrous; staminal ring 0.2–0.5 mm wide, puberulent, stamens 30–107, filament 2.8–7.0 mm long, anther 0.16–0.24 × 0.24–0.40 mm, gland at the apex; ovary 0.6–1.0 × 0.6–1.0 mm, obconic, style 4.4–8.0 mm long, glabrous, stigma punctiform (often slightly swollen). Fruit immature green, mature black, 1.2–1.8 × 1.4–1.8 cm, globose, base rounded, puberulent to glabrous, totally covered by pellucid dots, remnants of hypanthium and calyx present; seeds 1 or 2 per fruit.</p><p>Distribution and Habitat: — Myrcia tenondeporan is distributed within the Atlantic Forest domain along the Serra do mar escarpments, ranging from northern Santa Catarina to southern Rio de Janeiro states (Figure 20). It grows in rainforest, riparian forest and restinga, at elevations from 0 to 900 meters. This species generally inhabits the undercanopy but has also been recorded at forest edges and riparian zones (Santos &amp; Barretto 2023).</p><p>Phenology: —Flowering occurs from September to February, with a peak from November to December. Fruiting has been recorded throughout the year, though mature fruits were found only from August to December.</p><p>Conservation Status: —According to Santos &amp; Barretto (2023), based on the Extent of Occurrence (33,533 km 2) and the Area of Occupancy (276 km 2), the species is preliminarily considered as Vulnerable (VU, criteria B2a, bii, biii; IUCN 2022).</p><p>Taxonomy: — Myrcia tenondeporan is distinctively characterized by its monopodial (often sympodial) branching, leaves with elongated (acuminate or caudate) apices, inflorescence typically paired at the subterminal node of the SGU with deciduous pherophyll, and small flower bud with a constrict apex (Figure 32E–H). Specimens of M. tenondeporan were previously often identified as M. vellozoi, but M. tenondeporan can be distinguished by its smaller tree size, more elongated leaf apices, and smaller floral bud (Santos &amp; Barretto 2023). Myrcia tenondeporan also shares similarities with Myrcia eugeniopsoides in leaf blade shape and size as well as much of its range, but M. eugeniopsoides has a closed floral bud with indistinct calyx lobes, which open irregularly during anthesis (Santos &amp; Barretto 2023). Myrcia tenondeporan is also similar to Myrcia clausseniana but differs by the distribution, the inflorescence with usually only one pair per seasonal growth unit, and the bigger mature fruit. Vernacular names include “guamirim-branco” in São Paulo state and “guamirim” or “guamirim-mamão” in Paraná state.</p><p>Illustrations and images in previous works: — Santos &amp; Barretto (2023).</p><p>Additional specimens examined: — BRAZIL. Unknown province /state, no date (fl.), W. J . Burchell 3662 (K, P). Paraná: Mun. Guaraqueçaba, 20 November 1974 (fl.), G . Hatschbach 35507 (MBM, UEC); ibidem, May 2003 (fr.), L . D.Meireles 1420 (UEC). Mun. Guaratuba, 12 December 1993 (fl.), J. M . Silva 1287 (HUEFS, MBM); ibidem, 8 November 2003 (fl.), E. J . Lucas 190 (K, MBM). Mun. Matinhos, 17 May 2003 (fr.), J . Sonehara 3 (BHCB). Mun. Paranaguá, 21 November 1967 (fl.), G . Hatschbach 17896 (MBM, NY, P, US); ibidem, 24 November 1962 (fl.), G . Hatschbach 10169 (K, MBM, SPSF); ibidem, 25 November 1967 (fl.), G . Hatschbach 17969 (MBM); ibidem, 29 May 1966 (fr.), G . Hatschbach 14393 (MBM); ibidem, 6 June 1995 (fr.), S. R . Ziller 756 (SPSF); ibidem, 6 June 1995 (fr.), S. R . Ziller 773 (ESA); ibidem, 8 November 1986 (fl.), R. M . Britez 24578 (UEC); ibidem, 17 April 1987 (fr.), S. M . Silva 24577 (UEC); ibidem, 5 December 1992 (fl.), S. M . Silva s.n. (UEC75574). Mun. Pontal do Paraná, 26 November 2012 (fl.), R. A . Bonaldi 611 (MBM). Rio de Janeiro: Mun. Paraty, 16 May 1995 (fr.), R . Marquete 2144 (RB); ibidem, 23 November 1990 (fl.), C . Farney 2517 (RB); ibidem, 8 September 2013 (fr.), A. G . Silva 3 (SPSF); ibidem, 12 April 1994 (fr.), R . Marquete 1578 (RB, UEC). Santa Catarina: unknown municipality, 8 May 2004 (fr.), K . Bourscheio 66 (BHCB); 25 May 2004 (fr.), Equipe Acaraú 96 (BHCB). Mun. Garuva, 4 November 2004 (fl.), A. C . Cervi 8725 (MBM). Mun. Itapoá, 6 November 1992 (fl.), R . Negrelle 542 (CRI). São Paulo: unknown municipality, 16 November 1826 (fl.), W. J . Burchell 3649 (NY, US). Mun. Bertioga, 10 February 2000 (fl.), P. S. P . Sampaio 435 (SP); ibidem, 11 March 1999 (fr.), S. E . Martins 394 (SP, SPSF); ibidem, 12 August 1999 (fr.), P. S. P . Sampaio 315 (SP); ibidem, 14 December 1972 (fl.), G . Gottsberger 18-141272 (MBM, SP); ibidem, 14 January 1999 (fl.), M. A. G . Magenta 77 (SP); ibidem, 19 January 1999 (fl.), E. A . Anunciação 679 (SP, SPSF); ibidem, 22 April 1999 (fr.), P. S. P . Sampaio 247 (SP); ibidem, 22 June 1999 (fr.), S. E . Martins 501 (SP); ibidem, 26 August 1999 (fr.), L . Rossi 2056 (SP); ibidem, 27 November 1989 (fl.), M. T . Grombone 22449 (UEC); ibidem, 28 November 1989 (fl.), L. T . Silveira 22514 A (UEC); ibidem, 29 May 1990 (fr.), M . Kirizawa 2291 (SP, UEC); ibidem, 29 November 1989 (fl.), M. T . Grombone 22862 (UEC); ibidem, 6 May 1999 (fr.), S. E . Martins 466 (SP). Mun. Cananéia, 9 June 2012 (fr.), V. G . Staggemeier 809 (K); ibidem, 5 November 1979 (fl.), D. A . Grande 214 (SP); ibidem, 6 July 1989 (fr.), F . Barros 1711 (SP); ibidem, 28 November 1974 (fl.), J . Mattos 16174 (SP, SPF); ibidem, 10 November 1977 (fl.), D. A . Grande 13 (SP, SPSF); ibidem, no date (fl.), D.Sampaio 117 (ESA, RB, SPSF); ibidem, 30 October 1985 (fl.), M . Kirizawa 1520 (SP); ibidem, 9 June 2012 (fr.), V. G . Staggemeier 828 (K); ibidem, 16 December 2006 (fl.), V. G . Staggemeier 98 (MBM). Mun. Cubatão, 8 June 2010 (fr.), M. F . Santos 580 (SPF); ibidem, 14 December 1988 (fl.), O. T . Aguiar 292 (SPSF); ibidem, 29 November 1988 (fl.), R . Esteves 1 (SPSF); ibidem, 21 December 1994 (fl.), S. A. C . Chiea 824 (SP, SPF). Mun. Iguape, 11 December 1985 (fl.), E. L. M . Catharino 538 (ESA, SP); ibidem, 30 March 1986 (fr.), E. L. M . Catharino 779 (ESA, SP); ibidem, 8 November 1986 (fl.), C. B. J . Jaramillo 4 (ESA, SP); ibidem, 23 June 1992 (fr.), D. F . Pereira 179 (SP); ibidem, 13 June 1991 (fr.), E. A . Anunciação 75 (SP); ibidem, 15 December 1991 (fl.), E. A . Anunciação 118 (SP, SPF); ibidem, 29 August 1993 (fr.), E. A . Anunciação 357 (MBM, SP); ibidem, 14 December 1993 (fl.), E. A . Anunciação 434 (MBM, SP); ibidem, 11 December 1990 (fl.), I . Cordeiro 779 (MBM, SP); ibidem, 8 December 1994 (fl.), I . Cordeiro 1489 (SP); ibidem, 14 May 1991 (fr.), L . Rossi 869 (SP); ibidem, 12 June 1991 (fr.), L . Rossi 904 (SP); ibidem, 17 December 1991 (fl.), L . Rossi 1026 (SP); ibidem, 13 March 1990 (fr.), M. C. H . Mamede 225 (SP); ibidem, 21 June 1990 (fr.), M. C. H . Mamede 292 (SP); ibidem, 27 