identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03B8834CFF87F62BFDABFED3A4F2FB79.text	03B8834CFF87F62BFDABFED3A4F2FB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum	<html xmlns:mods="http://www.loc.gov/mods/v3">
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        <div> KEY TO THE CARIBBEAN  PHAENONOTUM1. Eyes small, interocular distance 5.0–6.0× the width of one eye in dorsal view (Fig. 6A–C). Metaventrite very short, its total width 5.0–6.0× the length behind mesocoxae (Fig. 7A–C). Species without hind wings (apterous)....................................................................................................................................................... 2– Eyes moderately large, interocular distance 3.5–4.0× the width of one eye in dorsal view (Fig. 6D–F). Metaventrite moderately long, its total width 3.0–4.0× the length behind mesocoxae (Fig. 7D–E). Species with fully developed hind wings (macropterous)..........................................................................................4 2. Metaventrite with pubescent cavities at sides of meso-metaventral process (Fig. 7A). Elytral suture strongly elevated posteriorly; elytral punctation very coarse (Fig. 5A). Pronotum with moderately coarse punctation, without microsculpture. Eastern Cuba (Fig. 4). ..................................  P. delgadoi Deler-Hernández et al., 2013 . – Metaventrite without pubescent cavities at sides of meso-metaventral process (Fig. 7B–F). Elytral suture weakly elevated posteriorly; elytral punctation never extremely coarse. Pronotum either with distinct microsculpture or with extremely fine (nearly invisible) punctation..........................................................3 3. Body length 3.1–3.4 mm. Head and pronotum with mesh-like microsculpture and distinct punctation. Elytral punctation rather coarse, yellowish margins of elytra present in apical half only (Fig. 5B). Mesoventral process moderately wide, median portion of metaventrite with narrow highly elevated median keel (Fig. 7C). Aedeagus 0.6 mm long (Fig. 6H). Jamaica (Fig. 4). .................................................................  P. ondreji sp. nov. – Body length 2.5–2.7 mm. Head and pronotum without microsculpture, pronotal punctation extremely fine, almost invisible. Elytral punctation fine, whole lateral margins of elytra with yellowish stripe (Fig. 5C). Mesoventral process very narrow, metaventrite with slightly elevated median portion, without median keel. Aedeagus 0.35 mm long (Fig. 6I). Hispaniola (Fig. 4). .............................................  P. laterale sp. nov. 4. Median lobe of the aedeagus wide; gonopore large, situated in apical third of the median lobe. Bases of lateral struts of the median lobe weakly expanded laterally (Fig. 6L). Widespread in the Greater Antilles (Fig. 4). ......................................................................................... .. ..................  P. laevicolle complex – Median lobe of the aedeagus narrow; gonopore small, situated subapically. Bases of lateral struts of the median lobe largely expanded laterally (Fig. 6J, K).................................................................................... 5 5. Body length 3.3–4.0 mm. Bases of parameres meeting in single point (Fig. 6K). Widespread in Greater and Lesser Antilles (Fig. 4).............................................................................................  P. exstriatum (Say, 1835) – Body length 3.2– 3.3 mm. Bases of parameres widely joined together (Fig. 6J). Puerto Rico (Fig. 4). ...............................................................................................................  P. borinquenum sp. nov.</div>
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	https://treatment.plazi.org/id/03B8834CFF87F62BFDABFED3A4F2FB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
03B8834CFF87F62FFF01FAEDA2D9F88D.text	03B8834CFF87F62FFF01FAEDA2D9F88D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum borinquenum Deler-Hernández & Sýkora & Seidel & Cala-Riquelme & Fikáček 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PHAENONOTUM BORINQUENUM SP. NOV.</p>
            <p>(FIGS 4A, 5D, 6D, J, 7D)</p>
            <p>
                  Type locality: Puerto Rico, Naguabo,  
                <a title="Search Plazi for locations around (long -65.80167/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.80167&amp;materialsCitation.latitude=18.268333">El Yunque National Forest</a>
                 , 18°16.1′N 65°48.1′W, 575 m. 
