taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453375	https://doi.org/10.3897/vz.75.e149399.figure2	Figure 2. General skull morphology of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female, 206 mm total length). 3 D reconstruction of the skull is shown with colored rendering in dorsal (A), ventral (B), anterior (C), posterior (D) and left lateral (E) views, and of the left lower jaw in labial (F) and lingual (G) views. Scale bar equals 3 mm. Abbreviations: bas – os basale; car – foramen for the carotid artery; ch – choana; corb – circumorbital; fm – foramen magnum; front – frontal; jf – jugular foramen; ltf – lower temporal fossa; mpc - mediopalatinal cavity; mxpal – maxillopalatine; nas – nasal; nos – nostrils; oc – occipital condyles; orb + tc – orbit and tentacular canal; par – parietal; pmx – premaxilla; psa – pseudoangular; prfr – prefrontal; psd – pseudodentary; pt – pterygoid; q – quadrate; smx – septomaxilla; sphen – sphenethmoid; st – stapes; sq – squamosal; utf – upper temporal fossa; vom – vomer; vf – vomeral foramen.	Figure 2. General skull morphology of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female, 206 mm total length). 3 D reconstruction of the skull is shown with colored rendering in dorsal (A), ventral (B), anterior (C), posterior (D) and left lateral (E) views, and of the left lower jaw in labial (F) and lingual (G) views. Scale bar equals 3 mm. Abbreviations: bas – os basale; car – foramen for the carotid artery; ch – choana; corb – circumorbital; fm – foramen magnum; front – frontal; jf – jugular foramen; ltf – lower temporal fossa; mpc - mediopalatinal cavity; mxpal – maxillopalatine; nas – nasal; nos – nostrils; oc – occipital condyles; orb + tc – orbit and tentacular canal; par – parietal; pmx – premaxilla; psa – pseudoangular; prfr – prefrontal; psd – pseudodentary; pt – pterygoid; q – quadrate; smx – septomaxilla; sphen – sphenethmoid; st – stapes; sq – squamosal; utf – upper temporal fossa; vom – vomer; vf – vomeral foramen.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453376	https://doi.org/10.3897/vz.75.e149399.figure3	Figure 3. Braincase morphology of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female). 3 D reconstruction of the braincase is shown with colored rendering in dorsal (A), left lateral (B), and right medial (C) views, and of the isolated sphenethmoid in posterior (D) and anterior (E) views. Scale bar equals 1 mm. Abbreviations: Id – incisure for the dorsal branch of the olfactory nerve; Iv – foramen for the ventral branch of the olfactory nerve; II – incisure for the optic nerve; Vop – foramen for the deep ophthalmic branch of the trigeminal nerve; VII – foramen for the trunk of the facial nerve; VIIIa – foramen for the anterior branch of the auditory nerve; VIImd – foramina for the medial branches of the auditory nerve; VIIIp – foramen for the posterior branch of the auditory nerve; bca – basicranial articulation; car + VIIpal – foramen for the carotid artery and the palatal branch of the facial nerve; dmp – dorsomedial process of the sphenethmoid; dv? – incisure for the dorsal vein; fe – endolymphatic foramen; fper – perilymphatic foramen; fv – fenestra vestibuli; jf – jugular foramen; ns – nasal septum; sn – sola nasi (processus conchoides).	Figure 3. Braincase morphology of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female). 3 D reconstruction of the braincase is shown with colored rendering in dorsal (A), left lateral (B), and right medial (C) views, and of the isolated sphenethmoid in posterior (D) and anterior (E) views. Scale bar equals 1 mm. Abbreviations: Id – incisure for the dorsal branch of the olfactory nerve; Iv – foramen for the ventral branch of the olfactory nerve; II – incisure for the optic nerve; Vop – foramen for the deep ophthalmic branch of the trigeminal nerve; VII – foramen for the trunk of the facial nerve; VIIIa – foramen for the anterior branch of the auditory nerve; VIImd – foramina for the medial branches of the auditory nerve; VIIIp – foramen for the posterior branch of the auditory nerve; bca – basicranial articulation; car + VIIpal – foramen for the carotid artery and the palatal branch of the facial nerve; dmp – dorsomedial process of the sphenethmoid; dv? – incisure for the dorsal vein; fe – endolymphatic foramen; fper – perilymphatic foramen; fv – fenestra vestibuli; jf – jugular foramen; ns – nasal septum; sn – sola nasi (processus conchoides).	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453377	https://doi.org/10.3897/vz.75.e149399.figure4	Figure 4. 3 D reconstruction of the atlas of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female) in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar equals 1 mm. Abbreviations: acot – anterior cotyles; mo – medial outgrowth; na – neural arch; pcot – posterior condyle; pzp – postzygopophyseal processes; scs – spinal cord support; snf – spinal nerve foramen.	Figure 4. 3 D reconstruction of the atlas of the holotype of Ichthyophis griseivermis sp. nov. (ZMMU A- 8208, adult female) in anterior (A), posterior (B), left lateral (C), dorsal (D), ventral (E), and right lateral (F) views. Scale bar equals 1 mm. Abbreviations: acot – anterior cotyles; mo – medial outgrowth; na – neural arch; pcot – posterior condyle; pzp – postzygopophyseal processes; scs – spinal cord support; snf – spinal nerve foramen.