taxonID	type	description	language	source
057387FDFF84FFB0B7D4FA3BFB29FE62.taxon	vernacular_names	[Japanese name: Yunohama-himegumo-zoku; Korean name: Tam-ra-kko-ma-geo-mi-sok]	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF84FFB0B7D4FA3BFB29FE62.taxon	type_taxon	Type species. Theridium yunohamense Bösenberg & Strand, 1906	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF84FFB0B7D4FA3BFB29FE62.taxon	diagnosis	Diagnosis. Yunohamella species resemble those of Takayus Yoshida, 2001, both with a large tegulum that covers most of the palp in retrolateral view, relatively small and globular theridiid tegular apophysis, and no distinct groove or boundary between the surfaces of the conductor and tegulum (Yunohamella: Figs 2, 4 A – F; Yoshida 2003, figs 235, 238, 242; Takayus: Yoshida 2003, figs 225, 229, 232). However, Yunohamella can be distinguished from Takayus by the following characteristics: generally dark and cryptic body coloration (Fig. 1 B – I; Yoshida 2003, figs 572, 573) (vs bright yellowish to orangish in Takayus) (Yoshida 2003, figs 569 – 571); wavy cardiac pattern on the dorsum of the abdomen (Fig. 1; Yoshida 2003, figs 572, 573) (vs feather-like flecks in Takayus) (Yoshida 2003, figs 569 – 571); females with highly variable epigynal morphology among species, with the type species Y. yunohamensis bearing a large angular scape, while others lack a scape or possess a pair of semicircular epigynal processes (Figs 3, 4 G, I; Yoshida 2003, figs 233, 236, 239) (vs relatively uniform epigynal morphology in Takayus, with a small triangular scape on the posterior margin of the epigynal plate) (Yoshida 2003, figs 222, 227); males with palps bearing a thinner and more slender embolus (Figs 2 B, E, 3 B, E; Yoshida 2003, figs 235, 238, 241) (vs thicker embolus in Takayus) (Yoshida 2003, figs 224, 229, 232); and a relatively slender conductor with shape varying among species, branching, twisted or spatulate (Figs 2, 4 A – F; Yoshida 2003, figs 235, 238, 241) (vs conductor relatively broad, short, and blunt in Takayus) (Yoshida 2003, figs 224, 229, 232). Additionally, both genera differ in microhabitat preference: Takayus spiders typically build tangle webs under leaves or among branches of shrubs, whereas Yunohamella species inhabit shaded, moist, and cryptic environments such as rocky precipices along streams and waterfalls or the surfaces of tree trunks (Yoshida 2007).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	vernacular_names	[Japanese name: Haraobi-himegumo; Korean name: Heo-ri-ddi-kko-ma-geo-mi]	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	description	Figs 1 A – C, F – H; 2 A – C; 3 A – C; 4 A – C, G, H; 5	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	materials_examined	Type-material. Theridion mneon: ♀ holotype from Kompira, Saga, Northern Kyushu, Japan, deposited in Senckenberg Museum, Frankfurt am Main (SMF 3034), Germany [photograph examined via Dupérré (2023)]. Yunohamella varietas: ♂ holotype from Ulsan, Korea, deposited in the National Institute of Biological Resources (NIBRIV 0000879011), Incheon, Republic of Korea [photographs and illustrations examined via Lee & Kim (2021)]. Material examined. JAPAN, Akita Pref.: 1 ♀, Kisakata-machi, Nikaho-shi, 24. VII. 1993, A. Fukushima leg. [reported as Y. lyrica in Fukushima (1997)] (NSMT-Ar 25444). Ibaraki Pref.: 2 ♂, 1 ♀, lakeshore of Kasumigaura, Okijuku-machi, Tsuchiura-shi, 22. V. 2019, Y. Suzuki leg. (TKPM-AR 3251). Chiba Pref.: 2 ♀, Risoukai-kinenshizen-koen