            </p>
            <p>
                 Type material:   Holotype: male (NMPC): ‘ PUERTO RICO: Naguabo: El Yunque Nat. Forest, S part La Sabana recr area 5.25 km N of Río Blanco above rd. PR191, 18°16.1′N 65°48.1′W, elevation 575 m, 21.vi.2016, A. Deler-Hernández lgt., PR05/DNA isolation: MF1730, isolated by A. Deler-Hernández 2016, isolate deposited at  
                <a title="Search Plazi for locations around (long -65.80167/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.80167&amp;materialsCitation.latitude=18.268333">Department of Entomology</a>
                 ,  
                <a title="Search Plazi for locations around (long -65.80167/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.80167&amp;materialsCitation.latitude=18.268333">National Museum in Prague</a>
                 (green printed)’ (molecular voucher MF1730)  .   Paratype: one female (NMPC; associated with the holotype by cox1 sequence): Puerto Rico: Naguabo, El Yunque Nat. Forest, S part La Sabana recr area 5.6 km N of Río Blanco at rd. PR191, 18°16.1′N 65°47.6′W, 510 m a.s.l., 21.vi.–2. vii.2016, Deler-Hernández,  
                <a title="Search Plazi for locations around (long -65.793335/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.793335&amp;materialsCitation.latitude=18.268333">Fikáček</a>
                 ,  
                <a title="Search Plazi for locations around (long -65.793335/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.793335&amp;materialsCitation.latitude=18.268333">Seidel</a>
                 lgt., PR03/DNA isolation: MF1729, isolated by A. Deler-Hernández 2016, isolate deposited at  
                <a title="Search Plazi for locations around (long -65.793335/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.793335&amp;materialsCitation.latitude=18.268333">Department of Entomology</a>
                 ,  
                <a title="Search Plazi for locations around (long -65.793335/lat 18.268333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.793335&amp;materialsCitation.latitude=18.268333">National Museum in Prague</a>
                 (green printed)’ (Molecular voucher MF1729)  . 
            </p>
            <p>Description: Habitus as in Figure 5D. Body size 3.2–3.3 mm (holotype: 3.3 mm). Body elongate oval, moderately convex, elytral suture not elevated. Dorsum black (dark brown in the teneral paratype), lateral margins of pronotum and elytra without distinct paler stripe; ventral surface dark brown to black; femora and tibiae black; antennae, maxillary palpi and tarsi yellowish. Head with sparse and moderately coarse punctation, without microsculpture (except posteriorly on frons); eyes moderately large, separated by 4.0× dorsal width of one eye (Fig. 6D). Pronotum with sparse punctures slightly finer than on head, interstices without microsculpture. Elytral punctation strongly impressed, coarser than on pronotum and head; elytral interstices without microsculpture. Wings present, fully developed. Mesoventral elevation as wide as metaventral process posteriorly, narrowing anteriorly, with distinct anterior hood; metaventrite without pubescent pits on sides of metaventral process; metaventrite c. 4.0× wider than its length posterior of mesocoxae; median moderately elevated part of metaventrite narrow throughout (Fig. 7D). Aedeagus 0.5 mm long (Fig. 6J). Median lobe rather widely triangular, c. 1.8× longer along midline than wide; apex not reaching apices of parameres; gonopore moderately large, subapical; lateral struts projecting laterad. Parameres distinctly sinuate on lateral margin, slightly expanded subapically; widely meeting each other basally. Phallobase longer than wide.</p>
            <p>Etymology: The species name is a Latinized adjective derived from the Spanish version of the indigenous Taíno name of the islands of Puerto Rico.</p>
            <p> Diagnosis:  Phaenonotum borinquenum may be distinguished from other Caribbean species by the combination of moderately large eyes, moderately long metaventrite, presence of wings and morphology of the aedeagus. It is extremely similar to  P. exstriatum in its external morphology and genital morphology, and may be distinguished from it only by its smaller body size and widely meeting bases of parameres only. Despite the strong morphological similarity, it is not closely related to  P. exstriatum . </p>
            <p>D i s t r i b u t i o n: P h a e n o n o t u m b o r i n q u e n u m i s only known from the eastern part of Puerto Rico (Fig. 4A).</p>
            <p> PHAENONOTUM DELGADOI DELER-HERNÁNDEZ, CALA-RIQUELME &amp; FIKÁČEK, 2013</p>
            <p>(FIGS 4A, 5A, 6A, G, 7A)</p>
            <p>Type material examined: See Deler-Hernández et al. (2013), including one sequenced paratype from the type locality (molecular voucher MF455).</p>
            <p>
                 Additional material examined:   Cuba: Holguín Prov.: 5 spec. (NMPC): La Melba, Parque Nacional Alexander von  
                <a title="Search Plazi for locations around (long -74.82507/lat 20.43352)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.82507&amp;materialsCitation.latitude=20.43352">Humboldt</a>
                 , 20.43352°N 74.82507°W, 336 m, rainforest litter, 21.ix.2014, R. Anderson, F. Cala-Riquelme, A. Deler-Hernández lgt. (2014-002)  ;  4 spec. (NMPC): 20.45396°N 74.82342°W, 510 m, pluviselva litter, 22.ix.2014, R. Anderson, F. Cala-Riquelme, A. Deler-Hernández lgt. (2014-011) ;   19 spec. (NMPC): road out  
                <a title="Search Plazi for locations around (long -74.81844/lat 20.51524)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.81844&amp;materialsCitation.latitude=20.51524">La Melba</a>
                 , 20.51524°N 74.81844°W, 407 m, elfin forest litter, 24.ix.2014, R. Anderson, F. Cala-Riquelme, A. Deler-Hernández lgt. (2014-015)  ;   8 spec. (NMPC): road out La Melba, road out of  
                <a title="Search Plazi for locations around (long -74.83627/lat 20.59086)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.83627&amp;materialsCitation.latitude=20.59086">La Melba</a>
                 , 20.59086°N 74.83627°W, 130 m, scrub forest litter, 24.ix.2014, R. Anderson, F. Cala-Riquelme, A. Deler-Hernández lgt. (2014-016)  . 
