identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03BAF97BFFC1FF90FF46F97EC31F7C92.text	03BAF97BFFC1FF90FF46F97EC31F7C92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ochodaeus montanus Fuente Hind 1912	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ochodaeus montanus Fuente, 1912 (Fig. 1–5, 6–8, 16, 17, 20, 23, 25, 28, 34) </p>
            <p> Type material.  Ochodaeus montanus Fuente, 1912 . NEOTYPE ♀ [erroneously considered a male!], designated by Alonso-Zarazaga and López-Colón (1996) (MNCN). </p>
            <p> Labels: (a) white rectangular paper, handwritten: Molinicos, 29.V.1938; (b) white label, rectangular, handwritten:  Ochodaeus /  pocadioides Mots. , ♂ / J.J. L. Colón. det., 1980; (c) white label, rectangular, framed in black, handwritten and printed:  Ochodaeus /  montanus / Fuente, 1912 / F.M. Piera det. 1988; (d) red label, rectangular, handwritten:  Ochodaeus montanus / FUENTE, 1912 / M.A. Alonso Zarazaga / &amp; J.I. López-Colón des. 97 / – </p>
            <p>NEOTYPUS –; (e) red label, rectangular, framed in black, printed: MNCN / Cat. Tipos Nº / 8380; (f) white label,</p>
            <p>rectangular, framed in black, printed: MNCN_Ent / 2957; (g) white label, rectangular, handwritten and printed:</p>
            <p> Ochodaeus /  montanus Fuente ♀ / J. B Huchet det. 2024. Other specimens (3 ex.): </p>
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                  SPAIN: ALBACETE:  River Guadalmena (Villapalacios), UTM: 30SWH 2871, 730m., 18-V-2003, attracted by artificial light in a meadow on the riverbank, in a holm oak forest environment with  Quercus ilex L.  ssp. ballota (Desf.) Samp., J.L. Lencina &amp; F. Lencina leg. (1 ♂ / CJLL)  ;   JAEN: Sierra de Cazorla: Santiago-Pontones,  
                <a title="Search Plazi for locations around (long -2.852325/lat 38.04569)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.852325&amp;materialsCitation.latitude=38.04569">Coto Ríos</a>
                 , 38.045688 -2.852325 (650 mts), Light trap,  
                <a title="Search Plazi for locations around (long -2.852325/lat 38.04569)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-2.852325&amp;materialsCitation.latitude=38.04569">Ribera</a>
                 , 7.VI.2021, F.J. Campillo leg. (1 ♀ / CJBH)  .   MURCIA: Collado de las Cabellas,  Sierra de Villafuerte (Moratalla), UTM: 30SWH 7422, 1650 m., 9-VI to 8-VII-2017, in a clearing of pine forest  Pinus nigra Arnold , in pitfall trap, J.L. Lencina &amp; J. Miñano leg. (1 ♂ / CJLL)  . 