Park, Yamazaki, Noda-shi, 11. VII. 2010, A. Tanikawa leg. (TKPM-AR 3264); 3 ♂, 2 ♀, same locality as previous, 16. V. 2010, A. Tanikawa leg. (TKPM-AR 3265); 2 ♂, 2 ♀, lakeshore of Tega-numa, Tobarishinden, Kashiwa-shi, 13. VII. 2025, R. Serita leg. (RS, per. coll.); 1 ♀ with an egg sac, same locality as previous, 13. VII. 2025, R. Hidaka leg. (RH, per. coll.). Saitama Pref.: 1 ♀ (collected as a juvenile and matured under rearing), riverbed of Edo-gawa River, Fukaishinden, Yoshikawa-shi, Saitama, 05. I. 2024, A. Kikuchi leg. (AK, per. coll.). Kanagawa Pref.: 1 ♂ (collected as a juvenile and matured under rearing), riverbed of Sagami-gawa River, Kawaraguchi, Ebinashi, 29. II. 2024, A. Kikuchi leg. (AK, per. coll.). Mie Pref.: 1 ♂, 4 ♀, riverbed under Machiya-bashi Bridge, Nao, Asahi-cho, Mie-gun, 29. VII. 2023, T. Shiozaki leg. (TKPM-AR 3201 – 3205); 1 ♀, Kawarada-cho, Yokkaichi-shi, 18. VII. 1977, K. Kaihotsu leg. (TKPM-AR 3215); 1 ♀, Oe, Minamiise-cho, Watarai-gun, 09. VI. 2022, K. Kumada leg. (TKPM-AR 3255); 1 ♂, Sekicho-shinjo, Kameyama-cho, 30. V. 2006, K. Kumada leg. (TKPM-AR 3256). Shimane Pref.: 1 ♂, Isotake, Isotake-cho, Ota-shi, 23. VII. 2006, A. Tanikawa leg. [reported as Takayus lyricus by Shinkai & Tanikawa (2006)] (TKPM-AR 3263). Tokushima Pref.: 1 ♂, 1 ♀, riverbed of Yoshino-gawa River, Tokushima-shi, 03. V. 2023, Y. Suzuki leg. (TKPM-AR 3194); 1 ♀, same data as previous (TKPM-AR 3195); 2 ♀, same locality as previous, 18. V. 2023, Y. Suzuki leg. (TKPM-AR 3196); 1 ♀, same locality as previous, 13. X. 2023, Y. Suzuki leg. (TKPM-AR 3197); 1 ♂, 1 ♀, riverbed of Yoshino-gawa River under Nada-bashi Bridge, Nada, Tokumei, Aizumi-cho, Itano-gun, 18. IV. 2024, Y. Suzuki leg. (TKPM-AR 3257); 1 ♀, riverbed at estuary of Akui-gawa River, Fudohigashimachi, Tokushima-shi, 18. IV. 2024, Y. Suzuki leg. (TKPM-AR 3258); 1 ♂, riverbed at estuary of Katsuura-gawa River, Wadabarisoto, Ronden-cho, Tokushima-shi, 22. IV. 2024, Y. Suzuki leg. (TKPM-AR 3259); 2 ♂, 2 ♀, riverbed at Naka-gawa River, Sumiyoshi-cho, Anan-shi, 02. V. 2024, Y. Suzuki leg. (TKPM-AR 3260). Saga Pref.: 1 ♂, 3 ♀, Kosegawa-choseichi balancing reservoir, Kinryu-machi Chifu, Saga-shi, 18. V. 2025, A. Nakamura leg. (TKPM-AR 3417). Oita Pref.: 4 ♀, Otsuka, Nakatsu-shi, 21. VI. 2020, A. Nakamura leg. (TKPM-AR 3253); 2 ♀, Kaizaki, Saiki-shi, 27. VIII. 2024, Y. Hino leg. (TKPM-AR 3252). Kumamoto Pref.: 1 ♂, 1 ♀, Iwasaka, Ozumachi, 21. VII. 2024, A. Nakamura leg. (TKPM-AR 3254). REPUBLIC OF KOREA, Gyeongsangnam-do: 1 ♂, Gyeongsangnamdo Arboretum, Ibanseong-myeon, Jinju-si (35 ° 09 ' 40 '' N, 128 ° 17 ' 41 '' E), 1 – 17. VI. 2011, J. H. Hwang leg., Malaise trap. (KNU). Jeonbuk: 1 ♂, Kunsan National University, Miryong-dong, Gunsan-si (35 ° 57 ' 01 " N, 126 ° 40 ' 55 " E), 11 – 25. IV. 2020, J. H. Sohn et al. leg., Malaise trap. (KNU). Jeollanam-do: 1 ♂, Hakgu-ri, Seo-myeon, Suncheon-shi (35 ° 02 ' 29 " N, 127 ° 26 ' 53 " E), 03. V – 10. VI. 2021, S. W. Kim leg., Malaise trap. (KNU); 1 ♂, same locality as previous, 10. VI – 21. VII. 2021, S. W. Kim leg. (KNU).