            </p>
            <p> Published records:   Cuba: Guantánamo Prov.:  El Yunque de Baracoa (Deler-Hernández et al., 2013a)  .  Holguín Prov.: La Melba (Deler-Hernández et al., 2013a) . </p>
            <p>Redescription: Habitus as in Figure 5A. Body length 2.2–2.5 mm (holotype: 2.3 mm). Body oval, strongly convex, elytral suture distinctly elevated posteriorly. Dorsum black, lateral margins of pronotum and elytra without distinct paler stripe; ventral surface brown to dark brown; femora and tibiae reddish; antennae, maxillary palpi and tarsi yellowish. Head with sparse fine punctation, without microsculpture (except posteriorly on frons); eyes small, separated by 5.2× dorsal width of one eye (Fig. 6A). Pronotum with very sparse minute punctures much smaller than on head, interstices without microsculpture. Elytral punctation very strongly impressed, much coarser than on pronotum and head; elytral interstices without microsculpture. Wings completely absent. Mesoventral elevation as wide as metaventral process throughout, not narrowing anteriorly, with distinct anterior hood; metaventrite with a deep pubescent pit on each side of metaventral process; metaventrite c. 5.8× wider than its length posterior of mesocoxae; median moderately elevated part of metaventrite narrow anteriorly, widening posteriorly (Fig. 7A). Aedeagus 0.4 mm long (Fig. 6G). Median lobe rather narrowly triangular, c. 1.9× longer along midline than wide; apex not reaching apices of parameres; gonopore small, subapical; lateral struts simple, not expanded. Parameres indistinctly sinuate, nearly evenly arcuate on lateral margin, not expanded subapically, broadly meeting each other basally. Phallobase slightly wider than long.</p>
            <p> Diagnosis:  Phaenonotum delgadoi may be distinguished from all other  Phaenonotum known to us by the combination of extremely coarse elytral punctation, elevated elytral suture and deep pubescent pits on each side of metaventral process. See Identification Key for additional characters. </p>
            <p> Distribution:  Phaenonotum delgadoi is only known from eastern Cuba, all known localities are situated in the Nipe-Sagua-Baracoa mountain system (Fig. 4A). </p>
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	https://treatment.plazi.org/id/03B8834CFF87F62FFF01FAEDA2D9F88D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
03B8834CFF80F62DFF35F9FAA42AF8B1.text	03B8834CFF80F62DFF35F9FAA42AF8B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum exstriatum (Say 1835)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PHAENONOTUM EXSTRIATUM (SAY, 1835)</p>
            <p>(FIGS 4B, 5E, 6E, K, 7E)</p>
            <p> Type material: Neotype of  Hydrophilus exstriatus Say, 1835 designated by Smetana (1978) from southeastern states of USA (deposited in Museum of Comparative Zoology, Harvard University, Boston, USA): not examined in our study. Types of  Phaenonotum dubium Sharp, 1882 were examined by Deler-Hernández &amp; Fikáček (2016) and confirmed as being conspecific to North American specimens treated as  P. exstriatum (synonymy proposed by Smetana, 1978). </p>
            <p>
                 Material examined:   Barbados: 1 spec. (SBPC):  
                <a title="Search Plazi for locations around (long -59.571667/lat 13.183333)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-59.571667&amp;materialsCitation.latitude=13.183333">Jack-in-Box Gully</a>
                 , forest, 13°11’N 59°34.3’W, 230 m, UV light, 23.vi.2007, S &amp; J. Peck lgt. (07-25)  .  Cuba: Cienfuegos prov. :   1 spec. (NMPC):  
                <a title="Search Plazi for locations around (long -80.33289/lat 22.12682)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.33289&amp;materialsCitation.latitude=22.12682">Soledad</a>
                 , 22.12682°N 80.33289°W, 71 m, MV light, 21.v.2013, A.B.T. Smith lgt. Guantánamo prov.  :   2 spec. (NMPC): Baracoa,  
                <a title="Search Plazi for locations around (long -74.45599/lat 20.304998)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.45599&amp;materialsCitation.latitude=20.304998">Jobo Dulce</a>
                 , 20°18′18.00″N 74°27′21.60″W, 94 m, 29.x.2010, R. Correa lgt  ;   
                <a title="Search Plazi for locations around (long -74.566666/lat 20.316668)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-74.566666&amp;materialsCitation.latitude=20.316668">El Yunque</a>