June 1993 (fr.), M. C. H . Mamede 542 (NY, SP); ibidem, 25 April 1991 (fr.), M. M. R. F . Melo 954 (SP); ibidem, 7 January 1999 (fr.), M. R . Gorenstein 51 (ESA, RB, SPSF, UEC); ibidem, 16 January 1983 (fl.), N . Figueiredo 14400 (UEC); ibidem, 16 August 1990 (fr.), S. A. C . Chiea 573 (SP); ibidem, 12 December 1990 (fl.), S. J. G . Silva 112 (SP); ibidem, 12 December 1990 (fl.), S. J. G . Silva 120 (SP). Mun. Itanhaém, 7 July 2001 (fr.), N. M . Ivanauskas 4490 (ESA); ibidem, 17 April 2001 (fr.), F. M . Souza 197 (ESA, RB); ibidem, 17 April 2001 (fr.), F. M . Souza 205 (ESA, SPSF, UEC); ibidem, 29 March 2005 (fr.), F. M . Souza 592 (BHCB, SPSF); ibidem, 14 March 2005 (fr.), R . Cielo-Filho 383 (MBM, SPSF). Mun. Pariquera-Açu, 29 September 1998 (fl.), M . Sztutman 43 (ESA); ibidem, 23 November 1998 (fl.), M . Sztutman 79 (ESA, RB). Mun. Peruíbe, 11 October 1987 (fl.), G . Hashimoto 17152 (SP); ibidem, 9 October 1995 (fl.), V. C . Souza 9276 (ESA, MBM, RB, SP); ibidem, 21 June 1994 (fr.), I . Cordeiro 1505 (SP, UEC); ibidem, 6 November 1988 (fl.), V. C . Souza 341 (ESA). Mun. Praia Grande, 25 October 2007 (fl.), A . Lobão 1441 (RB). Mun. Salesópolis, 17 March 1958 (fr.), M . Kuhlmann 4351 (SP). Mun. Santo André, 13 December 2007 (fl.), R. J . Almeida-Scabbia 5219 (P, SP); ibidem, 5 June 2008 (fr.), F . Búgola-Silva 8201 (NY, P, SP); ibidem, 16 January 1920 (fl.), F. C . Hoehne s.n. (SP3607, SPF, US); ibidem, 6 December 1961 (fl.), J . Mattos 8682 (SP, SPF, US); ibidem, 30 August 1961 (fr.), J . Mattos 9093 (SP, SPF, US); ibidem, 6 August 1961 (fr.), J. R . Mattos s.n. (SP64710); ibidem, 5 November 1991 (fr.), M . Kirizawa 2534 (SP); ibidem, 20 November 1984 (fl.), M . Sugiyama 566 (SP); ibidem, 30 September 1988 (fr.), M . Sugiyama 787 (SP). Mun. Santos,1875 (fr.), [Mosiu] 2783 (P); ibidem, 22-23 October 1826 (fl.), W. J . Burchell 3266 (K); ibidem, 3 May 1994 (fr.), M. M. R. F . Melo 1009 (BHCB, SP). Mun. São Bernardo do Campo, 22 August 1990 (fr.), S . Ferreira s.n. (SP271139). Mun. São Paulo, September 1994 (fr.), I . Cordeiro 1642 (SP); ibidem, 20 October 1998 (fl.), A . Furlan s.n. (BHCB84697); ibidem, 22 June 1995 (fr.), S. A. P . Godoy 612 (MBM, SP); ibidem, 2 May 1996 (fr.), G. M. P . Ferreira 86 (SP); ibidem, 28 January 1999 (fr.), L. C. Q. M. P . Sampaio 149 (MBM, SPF, SPSF); ibidem, 13 October 2016 (fl.), M. F . Santos 872 (SORO); ibidem, 28 November 1980 (fl.), S. L . Jung 369 (MBM, SP). Mun. São Vicente, 21 November 2003 (fl.), C . Moura 43 (SPSF); ibidem, 16 February 2001 (fr.), J. A . Pastore 958 (SP, SPSF). Mun. Ubatuba, 12 January 1993 (fl.), M. A . Assis 71 (UEC); ibidem, 13 November 1993 (fl.), A. L. M . Franco 29358 (SP, SPF, UEC); ibidem, 17 November 1999 (fl.), D. C . Zappi 311 (SP, UEC); ibidem, 21 June 1996 (fr.), S. L. R . Castro 49 (SP, SPSF); ibidem, 6 November 1988 (fl.), A . Furlan 596 (SPSF, UEC); ibidem, 7 November 1988 (fl.), A . Furlan 639 (SPSF, UEC) .</p></div>	https://treatment.plazi.org/id/03B187B9FFF2FFD200FEFF5CA53CF92D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Santos, Matheus F.	Santos, Matheus F. (2025): Taxonomic monograph of Myrcia sect. Eugeniopsis (Myrciinae, Myrteae, Myrtaceae), an endemic clade of Eastern South America. Phytotaxa 703 (1): 1-100, DOI: 10.11646/phytotaxa.703.1.1, URL: https://doi.org/10.11646/phytotaxa.703.1.1