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	diagnosis	Diagnosis. Yunohamella mneon is similar to Y. lyrica in the shape of the epigyne, sharing a pair of semicircular sclerotized epigynal processes and a broad atrium, but can be distinguished from the latter by the following characteristics: carapace pale yellowish brown with a dark brown spot anteriorly to the fovea and dark stains on the lateral margins (Fig. 1 B, C) (vs uniformly reddish brown in Y. lyrica) (Fig. 1 D, E); dorsum of the abdomen pale yellowish brown with a longitudinal white band (Fig. 1 B, C, F – H) (vs dark reddish brown with a diamond-shaped whitish marking with pink spots, anterior side of abdomen with a pair of large blackish markings in Y. lyrica) (Fig. 1 D, E, I); legs with dark brown annulations on the terminal region of each segment (Fig. 1 B, C, F – H) (vs annulations on both the terminal and middle regions of each segment in Y. lyrica) (Fig. 1 D, E, I); male palp with conductor bearing a blunt, spoon-like tip (Figs 2 A – C, 4 A – C) (vs conductor twisted, flame-like, and with a distally sharper tip in Y. lyrica) (Figs 2 D – F, 4 D – F); sperm duct more intricate, with tight loops (Figs 2 A; 4 A) (vs sperm duct simpler, with more spread loops in Y. lyrica) (Figs 2 D, 4 D); female genitalia with longer copulatory ducts (CDL / SW: 2.20), more convoluted, entering the spermathecae at an angle nearly parallel to the orientation of the basal portion of the fertilization ducts (Figs 3 B, C, 4 H) [vs relatively shorter (CDL / SW: 1.35), simple C-shaped trajectory, entering the spermathecae at an angle nearly orthogonal to the orientation of the basal portion of the fertilization ducts in Y. lyrica.] (Figs 3 D, E, 4 I).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	description	Redescription. Male (TKPM-AR 3194). Measurements. Total length 2.24 long. Carapace 1.15 long, 0.96 wide. Eye diameter: AME, 0.124; ALE, 0.069; PME, 0.075; PLE, 0.078. Eye interdistances: AME-AME, 0.090; AME-ALE, 0.030; PME-PME, 0.085; PME-PLE, 0.085. Length of legs: I, 2.96 + 0.54 + 2.90 + 3.01 + 1.05 = 10.46; II, 1.97 + 0.53 + 1.47 + 1.77 + 0.77 = 6.51, III, 1.24 + 0.31 + 0.72 + 0.95 + 0.50 = 3.72; IV, 1.87 + 0.48 + 1.24 + 1.49 + 0.68 = 5.76. Abdomen 1.06 long, 0.89 wide. Coloration and markings (Fig. 1 B, F). Carapace pale yellowish brown with a dark brown spot anteriorly to fovea and mottled dark on lateral margins; chelicerae and labium yellowish brown; sternum yellowish brown with dark margins; legs pale yellowish brown with dark brown annulations on terminal region of each segment; dorsum of abdomen pale yellowish brown with longitudinal white band and small blackish spots; ventral side of abdomen with a large black marking on middle part and black transverse band running laterally; spinnerets surrounded by black markings. Palp (Figs 2 A – C, 4 A – C). Tegulum large, covering most part of palpal bulb in ventral view; embolus almost straight and slightly curved, broad at base and pointed at tip; conductor broad, beak-like, conjugated with the tegulum; tip of the conductor concave, spoon-like; trajectory of sperm ducts intricate with tight loops; theridiid tegular apophysis large, strongly sclerotized with angular corner; median apophysis small, most portion hidden under the theridiid tegular apophysis; subtegulum distinct. Female (TKPM-AR 3195). Measurements. Body 3.42 long. Carapace 1.25 long, 1.11 wide. Eye diameter: AME, 0.102; ALE, 0.092; PME, 0.101; PLE, 0.098. Eye interdistances: AME-AME, 0.095; AME-ALE, 0.036; PME-PME, 0.077; PME-PLE, 0.072. Length of legs: I, 2.13 + 0.51 + 1.92 + 2.03 + 0.81 = 7.40; II, 1.57 + 0.37 + 1.09 + 1.11 + 0.54 = 4.68; III, 0.97 + 0.33 + 0.57 + 0.66 + 0.45 = 2.98; IV, 1.32 + 0.35 + 1.07 + 1.09 + 0.61 = 4.44. Abdomen 2.00 long, 1.99 wide. Coloration and markings (Fig. 1 C, G – H). Same as the male, slightly brighter. Genitalia (Figs 3 A – C, 4 D, E): epigyne with oval atrium; pair of semicircular, pigmented, sclerotized