                 , 3.2 km SW of campismo, 20°19′N 74°34′W, 150 m, 13.vi.2012, A. Deler-Hernández lgt. Holguín Prov.  :   1 spec. (NMPC): Mayarí, Guatemala,  
                <a title="Search Plazi for locations around (long -75.673454/lat 20.701128)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.673454&amp;materialsCitation.latitude=20.701128">Guarina Nuñez</a>
                 , 20°42′4.06″N 75°40′24.45″W, c. 27 m, 26.iii.2013, A. Deler-Hernández lgt. Granma Prov.  :   1 spec. (NMPC): Cauto Cristo, Rio Cauto,  
                <a title="Search Plazi for locations around (long -76.48399/lat 20.272999)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.48399&amp;materialsCitation.latitude=20.272999">El Sitio</a>
                 , 20°16′22.80″N 76°29′2.40″W, 135 m, 1.v.2005, L. Chaves. Santiago de Cuba prov.  :   9 spec. (NMPC): El Vivero,  
                <a title="Search Plazi for locations around (long -75.778175/lat 20.180378)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.778175&amp;materialsCitation.latitude=20.180378">1.3 km NEE of Dos Caminos</a>
                 , 20°10′49.36″N 75°46′41.43″W, c. 170 m, at light, 23.v.2013, Deler-Hernández &amp; A. Smith lgt. (1 spec: molecular voucher MF654)  ;   1 spec. (NMPC):  Daiquirí , 1–4.vi.1985, S. Bílý lgt  .   Dominica: 8 spec. (BMNH):  La Plaine , 97-67, 22.i.1889, G. A. Ramage lgt  .  Dominican Republic: 1 spec. (NMPC): Samaná ,   6.4 km N of Samaná, road to  
                <a title="Search Plazi for locations around (long -69.337166/lat 19.263)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-69.337166&amp;materialsCitation.latitude=19.263">El Valle</a>
                 , 19°15.78′N 69°20.23′W, 21 m, 5.ix.2014, Deler, Fikáček, Gimmel (DR36)  .   Grenada: 1 spec. (SBPC):  
                <a title="Search Plazi for locations around (long -61.7027/lat 12.082533)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.7027&amp;materialsCitation.latitude=12.082533">Grand Etang Forest Reserve</a>
                 , 12°04.952′N 61°42.162′W, 434 m, nursery edge forest, UV trap, 14.viii.2010, S. Peck lgt. (10–69)  ;   25 spec. (BMNH): Grand Etang (  Windward side), 1900 ft, H. H. Smith  .;   1 spec. (SBPC): Par. St. Andrew, Mirabeau,  Agric. Lab , UV trap, 23.ii.1990, R. E. Woodruff lgt  .;   34 spec. (BMNH): Mount Gay Est. (Leeward side), H. H.  Smith lgt  .;   3 spec. (BMNH): Chantilly Est. (  Windward side), H. H. Smith lgt  .;   8 spec. (BMNH): Balthazar (  Windward side), H. H. Smith lgt  .;   1 spec. (BMNH): Vendome Est. (  Leeward side), H. H. Smith lgt  .   Cayman Islands: 1 spec. (BMNH):  Grand Cayman , by freshwater lake near George Town  ., UV light trap, 4.viii.1970, Joy Farradane lgt.   Haiti: 1 spec. (BMNH):  Port au Prince. , 1.iii.1908, M. Cameron lgt. Jamaica  :   1 spec. (BMNH):  Kingston. , 16.ii.1908, M. Cameron lgt  .;   9 spec. (SBPC): Ewarton, St. Cath. Par.,  St. Clair Cave , 27.xii.1972, S &amp; J. Peck. Monserrat  :   1 spec. (BMNH):  Salem 8.ix.1975, J. Cooter lgt  .   Saint Lucia: 1 spec. (SBPC):  
                <a title="Search Plazi for locations around (long -60.94/lat 13.875)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-60.94&amp;materialsCitation.latitude=13.875">Mon Repos</a>
                 , 6.5 km W of Fox Grovelnn, 13°52.5′N 60°56.4′W, 300 m, submontane forest litter, 22.vii.2007, S &amp; J. Peck lgt. (07-76)  .   Saint Vincent and the Grenadines: 1 spec. (SBPC): Saint Vincent,  
                <a title="Search Plazi for locations around (long -61.167667/lat 13.1328335)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-61.167667&amp;materialsCitation.latitude=13.1328335">Brighton Bay Village</a>
                 , 13°07.97′N 61°10.06′W, 1 m, streamside UV trap 8.vi.2007, S &amp; J. Peck lgt. (07-10)  ;   15 spec. (BMNH):  Saint Vincent , without additional data, H. H. Smith lgt  .  Puerto Rico: 7 spec. (NMPC): Naguabo ,   
                <a title="Search Plazi for locations around (long -65.795/lat 18.246666)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-65.795&amp;materialsCitation.latitude=18.246666">El Yunque Nat. Forest</a>
                 , S part 3.45 km N of Río Blanco at rd. PR191, 18°14.8′N 65°47.7′W, 170 m, 24.vi.2016, Deler, Fikáček &amp; Seidel lgt. (1 spec.: molecular voucher MF1728)  .   Trinidad &amp; Tobago: 2 spec. (BMNH): Trinidad,  St. Augustine , 15.iv.1926, C. L. Withycombe lgt  .;   1 spec. (BMNH):  Trinidad ,  St. Margarita , 11.vi.1942. E. C. Humphries lgt  .   USA: Florida: 1 spec. (NMPC): Highlands Co.,  Venus , 4 miles W of Fish Eating Creek, 15.viii.1965, W. Suter lgt., det. A. Smetana  ;   1 spec. (NMPC): Alachua Co.,  Gainesville , black light, 19.vii.1978, F. N. Young lgt., det. A. Smetana  ;  1 spec. (NMPC): Kansas :   Douglas Co.,  