            </p>
            <p> To the best of our knowledge, besides these four specimens, no other specimens of  O. montanus are known, highlighting the extreme uncommonness of this species. Additionally, three specimens previously identified as  O. montanus in the collections of the Museum of Madrid, as listed below (MNCN _Ent 2959, 211587, and 211588), were found to correspond to  O. inermis . </p>
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	https://treatment.plazi.org/id/03BAF97BFFC1FF90FF46F97EC31F7C92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gutierrez, Jean-Bernard Huchet José Luis Lencina	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
03BAF97BFFC2FF9DFF46FD1AC3EE781A.text	03BAF97BFFC2FF9DFF46FD1AC3EE781A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ochodaeus inermis Reitter, J. B Huchet 1893	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Ochodaeus inermis Reitter, 1893 (77 ex.) (Fig. 9, 10, 11-15, 18, 19, 22, 24, 26, 29) </p>
            <p> SPAIN: ALBACETE: Molinicos, 29.5.1938. [Escalera], MNCN_Ent 2959,  Ochodaeus pocadioides Mots. J.I.L. Colón det., 1980,  Ochodaeus montanus Fuente, 1912 , F.M. Piera det. 1988,  Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♀ / MNCN); Albacete: Molinicos, without data [Escalera], MNCN_Ent 211587,  Ochodaeus pocadioides Mots. J.I.L. Colón det., 1980,  Ochodaeus montanus Fuente, 1912 , F.M. Piera det. 1988 [erroneously considered a male!],  Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♀ / MNCN); Molinicos, 17.V.2022, F. Lencina and F. Albert leg. (1 ex. / CJLL); Ayna, 20.VI.2003, J.L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); same data and collectors but 27.VIII.2003 (2 ex. / CJLL); Férez, 13.V.2006, E. Aistleitner leg. (1 ex. / CJLL); Hellín, 30.V.2012, J.L. Lencina leg. (1 ex. / CJLL); same data but 30.V.2009, J.L. Lencina et al. leg. (1 ex. / CJLL); same data but 28.VIII.2007, J.L. Lencina and F. Lencina leg. (6 ex. / CJLL); same data but 18.V.2007, J.L. Lencina and F. Lencina leg. (1 ex. / CJLL); same data but 19.IV.2007, J.L. Lencina, F. Albert, and F. Lencina leg. (5 ex. / CJLL); E. Hellin, Saladar de Cordovilla, 18.V.2007, J.L Lencina leg.,  Ochodaeus inermis Reitter, 1893 , det. J.L Lencina 2007 (2 ex. / CJBH); E. Hellin, Embalse de Camarillas, 19.IV.2007, luz, J.L Lencina leg. (1 ex. / CJBH); E. Tobarra, Saladar de Cordovilla, 11.V.2007, J.L Lencina leg.,  Ochodaeus inermis Reitter, 1893 , det. J.L Lencina 2007 (1 ♂ / CJBH); Letúr, 02.II.2008, J.L.Lencina and A. Andújar leg., berlese (1 ex. / CJLL); Motilleja, 26.VI.2004, J.L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); Nerpio, 11.IX.2004, J.L. Lencina, F. Albert, and F. Lencina leg. (2 ex. / CJLL); Tobarra, 01.V.2004, J.L. Lencina, F. Albert, and F.Lencina leg. (2 ex. / CJLL); same data but 11.V.1999, J.L. Lencina and F. Lencina leg. (7 ex. / CJLL); same data but 11.V.2007, J.L. Lencina, F. Lencina, and P. Abarca leg. (3 ex. / CJLL); same data but 28.VII.2007, J.L. Lencina leg. (2 ex. / CJLL); Villapalacios, 04.VI.2004, J.L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); Yeste, 19.VI.2004, J.L. Lencina, F. Albert, and F. Lencina leg. (1 ex. / CJLL); CÁDIZ: Jerez, without data. L. Cepero, “ codocera ” MNCN_Ent 211572,  Ochodaeus pocadioides Mots. Jerez / L. Cepero – codocera (1 ex. / MNCN); CÓRDOBA: Córdoba, without data, Mora!. Col. del Sr. Perez Arcas. MNCN_Ent 211575 (1 ex. / MNCN); CUENCA: Villalba de la Sierra, 01.VII.1989, J.L. Lencina and M.C.Muñoz leg. (1 ex. / CJLL); GRANADA: Puebla de Don Fadrique, Nablanca, 22-23 VI 2012, Z. Laštùvka leg.,  Ochodaeus inermis Reitter, 1893 , J. B Huchet det. 2017 (1 ex. / NMPC); JAÉN: Úbeda, Jaén, without data, Ceballos, MNCN_Ent 211574 (1 ex. / MNCN); Cazorla, Coto Ríos, 7.VI.2021, F. J. Campillo leg., trampa de luz, 38.045688 / -2.852325 (650 mts), Ribera (1 ♂, 1 ♀ / CJBH); Santiago-Pontones, 07.VI.2021, F.J. Campillo leg. (2 ex. / CJLL); MADRID: El Regajal, Aranjuez, 28-05-2001, M.G. París, T. Luz. MNCN_Ent 211588,  Ochodaeus pocadioides Mots. J.I.L. Colón det., 1980,  