epigynal processes positioned anterolaterally on epigynal plate; copulatory openings located laterally on epigynal processes; copulatory ducts convoluted with tight loops, entering spermathecae at an angle nearly parallel to orientation of basal portion of the fertilization ducts; spermathecae spherical; fertilization ducts arising posteriorly from spermathecae, with simple curve and pointed tip. Variations. Some female specimens collected in Saga and Oita, Kyushu, Japan, had darker markings on the cephalic region and dark annulations on the middle part of the femur of legs, as well as darker abdomen compared with those of the type specimen and specimens from other regions (Fig. 5 C).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	discussion	Remarks. Zhong et al. (2025) treated Yunohamella mneon as a junior synonym of Y. lyrica, based on the redescription made by Dupérré (2023), citing the similarity in the epigynal morphology. However, we argue that the type specimen of Y. mneon can be clearly distinguished from Y. lyrica based on several morphological features. Yunohamella mneon has a dark marking anteriorly to the fovea of the carapace, and mottled lateral margins, the blackish annulations on the middle portion of the leg segment are absent (Fig. 1 A; Bösenberg & Strand 1906, fig. 286; Dupèrrè 2003, fig. 73 A), and has internal genitalia with more convoluted copulatory ducts with tight loops (Dupèrrè 2003, fig. 73 B, C). On the other hand, females of Y. lyrica possess uniformly reddish-brown carapace, legs with dark brown annulations on both terminals and middle region of each segment, and relatively short and less convoluted copulatory ducts (Figs 1 D, E, I, 3 E, F, 4 J). The type locality of Y. mneon is assumed to be Kinryu-cho, Saga Pref., where Kompira (Kotohira) Shrine is located (Ichiba 1975, fig. 6). Additional specimens were collected from Kosegawa-choseichi balancing reservoir, only 4 km south from Kompira Shrine (Fig. 6 C). These specimens showed consistency in key morphological characters with the type specimen illustrated by Bösenberg & Strand (1906), with some intraspecific variations in coloration and markings, as described above. In the molecular phylogeny (Fig. 7), specimens of Y. mneon in both Japan and Korea formed a monophyletic group with high support values, and the p-distances between specimens in Korea and Japan varied between 0.0018 and 0.0091. Judging from both morphological comparison and molecular phylogenetic analysis among numerous Japanese and Korean specimens of Yunohamella species, males identified as Yunohamella varietas are well-grouped with the females of Yunohamella mneon (Fig. 7). Therefore, they are considered conspecific, and Y. mneon should be treated as a senior synonym of Y. varietas syn. nov.	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	biology_ecology	Natural History. In Japan, Y. mneon has primarily been found in grasslands on riverbeds and lakeshores, where it was observed constructing a tangle web between grasses near the ground. In the Republic of Korea, Y. mneon was collected from paddy fields and forest plantations (Lee & Kim 2021; this study). In contrast, Y. lyrica and other Yunohamella species are typically associated with shaded, moist forest or cliffs and rocky walls along mountainous streams, and tree trunks (Yoshida 1987, 2003). These observations suggest that Y. mneon prefers open landscapes, in contrast to at least some of its congeners.	