                <a title="Search Plazi for locations around (long -95.18669/lat 38.81043)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.18669&amp;materialsCitation.latitude=38.81043">Bridenthal Ecological Reserve</a>
                 , 3.2 miles N of Baldwin City, 38.81043°N 95.18669°W, 270 m, ix.2009, Eldgedge &amp; Fikáček lgt  .;   1 spec. (NMPC): Douglas Co.,  Lawrence Baker Wetlands , 27.viii.2009, Gustafson, Eldgedge &amp; Fikáček lgt  .;  1 spec. (NMPC): Delaware :   New Castle Co.,  Frenchtown Woods Natural Area , 23.v.2004, at light, A.E. Z. Short lgt. (AS-04-065) (molecular voucher MF1063)  .  Costa Rica: 1 spec. (NMPC): Guanacaste : 6.6 km from main road, roadside gravel stream running through culvert, 10°09′26.7″N 85°22′47.5″W, 50 m, 13.i.2004, Short &amp; Lebbin lgt. (AS-04-037) (molecular voucher MF1738). 
            </p>
            <p>Redescription: Habitus as in Figure 5E. Body length 3.3–4.0 mm. Body elongate oval, moderately convex, elytral suture not elevated posteriorly. Dorsum black, lateral margins of elytra with very narrow indistinct paler stripe, pronotum paler in posterolateral corners; ventral surface dark brown to black; femora and tibiae black; antennae, maxillary palpi and tarsi yellowish. Head with sparse moderately coarse punctation, without microsculpture (except posteriorly on frons); eyes moderately large, separated by 4.0× dorsal width of one eye (Fig. 6E). Pronotum with sparse fine punctures, slightly smaller than on head, interstices without microsculpture. Elytral punctation moderately impressed, coarser than on pronotum and head; elytral interstices without microsculpture. Wings present, well developed. Mesoventral elevation as wide as metaventral process posteriorly, slightly narrowing anteriorly, with distinct anterior hood; metaventrite without pubescent pits on sides of metaventral process; metaventrite c. 3.9× wider than its length posterior of mesocoxae; median moderately elevated part of metaventrite moderately wide throughout (Fig. 7E). Aedeagus 0.4 mm long (Fig. 6K). Median lobe narrowly triangular, c. 2.1× longer along midline than wide; apex reaching apices of parameres; gonopore small, subapical; lateral struts expanded laterally. Parameres distinctly sinuate on lateral margin, slightly expanded subapically. Phallobase longer than wide.</p>
            <p> Diagnosis:  Phaenonotum exstriatum may be distinguished from Caribbean species except  P. borinquenum by the combination of moderately large eyes, moderately long metaventrite, presence of wings and morphology of the aedeagus. It may be distinguished from  P. borinquenum by larger body size and bases of parameres meeting in single point only. Despite the strong morphological similarity, it is not closely related to  P. borinquenum (see Fig. 2). </p>
            <p> Distribution:  Phaenonotum exstriatum is widespread in the eastern USA and in Central America (Smetana, 1978; Deler-Hernández &amp; Fikáček, 2016) as well as in Greater and Lesser Antilles (Fig. 4B). The only record from South America is from eastern Colombia Gonzáles-Rodríguez, García-Hernández &amp; Clarkson (2017). but the species is probably widespread in northern South America as it is common in southern islands of Lesser Antilles and it also occurs in Trinidad. </p>
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	https://treatment.plazi.org/id/03B8834CFF80F62DFF35F9FAA42AF8B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
03B8834CFF9EF632FF48FF3CA533F9FE.text	03B8834CFF9EF632FF48FF3CA533F9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum laevicolle SHARP 1882	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PHAENONOTUM LAEVICOLLE SHARP, 1882 COMPLEX </p>
            <p>(FIGS 4C, 5F, 6F, L, 7F)</p>
            <p> Type material examined: Types of  P. laevicolle Sharp, 1882 , see Deler-Hernández &amp; Fikáček (2016). </p>
            <p>
                 Material examined:   Cuba: Cienfuegos Prov.: 1 male (NMPC): Río Cabagan,  
                <a title="Search Plazi for locations around (long -80.08461/lat 21.93123)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-80.08461&amp;materialsCitation.latitude=21.93123">Gruta Mengoa</a>
