taxonID	type	description	language	source
03B887A8965F970DFF067A63FB69D27B.taxon	description	urn: lsid: zoobank. org: act: 2 A 8 E 2 E 4 A-FA 5 A- 4 E 45 - A 64 F-E 16 B 75 AE 8 F 56	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	materials_examined	Type material designated. Holotype ♂ MZUF: S-India, Kerala, Peryar [sic!] nat. Reserve: Kumily, alle luci 23. X. 1997 legit A. Sforzi & L. Bartolozzi (num. Mag. 2091) [printed] // Notoxus [printed] andrewesi Krek. [handwritten] det. G. Uhmann 19 [printed] 99 [handwritten] // “ La Specola „ Firenze [printed] 10935 [handwritten] [label black framed]. The correct name for the type locality is spelled as Periyar. Paratype 1 ♂ MZUF: same labels as holotype.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	etymology	Derivatio nominis. Toponymic. Named after the Yela Mala, the Tamil name for the Cardamom Hills, the type locality of this species. Noun in apposition.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	description	Measurements, holotype male, total body length 3.6 mm; head length 0.8 mm, head width across compound eyes 0.7 mm, pronotal length 1.3 mm, maximum pronotal width 0.8 mm, pronotal horn length 0.55 mm, maximum width 0.3 mm, elytral length 2.1 mm, combined maximum elytral width 1.2 mm. Male paratype is 3.8 mm long. Description. Forebody dorsally and ventrally pale rufous brown. Elytra uniformly brown with indistinct khaki metallic shine. Antenna yellowish rufous, two terminal antennomeres brown. Maxillary palpus and legs yellowish brown, tibiae shortly darkened at their articulations. Abdomen brown. Head elliptical, slightly longer than wide, moderately glossy. Frons shallowly impressed, vertex barely convex in dorsal aspect. Compound eyes large, irregularly oval in lateral view, strongly protruding from lateral outline of head, longer than converging tempora. Head base truncate. Head with dorsal punctures dense but shallow on frons, becoming sparser on vertex. Dorsal cranial setae yellowish, long and dense, not fully appressed. Several significantly longer erect tactile setae on tempora and along inner eye margin, some longer, some shorter than eye length. Antenna moderately long, slightly widened distally, extending slightly beyond base of elytra when directed posteriad. Terminal antennomere irregularly fusiform, nearly twice as long as penultimate antennomere. Pronotum slightly wider than head across eyes, slightly transverse (not counting pronotal ‘ horn’), dorsally slightly convex, laterally regularly rounded. Pronotal disc glossy, with complete narrow median longitudinal sulcus which is less prominent in the holotype compared to the paratype. Pronotal antebasal median bulge present but indistinct in dorsal and lateral view (Fig. 1 B). Pronotal punctures sparse, distributed irregularly on pronotal disc but distinctly denser medially at base, variably deep, intervening spaces generally significantly wider than puncture diameters. Pronotal setae yellowish, long and dense, not fully appressed. Several very long erect tactile setae present on lateral sides of pronotum. Antebasal sulcus with very dense whitishyellowish setae shorter than setae on pronotal disc, effectively concealing structure of sulcus but widely interrupted at median portion of pronotal base. Pronotal ‘ horn’ robust, moderately long, ~ 1.8 × as long as wide across its base, slightly narrowing towards apex, lateral margins with 3 - 5 distinctly protruding rather large lobules present for basal two-thirds of length. Apical third of pronotal ‘ horn’ continuously fringed. ‘ Horn’ crest moderately wide, distinctly raised, evenly sloping anteriorly, distinctly marked laterally by row of subcontiguous rugules, medially with same sized and dense rugules arranged into 2 – 3 irregular longitudinal rows. ‘ Horn’ area between crest and lateral edge with dense, rather large rugules, preapically flattened and with several radial notches, some of which continue into lateral fringe of horn. Scutellar shield small, subtriangular, rounded apically. Elytra elongate, about 1.7 – 1.8 × as long as wide, widest at midlength, dorsally slightly convex. Postbasal transverse impression shallow but present. Humerus broadly rounded. Apical sutural angle