Ochodaeus montanus Fuente, 1912 , F.M. Piera det. 1988,  Ochodaeus inermis Reitter, J. B Huchet det. 2024 (1 ♂ / MNCN); Alcalá, 10.5.94. [Escalera], MNCN_Ent 2955 (1 ex. / MNCN); MURCIA: Jumilla, Monte de Santa Ana, 10.V.2013, trampa de luz, 30SXH 4754, 650 m, J.L Lencina leg.,  Ochodaeus inermis Reitter, 1893 , det. J.L Lencina 2013 (1 unsexed, 1 ♂ / CJBH); Jumilla, 29.V.2004, F. Lencina leg. (1 ex. / CJLL); same data but 10.V.2013, J.L. Lencina and F. Lencina leg. (12 ex. / CJLL); same data but 23.VIII.2016, J.L. Lencina and M.C. Muñoz leg. (1 ex. / CJLL); Moratalla, 22.VI.2013, UMU (1 ex. / CJLL); SÉVILLE: Villamanrique, 10-IV-55, J. Baraud, Muséum Paris, 1993, Coll. J. Baraud (2 ex. / MNHN); TERUEL: Teruel, without data, B. Muñoz 935 </p>
            <p> m, MNCN_Ent 211573 (1 ex. / MNCN) ;   ZARAGOZA:  Val de Boquero el Ciego, la Retuerta, 11.VI.2013, UTM 30TYL2795, R. Macía leg. (1 ex. / CJBH)  . </p>
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	https://treatment.plazi.org/id/03BAF97BFFC2FF9DFF46FD1AC3EE781A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gutierrez, Jean-Bernard Huchet José Luis Lencina	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
03BAF97BFFC9FF98FF46FB0DC4C37C75.text	03BAF97BFFC9FF98FF46FB0DC4C37C75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ochodaeus montanus Fuente Hind 1912	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Distinguishing  O. montanus from  O. inermis : new comparative discriminating characters </p>
            <p> Ochodaeus montanus can be distinguished from  O. inermis by a series of morphological characters. The most distinctive feature is the presence of a prominent tooth on the distal part of the posterior margin of the hind femora in both sexes of  O. montanus (Fig. 5), which is always absent in both sexes of  O. inermis . In  O. montanus , the clypeus is consistently convex, forming a curve that reaches the upper edge of the eye on either side (Fig. 1, 2, 28). In contrast, the clypeus of  O. inermis is weakly sinuate on either side of the middle and typically slightly flattened anteriorly (Fig. 11, 12, 29). The pronotal punctation also exhibits notable differences between the two species:  O. montanus shows a simple, very dense, and strong punctation composed of tightly packed setose squamiform granules (Fig. 1, 2, 23), whereas the punctation in  O. inermis is double and more superficial, comprising both tight setose squamiform granules and distinct scattered umbilicated punctures (this distinction is most apparent under low-angle lighting, with the use of a calque layer) (Fig. 11, 12, 24). In  O. montanus , the strial punctures on elytra are very large, resulting in narrow intervals whose width is roughly equal to 2 or 3× the diameter of the punctures constituting the striae (Fig. 25). Conversely, in  O. inermis , the striae consist of medium-sized punctures with interval widths approximately 4 to 5× the diameter of the strial punctures (Fig. 26). In ventral view, the abdomen of  O. montanus exhibits large setiferous granules along the anterior edge of the sternites, with these granules being spaced approximately 2–3× their diameter, particularly on ventrites IV-VI (Fig. 6). In contrast,  O. inermis has more reduced granules that are closer together, separated by about 1–2× their diameter (Fig. 9). Additionally, in tergal view, the shape of the stridulatory peg (plectrum) located on sternite VI of the abdomen proves to be a discriminating character in differentiating the two species (Fig. 8, 10). </p>
            <p> Color patterns: within the  Ochodaeini tribe, which includes, among others, the genera  Ochodaeus Dejean, 1821 and  Parochodaeus Nikolajev, 1995 , the coloration of the habitus is relatively variable within a single species and thus not highly discriminatory. However,  O. inermis is typically yellowish-tan to light brown, sometimes darkened, whereas the four known specimens of  O. montanus exhibit a dark reddish-brown coloration. </p>
            <p>Male genitalia</p>