                 , 21.93123°N 80.08461°W, 651 m, 20.v.2014, A. Deler-Hernández lgt. (molecular voucher MF 1115)  ;   3 spec. (ZMHB):  Sierra del Escambrai , 5 km N Topes de Collantes, c. 600 m, sifting of leafs, 17.xii.2007, M. Schülke lgt. (CU7-4). Sancti Spíritus Prov.  :   9 spec. (ZMHB): Sierra del Escambrai, Topes de Collantes, c. 700 m,  Streu ,  Totholz , 17.xii.2007, M. Schülke lgt. (CU7-3). Granma Prov.  :   7 spec. (ZMHB):  Sierra Maestra , La Habanita, 35 km NE Pilón, 900–1000 m, sifted hay, 20.xii.2007, M. Schülke lgt. (CU7-8). Santiago de Cuba Prov.  :   1 spec. (NMPC):  
                <a title="Search Plazi for locations around (long -75.77333/lat 20.18)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-75.77333&amp;materialsCitation.latitude=20.18">El Vivero</a>
                 , 1.6 km E of Dos Caminos, 20°10.8′N 75°46.4′W, 150 m, 20–21.vi.2012, A. Deler-Hernández &amp; Fikáček (MF18)  .   Haiti: 3 spec. (BMNH):  Port au Prince , 1.iii.1908, M. Cameron lgt  .  Venezuela: 1 male (NMPC): Monagas small pond between Morichal Largo &amp; Temblador, 9°05 ′47.9″N 62°43′37.1″W, 29 m, 2.ii.2010, Short, García &amp; Joly lgt. (VZ10-0202-03A) (molecular voucher MF1740). 
            </p>
            <p>Redescription: (refers to the Caribbean specimens examined): Habitus as in Figure 5F. Body length 2.7– 3.1 mm. Body elongate oval, moderately convex, elytral suture not elevated posteriorly. Dorsum black, lateral margins of elytra with paler stripe reaching subapically, pronotum paler in posterolateral corners; ventral brown to dark brown; femora dark brown to reddish, tibiae reddish; antennae, maxillary palpi and tarsi yellowish. Head with sparse fine punctation, without microsculpture (except posteriorly on frons); eyes moderately large, separated by 3.6× dorsal width of one eye (Fig. 6F). Pronotum with sparse fine punctures similar to that on head, interstices without microsculpture. Elytral punctation moderately impressed, coarser than on pronotum and head; elytral interstices without microsculpture. Wings present, well developed. Mesoventral elevation slightly narrower than metaventral process posteriorly, slightly narrowing anteriorly, with distinct anterior hood; metaventrite without pubescent pits on sides of metaventral process; metaventrite c. 3.5× wider than its length posterior of mesocoxae; median moderately elevated part of metaventrite moderately wide throughout (Fig. 7F). Aedeagus 0.5 mm long (Fig. 6L). Median lobe rather triangular, c. 1.7× longer along midline than wide; apex reaching apices of parameres; gonopore large, wide, situated in apical third of median lobe; lateral struts very shortly expanded laterally. Parameres distinctly sinuate on lateral margin, strongly expanded subapically, broadly meeting each other basally. Phallobase longer than wide.</p>
            <p>Comments: Both sequenced specimens (MF1115 from Cuba and MF1740 from Venezuela) form a strongly supported clade in the molecular analysis and are evidently closely related (pairwise distance of cox1 sequences is 4.8%). Both specimens correspond with each other in external morphology and the characteristic shape of male genitalia, and only differ in body size: the Cuban specimen is smaller (2.8 mm), the Venezuelan specimen larger (3.4 mm). In this aspect, the sequenced Cuban specimen corresponds to all additional Greater Antillean specimens examined, which are also rather small (2.7–3.1 mm). Hence, it seems probable that Venezuelan specimen is not conspecific with the Greater Antillean ones, but additional material is necessary to evaluate the intraspecific genetic and morphological variability properly to decide whether the sequenced specimens represent one or two species.</p>
            <p> The external morphology, body size and the morphology of genitalia of the Caribbean specimens correspond to the types of  P. laevicolle Sharp, 1882 described from Guatemala and examined by Deler-Hernández &amp; Fikáček (2016). However, due to the absence of DNAgrade specimens of  P.laevicolle from Central America, we are unable to determine whether the Greater Antillean specimens are conspecific. For that reason, we consider all above specimens as members of the  Phaenonotum laevicolle species complex whose taxonomy needs to be addressed once more material will be available. </p>