obtuse angulate, elytral apex truncate, with small, nearly circular pore of secretory channel at outer apical angle of each elytron. Elytral surface moderately glossy, dorsal punctures large, deep and dense, becoming smaller and shallower on apical fourth. Intervening spaces in part smooth to microscopically wrinkled, about as wide as to twice as wide as puncture diameters. Elytral setae double, moderate and long: yellowish, long, rather dense, suberect and intermixed with significantly longer, erect tactile setae. Metathoracic wing fully developed (functional). Latero-basal fovea of sternite III rounded, moderately deep along basal sulcus of sternum. Legs long and slender, simple, setae moderately densely and appressed. Basal metatarsomere slightly longer than combined length of remaining metatarsomeres. Tergite VII broadly rounded at posterior margin (Fig. 2 A). Sternite VII subtruncate at posterior margin (Fig. 2 B). Tergite VIII as in fig. 2 C – D, microscopically mamillate dorsally at posterior margin. Sternite IX short and narrow, rod-like (Fig. 2 E). Aedeagus as in fig. 2 F – H, paramere apex in lateral view with small and shortly setose preapical bulge (Fig. 2 H). Sexual dimorphism. Female is unknown.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	diagnosis	Differential diagnosis. The new species is another representative of the informal N. ravana Kejval, 2011 species-group (see Kejval 2011 for the definition) confined to the Indian Subcontinent as presently known. Notoxus yelamala sp. nov. resembles N. brunneorufus Bonadona, 1989 (India: Madhya Pradesh, Rajasthan) in the general appearance, but the lateral margin of the pronotal crest is distinctly lobulate for the basal portion (only few small basal lobules present in N. brunneorufus) and the aedeagus is very different, with a strongly widened and peculiarly shaped median lobe.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	biology_ecology	Ecology. Attracted to light.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965F970DFF067A63FB69D27B.taxon	distribution	Distribution. So far known only from Periyar National Park in the Cardamom Hills of the Western Ghats, Kerala, India and is likely endemic to the South Western Ghats moist deciduous forests ecoregion.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965C970CFF0679F0FB54D4A7.taxon	materials_examined	New material examined. 2 specimens ISEA: КЫргЫЗстан [Kyrgyzstan], В скл. Ферганского хр., дол. р. Урумбаш, 20 км ЗЮЗ п. КаЗарман, 1550 м, 41 ° 22 ’ N, 73 ° 47 ’ E Р. Ю. Дудко 26.07.2003; 5 specimens HMNH & 1 DTC: KYRGYZSTAN, Issyk-Kul region, E of Balikhci, 1628 m, 11. VII. 2011, // 42 ° 19.837 ’ N, 76 ° 13.088 ’ E, leg. Sándor Ilniczky; 1 specimen HMNH: KYRGYZSTAN, 15 km W of Ottuk, 1699 m, 13. VII. 2013, leg. S. Ilniczky; 3 specimens NME & 1 DTC: E-TÜRKEI Van-See Südufer, Resadiye 1800 m, 38 ° 41 ’ N 42 ° 50 ’ N [sic! Must be E], 21. IV. - 20. V. 2014, lg. Chr. Reuter; 1 specimen HMC: Mongolia / Owörchangai 33 km SE Baruunbayan-Ulaan 31.7.2019 1900 m 44 ° 53.695 ’ N / 101 ° 39.060 ’ E leg. H. Mühle 31; 9 specimens NME & 3 DTC: AFGHANISTAN NE of Paryian thern end of Panjshir valley to Anjuman pass 30.5. - 13.6. 2021, leg. C. Reuter; 1 specimen DTC: IRAN, Elbourz Prov., near Gachsar vill., 36 ° 6 ’ 38.86 ’’ N 51 ° 19 ’ 55.60 ’’ E, 4 – 6. vi. 2023, // leg. D. Kasatkin.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965A970BFF067A10FC1DD247.taxon	materials_examined	New material examined. 1 specimen NME & 1 DTC: C-VIETNAM, Thua Thien-Hua Prov., c. 30 km NW Hue, 80 m, 16 ° 17 ’ 59 ” N, 107 ° 20 ‘ 22 “ E, 10. V. 2019, by light river valley leg. A. Weigel; 1 specimen NME: C-VIETNAM, Thua Thien-Hua Prov., c. 30 km NW Hue, Son O Lau River, 16 ° 31 ’ 3 ” N, 07 ° 15 ‘ 36 “ E [sic!], 30 m, 11. V. 2019, leg. A. Skale, LF.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965A970BFF067A10FC1DD247.taxon	discussion	Note. First records from Vietnam since the original description.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965A970BFF0679D4FDB7D397.taxon	materials_examined	New material examined. 