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453378	https://doi.org/10.3897/vz.75.e149399.figure5	Figure 5. Maximum Likelihood (ML) phylogenetic tree of the genus Ichthyophis derived from the analysis of 3967 bp of cyt b, 12 S rRNA, and 16 S rRNA mtDNA gene sequences. For voucher specimen information and GenBank accession numbers, see Table S 2. Numbers at the nodes correspond to ML UFBS / BI PP support values, respectively. Species symbols and locality numbers correspond to those provided on the map in Figure 1. Unstriped species of Ichthyophiidae are shaded in grey. The thumbnail shows the holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208; denoted by an asterisk in the tree); photograph by A. M. Bragin.	Figure 5. Maximum Likelihood (ML) phylogenetic tree of the genus Ichthyophis derived from the analysis of 3967 bp of cyt b, 12 S rRNA, and 16 S rRNA mtDNA gene sequences. For voucher specimen information and GenBank accession numbers, see Table S 2. Numbers at the nodes correspond to ML UFBS / BI PP support values, respectively. Species symbols and locality numbers correspond to those provided on the map in Figure 1. Unstriped species of Ichthyophiidae are shaded in grey. The thumbnail shows the holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208; denoted by an asterisk in the tree); photograph by A. M. Bragin.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453379	https://doi.org/10.3897/vz.75.e149399.figure6	Figure 6. The holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208, adult female). A Dorsal view of annuli in the middle of the body; B ventral view of annuli in the middle of the body; C general dorsolateral view of the body, right side; D general ventrolateral view of the body, right side. Scale bar equals 5 mm. Photographs by A. M. Bragin.	Figure 6. The holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208, adult female). A Dorsal view of annuli in the middle of the body; B ventral view of annuli in the middle of the body; C general dorsolateral view of the body, right side; D general ventrolateral view of the body, right side. Scale bar equals 5 mm. Photographs by A. M. Bragin.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453380	https://doi.org/10.3897/vz.75.e149399.figure7	Figure 7. Details of external morphology of the holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208, adult female). A Lateral view of the head, right side; B lateral view of the head, right side; C dorsal view of the head; D ventral view of the head; E lateral view of the tail, left side; F lateral view of the tail, right side; G dorsal view of the tail; H ventral view of the tail. Scale bar equals 5 mm (all photographs shown in one scale). Photographs by A. M. Bragin.	Figure 7. Details of external morphology of the holotype of Ichthyophis griseivermis sp. nov. in life (ZMMU A- 8208, adult female). A Lateral view of the head, right side; B lateral view of the head, right side; C dorsal view of the head; D ventral view of the head; E lateral view of the tail, left side; F lateral view of the tail, right side; G dorsal view of the tail; H ventral view of the tail. Scale bar equals 5 mm (all photographs shown in one scale). Photographs by A. M. Bragin.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453381	https://doi.org/10.3897/vz.75.e149399.figure8	Figure 8. The holotype of Ichthyophis griseivermis sp. nov. in life in situ (ZMMU A- 8208, adult female). Photograph by A. M. Bragin.	Figure 8. The holotype of Ichthyophis griseivermis sp. nov. in life in situ (ZMMU A- 8208, adult female). Photograph by A. M. Bragin.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
7E85E3AC41CC519B9D6F9D682BFCF2C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1453374	https://doi.org/10.3897/vz.75.e149399.figure1	Figure 1. Distribution of the unstriped species of the genus Ichthyophis from Indochina and southern China. Symbols indicating Ichthyophis species are identical to those used in Figure 5; a dot in the center of a symbol indicates the type locality of a species. Localities: 1 Xuan Lien NP, Thanh Hoa Province, Vietnam; 2 Pu Hoat NR, Nghe An Province, Vietnam; 3 Maandi, Jinping County, Yunnan Province, China; 4 Cha Lo, Quang Binh Province, Vietnam; 5 Cat Loc, Cat Tien NP, Lam Dong Province, Vietnam; 6 Phnum Dalai, Phnom Samkos WS, Pursat Province, Cambodia; 7 Luang Prabang, the former French administrative centre of “ Haute Laos ” (following Geissler et al. 2015; but see the Discussion for the problem of the I. laosensis type locality); 8 Mae Wang, Chiang Mai Province, Thailand; 9 Muang Liep, Sayaboury Province, Laos; 10 Bhuping summer palace, Chiang Mai Province, Thailand; 11 Doi Ang Khang, Chiang Mai Province, Thailand.	Figure 1. Distribution of the unstriped species of the genus Ichthyophis from Indochina and southern China. Symbols indicating Ichthyophis species are identical to those used in Figure 5; a dot in the center of a symbol indicates the type locality of a species. Localities: 1 Xuan Lien NP, Thanh Hoa Province, Vietnam; 2 Pu Hoat NR, Nghe An Province, Vietnam; 3 Maandi, Jinping County, Yunnan Province, China; 4 Cha Lo, Quang Binh Province, Vietnam; 5 Cat Loc, Cat Tien NP, Lam Dong Province, Vietnam; 6 Phnum Dalai, Phnom Samkos WS, Pursat Province, Cambodia; 7 Luang Prabang, the former French administrative centre of “ Haute Laos ” (following Geissler et al. 2015; but see the Discussion for the problem of the I. laosensis type locality); 8 Mae Wang, Chiang Mai Province, Thailand; 9 Muang Liep, Sayaboury Province, Laos; 10 Bhuping summer palace, Chiang Mai Province, Thailand; 11 Doi Ang Khang, Chiang Mai Province, Thailand.	2025-10-29	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.		Zenodo	biologists	Poyarkov, Nikolay A.;Skorinova, Dana D.;Bragin, Andrey M.;Kolchanov, Veniamin V.;Gorin, Vladislav A.;Trofimets, Alexey V.;Yuzefovich, Alexander P.;Le, Dac Xuan;Nguyen, Tan Van;Skutschas, Pavel P.			