            <p> Distribution: The Caribbean specimens of the  Phaenonotum laevicolle complex are known from central and eastern Cuba and western Hispaniola (Haiti). Outside the Caribbean, the species complex clearly occurs in southernmost Northern and Central Americas (types of  P. laevicolle ) and in northern South America [as  P. globulosum (Mulsant, 1844) from Colombia (Hansen, 1999; not examined by us) and the Venezuelan specimen sequenced by us]. The records from Argentina (types of  P. spegazziinii Bruch, 1915 not examined by us) seem doubtful (Oliva et al., 2002) and are not considered here. Based on these sources, we estimate the distribution of the species complex to be as shown in Figure 4C. </p>
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	https://treatment.plazi.org/id/03B8834CFF9EF632FF48FF3CA533F9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
03B8834CFF9EF633FC64F95CA4DFFE42.text	03B8834CFF9EF633FC64F95CA4DFFE42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum laterale Deler-Hernández & Sýkora & Seidel & Cala-Riquelme & Fikáček 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PHAENONOTUM LATERALE SP. NOV.</p>
            <p>(FIGS 4A, 5C, 6C, I, 7B)</p>
            <p>
                  Type locality: Dominican Republic, Barahona, Monumento Natural Miguel Domingo Fuerte ‘  
                <a title="Search Plazi for locations around (long -71.18916/lat 18.0985)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-71.18916&amp;materialsCitation.latitude=18.0985">Cachote’</a>
                 18°5.91′N 71°11.35′W, 1188 m  . 
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            <p> Type material:   Holotype: male (NMPC): ‘ Dominican Republic: Barahona, MN Domingo Fuerte “Cachote,” 18°5.91′N 71°11.35′W, 1188 m, 14.viii.2014, Deler, Fikáček, Gimmel DR 03/montane broadleaf cloud forest with numerous  Cyathea : sifting of thin layer of wet leaf litter and mosses’  .  Paratypes: Dominican Republic: 10 spec. (NMPC): same data as holotype (including molecular voucher MF1013) ;  25 spec. (CMN, NMPC, MNHNSD, NHMW): MN Domingo Fuerte ‘ Cachote’, 18°5.21′N 71°11.46′W, 1205 m, sparse montane cloud forest with ferns and mosses here and there, sifting of thin layer of leaf litter, 14.viii.2014, M.Fikáček lgt. (DR03a) ;  49 spec. (BMNH, DZRJ, MCZ, MNHNSD, NMPC, SBMN): MN Domingo Fuerte ‘ Cachote’, 18°4.48–5.37′N 71°11.03–11.54′W, 1188 m, 14.viii.2014, secondary montane broadleaf forest with sparse ferns and moss, sifting of leaf litter, Deler-Hernández, Fikáček &amp; Gimmel lgt. (DR04) . </p>
            <p>Description: Habitus as in Figure 5C. Body length 2.5–2.7 mm (holotype: 2.7 mm). Body oval, moderately convex, elytral suture slightly elevated posteriorly. Dorsum brown to dark brown, lateral margins of pronotum and lateral and sutural margins of elytra with distinct yellowish stripe; ventral surface brown to reddish brown; femora and tibiae reddish; antennae, maxillary palpi and tarsi yellowish. Head with sparse fine punctation, without microsculpture (except posteriorly on frons); eyes small, separated by 5.1× dorsal width of one eye (Fig. 6C). Pronotum with very sparse and very fine punctures much smaller than on head, interstices with not very distinct mesh-like microsculpture. Elytral punctation sparse and moderately impressed, similar to that on head, much coarser than on pronotum; elytral interstices without microsculpture. Wings completely absent. Mesoventral elevation extremely narrow, as wide as metaventral process throughout, not narrowing anteriorly, without distinct anterior hood; metaventrite without deep pubescent pits at sides of metaventral process; metaventrite c. 5.0× wider than its length posterior of mesocoxae; median weakly elevated part of metaventrite narrow throughout (Fig. 7B). Aedeagus 0.35 mm long (Fig. 6I). Median lobe rather narrowly triangular, c. 2.1× longer along midline than wide; apex reaching apices of parameres; gonopore small, subapical; lateral struts simple, not expanded. Parameres indistinctly sinuate on lateral margin, not expanded subapically, widely meeting each other basally. Phallobase as long as wide.</p>
            <p>Etymology: The species name refers to the conspicuous yellow stripe along lateral margins of pronotum and elytra characteristic of this species.</p>
            <p> Diagnosis:  Phaenonotum laterale may be distinguished from other Caribbean species, based on its brown-yellow coloration, obsolete pronotal punctation and extremely narrow meso-metaventral keel. See Identification Key for further diagnostic characters. </p>