            <p> Ochodaeus montanus : The phallobase is elongated and dorsally curved, tapering to a point at the apex. The parameres are short, symmetrical, and rounded apically. The internal sac contains a single, large, boomerang-shaped endophallite (Fig. 16). The genital segment (urite IX) is elongated, with the base being nearly straight (Fig. 17). </p>
            <p> Ochodaeus inermis : The phallobase resembles that of the previous species. The parameres are short, symmetrical, and only slightly pointed at the apex. The internal sac contains two endophallites: one medium-sized, arched, and serrated along its convex surface, and another smaller one located beside it. The surface of the internal sac is partially covered with raspulae and fine spicules (Fig. 18). The genital segment (urite IX) is relatively wide and transverse, with the base distinctly sinuate in the middle (Fig. 19). </p>
            <p>Female terminalia</p>
            <p> Ochodaeus montanus (Fig. 20): The segment is complete. Tergite IX is arch-shaped, narrow, poorly sclerotized dorsally, and glabrous. Gonopod IX is trimeric, consisting of the subcoxite, coxite, and stylus. The subcoxite is roughly triangular and concave, with four to five long setae aligned at the tip of the ventral edge. The coxite is elongated, roughly cylindrical, slightly flattened at the apex, and bears 10 long setae. The style is elongated, digitiform, and bears eight thin setae apically. The trimeric gonopod IX was shown by Huchet (2021) to be a character of  Ochodaeini , in contrast to the same structure in  Nothochodaeini Nikolajev, 2015 (  Nothochodaeus Nikolajev, 2005 and  Ceratochodaeus Huchet, 2017 ) that is dimeric and lacks a terminal stylus. </p>
            <p> Ochodaeus inermis (Fig. 22): The gonopod IX is trimeric. The subcoxite is roughly triangular and concave, with five to six long setae arranged in staggered rows at the tip of the ventral edge. The coxite is elongated, roughly cylindrical, and slightly flattened at the apex, bearing 11 long setae. The style is short, digitiform, and bears five thin setae apically. </p>
            <p>Sexual dimorphism</p>
            <p> In  O. inermis , the anterior tibiae of males are significantly narrower with sharper teeth, whereas in females, the tibiae are enlarged, and the teeth are wider (Fig. 14). To our knowledge, these characteristics associated with sexual dimorphism encountered across multiple species within the genus  Ochodaeus have not been documented in the literature. Additionally, the clypeus in males is typically more markedly sinuate on either side of the midline, whereas in females, the clypeus exhibits a more smoothly curved anterior margin. </p>
            <p> In  O. montanus , we note the presence of a small tooth located on the anterior margin of male profemora. Although this character was mentioned in Baguena Corella’s (1967) diagnosis of this species and later included by Baraud (1992) in his identification key, neither of these authors had seen specimens of  O. montanus . As pointed out by López-Colón (1986: 34), Baguena Corella made a number of errors, mentioning  O. montanus as follows [translated from Spanish]: “ It differs from the preceding one by its anterior femora provided with a small tooth ...”, “ this obviously refers to the posterior femora, as indicated in the species key on the same page (page 212), an error that went unnoticed and, although evident, should be clarified to avoid confusion ”. </p>
            <p> Key to genera and species of  Ochodaeidae of the Iberian Peninsula </p>
            <p> 1. Apex of elytra dentate, interlocking with 2 tubercles on tergite VII (propygidium) margin (Fig 33). Anterior tibiae tridentate on outer margin, teeth equidistant, basal tooth well developed (Fig. 31). First metatarsomere strongly dilated in males. Size: 4–8 mm.  Parochodaeus pocadioides (Motschulsky)</p>
            <p> — Elytral apex and tergite VII without coaptation mechanism. Anterior tibiae tridentate on outer margin, basal tooth very small, located in proximal third (Fig. 32). Male first metatarsomere not dilated......................................................................... (  Ochodaeus ) 2 </p>