1 specimen NHMW: [Feschar] 21 7 49 [Iran]; 1 specimen NMW: SAUDI ARABIA: Asir, Abha, Al-Menhel 18 ° 12 ‘ N 42 ° 29 ’ E // 2214 m. 1 - 26. v. 2013. Malaise trap. H. A. Dawah; 1 specimen SAC & 1 DTC: Iran, Hormozgan province, Fin env., 3. II. 2015, 27 ° 38 ’ 34 ” N, 55 ° 53 ’ 58 ” E, 330 m, leg. S. Azadbakhsh; 1 specimen PPC: IRAN (ESFAHAN) Qamsar & Barzok prot. area, alt. 2757 m // Takht-e Jowr; peatland 33 ° 39 ’ 26.16 N 51 ° 21 ’ 19.5 E 19 - V- 2016 P. PONEL; 2 specimens MZLU & 1 DTC: IRN: Fars, 4 km E Paqalat 27.281998, 53.763504 31. iii. 2019 Moth light, wetland shore, desert Leg C. Fägerström; 1 specimen ABC: PAKISTAN 13. - 15.4. W, BALOCHISTAN Tump (90 km W Turbat) lgt. S. Becvar [the year of collecting is missing].	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965A970BFF0679D4FDB7D397.taxon	discussion	Note. First record from Pakistan.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067FC4FDD1D530.taxon	materials_examined	New material examined. 1 specimen ISEA: КЫргЫЗстан [Kyrgyzstan], В скл. Ферганского хр., дол. р. Урумбаш, 20 км ЗЮЗ п. КаЗарман, 1550 м, 41 ° 22 ’ N, 73 ° 47 ’ E Р. Ю. Дудко 26.07.2003.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067FC4FDD1D530.taxon	discussion	Note. First record from Kyrgyzstan.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067EC0FF75D613.taxon	materials_examined	New material examined. 2 specimens BMNH: NE INDIA, ASSAM, Bhalugpong 27 ° 02 ′ N 92 ° 35 ′ E, 150 m, L. Dembický leg., 26. v. - 3. vi. 2005 BMNH 2006 – 38.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067EC0FF75D613.taxon	discussion	Note. Hitherto this species was known only from the holotype collected in the vicinity of Jamari in Arunachal Pradesh.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067DE0FF73D7F3.taxon	materials_examined	New material examined. 7 specimens NME & 2 DTC: S-VIETNAM, Bin Phuoc Prov. Bu Gia Map NP, 16. - 21.4.2024, N 12 ° 11 ’ 38 ’’ E 107 ° 12 ’ 21 ’’, 450 – 600 m beaten from vegetation / UV LT, leg. R. Gerstmeier # 1.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067DE0FF73D7F3.taxon	discussion	Note. Hitherto this species was known only from the small type series collected in Cat Tien National Park in S Vietnam.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067C80FEACD0D3.taxon	materials_examined	New material examined. 6 specimens NME & 2 DTC: E-TÜRKEI Van-See Südufer, Resadiye 1800 m, 38 ° 41 ’ N 42 ° 50 ’ N [sic! Must read ‘ E’], 21. IV. - 20. V. 2014, lg. Chr. Reuter.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067C80FEACD0D3.taxon	discussion	Note. Hitherto this species was known only from the small type series also collected in the vicinity of Van Lake in E Turkey.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067BA0FED8D1B0.taxon	materials_examined	New material examined. 2 specimens DTC: IRAN, East Azerbaijan Prov., near Kandovan vill., 37.748092 N 46.286895 E, 21 – 22. v. 2017, leg. D. Kasatkin.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067BA0FED8D1B0.taxon	discussion	Note. Hitherto this species was known only from the extensive type series also collected in the Azerbaijan Province of Iran.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067940FC38D207.taxon	materials_examined	New material examined. 1 specimen NHMW: Iskere Bach [Pakistan].	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067994FDA4D33F.taxon	materials_examined	New material examined. 1 specimen MEC: Türkei, Manavgat Side, ex larva VI- 2010 leg. Manfred Egger.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B970AFF067994FDA4D33F.taxon	discussion	Note. First record from Turkey.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B9709FF0678CCFD60D56F.taxon	materials_examined	New material examined. 1 specimen NHMW: Lahor // Österr. Karakorum-Exp. 1958 (Pakistan) Dr. J. Piffi; 1 specimen NME: E — AFGHANISTAN Nuristan prov. 2900 m, Parun, pine forest 35 ° 18 ’ 35,6 ’’ N 70 ° 52 ’ 26,7 ’’ E 23. - 25.5. 2022, leg. C. Reuter.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
03B887A8965B9709FF0678CCFD60D56F.taxon	discussion	Note. First records since the original description.	en	Telnov, Dmitry (2025): First Notoxus Geoffroy, 1762 (Coleoptera: Anthicidae) from the Cardamom Hills of the Western Ghats with new faunistic records. Zootaxa 5665 (1): 144-150, DOI: 10.11646/zootaxa.5665.1.11, URL: https://doi.org/10.11646/zootaxa.5665.1.11