            <p> Distribution:  Phaenonotum laterale sp. nov. is only known from a high altitude cloud forest region in the southwestern Dominican Republic, geologically situated on the southern paleoisland of Hispaniola (Fig.4A). </p>
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	https://treatment.plazi.org/id/03B8834CFF9EF633FC64F95CA4DFFE42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
03B8834CFF9FF630FC4DFE4FA0BDFE45.text	03B8834CFF9FF630FC4DFE4FA0BDFE45.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaenonotum ondreji Deler-Hernández & Sýkora & Seidel & Cala-Riquelme & Fikáček 2018	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> PHAENONOTUM ONDREJI SP. NOV.</p>
            <p>(FIGS 4A, 5B, 6B, H, 7C)</p>
            <p>
                  Type locality: Jamaica,  
                <a title="Search Plazi for locations around (long -76.7271/lat 18.08726)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.7271&amp;materialsCitation.latitude=18.08726">Blue Mountains National Park</a>
                 , 18°5′14.13″N 76°43′37.55″W, 1279 m  . 
            </p>
            <p>
                 Type material:   Holotype: male (NMPC): Jamaica:  
                <a title="Search Plazi for locations around (long -76.7271/lat 18.08726)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-76.7271&amp;materialsCitation.latitude=18.08726">Blue Mountain</a>
                 , 18°5′14.13″N 76°43′37.55″W, 1279 m, 14.xi.2013, F. Cala-Riquelme leg.  ’.  Paratypes: Jamaica: 19 spec. (CMN, MCZ, NMPC, NHMW) same data as holotype (including molecular voucher MF980) ;  2 spec. (BMNH): Newcastle, 19–22.viii.1908, M. Cameron lgt . 
            </p>
            <p>Description: Habitus as in Figure 5B. Body length 3.2–3.4 mm (holotype: 3.1 mm). Body oval, highly convex, elytral suture slightly elevated posteriorly. Dorsum dark brown to black, posterolateral corners of pronotum and apical part of lateral margins of elytra distinctly yellowish; ventral surface brown to reddish brown; femora and tibiae yellowish to reddish; antennae, maxillary palpi and tarsi yellowish. Head with sparse moderately coarse punctation, with mesh-like microsculpture; eyes small, separated by 6.9× dorsal width of one eye (Fig. 6B). Pronotum with sparse and fine punctures, smaller than on head, interstices with distinct mesh-like microsculpture. Elytral punctation sparse, moderately impressed, coarser than on head and pronotum; elytral interstices without microsculpture. Wings completely absent. Mesoventral elevation very narrow, as wide as metaventral process throughout, not narrowing anteriorly, without distinct anterior hood; metaventrite without deep pubescent pits at sides of metaventral process; metaventrite c. 5.1× wider than its length posterior of mesocoxae; median highly elevated portion narrow throughout (Fig. 7C). Aedeagus 0.6 mm long (Fig. 6H). Median lobe very narrowly triangular, c. 2.3× longer along midline than wide; apex slightly overlapping apices of parameres; gonopore small, subapical; lateral struts weakly expanded. Parameres sinuate on lateral margin, weakly expanded subapically, narrowly meeting each other basally. Phallobase as long as wide.</p>
            <p>Etymology: This species is named in honour of Ondřej Jelínek, a good friend of the senior author, in appreciation of his friendship.</p>
            <p> Diagnosis:  Phaenonotum ondreji may be distinguished from other Caribbean species by the combination of small eyes, head and pronotum with distinct mesh-like microsculpture, and narrow highly elevated meso-metaventral keel. See Identification Key for additional diagnostic characters. </p>
            <p> Distribution:  Phaenonotum ondreji sp. nov. is only known from the eastern part of the island (Blue Mountains range, Fig. 4A). </p>
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	https://treatment.plazi.org/id/03B8834CFF9FF630FC4DFE4FA0BDFE45	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Deler-Hernández, Albert;Sýkora, Vít;Seidel, Matthias;Cala-Riquelme, Franklyn;Fikáček, Martin	Deler-Hernández, Albert, Sýkora, Vít, Seidel, Matthias, Cala-Riquelme, Franklyn, Fikáček, Martin (2018): Multiple origins of the Phaenonotum beetles in the Greater Antilles (Coleoptera: Hydrophilidae): phylogeny, biogeography and systematics. Zoological Journal of the Linnean Society 183: 97-120