            <p> 2. Anterior margin of clypeus distinctly incised in the middle (Fig. 30). Size: 4.0–6.5 mm..........................................................................  O. chrysomeloides (Schrank)</p>
            <p>— Anterior margin of clypeus not incised....................................................... 3</p>
            <p> 3. Metafemur with a distinct acute tooth at the distal part of the posterior margin in both sex (Fig. 4, 5). Profemur with a small tooth along the anterior margin in male. Anterior margin of the clypeus regularly convex (Fig. 28). Elytral intervals slightly convex, their width about 2–3 × the diameter of the strial punctures (Fig. 25). Size: 4.8–5.7 mm.............................  O. montanus Fuente</p>
            <p> — Metafemur unarmed, or at most with a blunt lamina at the distal part of the posterior margin in both sex (Fig. 15). Profemur lacking a small tooth along the anterior margin in male. Anterior margin of the clypeus hardly flattened in front and slightly sinuate on each side of the middle (Fig. 29). Elytral intervals width about 4–5 × the diameter of the strial punctures (Fig. 26). Size: 4.0–5.7 mm.............................................................................  O. inermis Reitter</p>
            <p> Clave de los géneros y especies de los  Ochodaeidae de la Península Ibérica </p>
            <p> 1. Ápice de los élitros dentado, entrelazado con dos tubérculos en el margen del VII tergito (propigidio) (Fig 33). Tibias anteriores tridentadas en el margen externo, dientes equidistantes, diente basal bien desarrollado (Fig. 31). Primer metatarsómero fuertemente dilatado en los machos. Tamaño: 4–8 mm ............................................  Parochodaeus pocadioides (Motschulsky)</p>
            <p> — Ápice elitral y VII tergito sin mecanismo de coaptación. Tibias anteriores tridentadas en el margen externo, diente basal muy pequeño, situado en el tercio proximal (Fig. 32). Machos con el primer metatarsómero no dilatado................................................. (  Ochodaeus ) 2 </p>
            <p> 2. Margen anterior del clípeo claramente inciso en el centro (Fig. 30). Tamaño: 4.0–6.5 mm........................................................................  O. chrysomeloides (Schrank)</p>
            <p>— Margen anterior del clípeo no inciso........................................................ 3</p>
            <p> 3. Metafémures con un diente afilado distinto en la parte distal del margen posterior en ambos sexos (Fig. 4, 5). Profémures con un pequeño diente a lo largo del margen anterior en el macho. Margen anterior del clípeo regularmente convexo (Fig. 28). Intervalos élitrales ligeramente convexos, de una anchura de 2–3 veces el diámetro de los puntos estriales (Fig. 25). Tamaño: 4.8–5.7 mm............................................................................  O. montanus Fuente</p>
            <p> — Metafémures inermes, o a lo sumo con una lámina roma en la parte distal del margen posterior en ambos sexos (Fig. 15). Profémur carente de un pequeño diente a lo largo del margen anterior en el macho. Margen anterior del clípeo apenas aplanado en el frente y ligeramente sinuado a cada lado de la parte media (Fig. 29). Anchura de los intervalos elitrales aproximadamente 4–5 veces el diámetro de los puntos estriales (Fig. 26). Tamaño: 4.0–5.7 mm............................  O. inermis Reitter</p>
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	https://treatment.plazi.org/id/03BAF97BFFC9FF98FF46FB0DC4C37C75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Gutierrez, Jean-Bernard Huchet José Luis Lencina	Gutierrez, Jean-Bernard Huchet José Luis Lencina (2024): New insights and comparative morphology of the rare Western Palearctic species Ochodaeus montanus Fuente, 1912 and its closest relative, O. inermis Reitter, 1893 (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 2024 (79): 1-16, DOI: 10.5281/zenodo.14662556