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF85FFBBB7D4FE2BFF2BF9DF.taxon	distribution	Distribution. Republic of Korea (Jeonbuk, Jeollanam-do, Gyeongsangnam-do), Japan (Honshu to Kyushu) (Fig. 6).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	vernacular_names	[Japanese name: Shimofuri-himegumo; Korean name: Seo-ri-kko-ma-geo-mi]	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	description	Figs 1 D, E, I; 2 D – F; 3 D – F; 4 D – F, I, J	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	materials_examined	Type-material: Theridion lyricum: female holotype from Georgia, USA; not examined. Material examined. JAPAN, Hokkaido: 2 ♀, Koetoi-jinjya, Koetoimura-koetoi, Wakkanai-shi, 11. VII. 1990, K. Kumada leg. (NSMT-Ar 18130); 1 ♀, Okikineushi, Biei-cho, Kamikawa-gun, 10. IX. 2016, Y. Suzuki leg. (TKPM-AR 3198). Miyagi Pref.: 1 ♂, 1 ♀, Uwahara, Hanayama-mura, Kurihara-gun, 04. VII. 1997, K. Kumada leg. (NSMT-Ar 3766); 3 ♀, Ushibuchikoen, Ichihazama-cho, Kurihara-gun, 29. VI. 1997, K. Kumada leg. (NSMT-Ar 3767); 1 ♀, Izunuma, 06. VII. 1986, A. Tanikawa leg. (TKPM-AR 3266); 1 ♀, same locality as previous, 20. VII. 1986, A. Tanikawa leg. (TKPM-AR 3267); 1 ♀, same locality as previous, 07. VI. 1987, A. Tanikawa leg. (TKPM-AR 3268). Fukushima Pref.: 1 ♀, Shirakawanoseki, Hatajuku, Shirakawa-shi, 24. VI. 1990, K. Kumada leg. (NSMT-Ar 18575). Ibaraki Pref.: 1 ♀, Oda, Tsukuba-shi, 27. V. 2017, Y. Suzuki leg. (YS. per. coll.). Tokyo: 1 ♂, Mitsuzawa, Hinode-machi, Nishitama-gun, 26. V. 1990, K. Kumada leg. (NSMT-Ar 2185); 2 ♀, Mt. Takao-san, Hachioji-shi, 28. V. 1994, M. Ban leg. (NSMT-Ar 10566); 1 ♀, Okutama-cho, 22. VII. 2007, A. Tanikawa leg. (TKPM-AR 3269). Kanagawa Pref.: 1 ♀, Shinsei-ko Lake, Hadano-shi, 07. VIII. 1983, S. Matsumoto leg. (NSMT-Ar 2101); 1 ♀, Fudojiri, Atsugi-shi, 21. VI. 1997, M. Ban leg. (NSMT-Ar 10910); 2 ♂, 1 ♀, Mikarube-rindo, Mikarube, Hadano-shi, 06. VI. 1993, K. Kumada leg. (NSMT-Ar 15023); 1 ♀, 1 juvenile, Totsuka-ku, Yokohama-shi, 19. IX. 1998, A. Tanikawa leg. (TKPM-AR 3270); 1 ♂, Kuzuha-ryokuchi, Hadano-shi, 12. VI. 1996, A. Tanikawa leg. (TKPM-AR 3271); 1 ♀, Fudakake-morino-ie, Susugaya, Kiyokawa-mura, Aiko-gun, 09. VI. 1991, A. Tanikawa leg. (TKPM-AR 3272). Yamanashi Pref.: 1 ♂, Ogiyama, Otsuki-shi, 05. VI. 1999, M. Ban leg. (NSMT-Ar 11173); 1 ♀, Fujikawa-cho (previously “ Masuho-cho ”), Minamikoma-gun, 24 – 26. VII. 2004, A. Tanikawa leg. (TKPM-AR 3273). Shizuoka Pref.: 1 ♂, Ichiyama, Amagiyugashima-cho, Tagata-gun, 02. VI. 1984, K. Kumada leg. (NSMT-Ar 17803). Nagano Pref.: 1 ♂, Mitsumata, Horikane-mura, Minamiazumi-gun, 11. VII. 1984, K. Kumada leg. (NSMT-Ar 8536); 1 ♂, Yatsugatake-kogen, 18. VI. 1994, M. Ban & H. Ikeda, leg. (NSMT-Ar 10728); 8 ♀, Susado, Karasugawa, Horikane-mura, Minamiazumi-gun, 20. VI. 1976, K. Kumada leg. (NSMT-Ar 15754); 1 ♂, 14 ♀, Karasugawa Susado, Horikane-mura, Minamiazumi-gun, 30. VII. 1977, K. Kumada leg. (NSMT-Ar 15755); 1 ♀, Megami-ko Lake, Tateshina-machi, Kitasaku-gun, 18. VII. 2009, A. Tanikawa leg. (TKPM-AR 3274); 1 ♀, Yumenodaira, Ashida-hakkano, Tateshina-machi, Kitasaku-gun, 20. VII. 2009, A. Tanikawa leg. (TKPM-AR 3275); 1 ♀, Sugadaira-kogen, Ueda-shi, 27. VII. 1980, A. Tanikawa leg. (TKPM-AR 3276); 1 ♀, same locality as previous, 24. VII. 1988, A. Tanikawa leg. (TKPM-AR 3277). Fukui Pref.: 2 ♀, Karikomiike, Kamiuchinami, Oono-shi, 18. VII. 2002, K. Kumada leg. (NSMT-Ar 15154); Mie Pref.: 1 ♀, Ise Jingu, 81 rinpan, Ujitachi-cho, collected in 2003, K. Kumada leg. (NSMT-Ar 13759); 1 ♂, 1 ♀, Nonoboriyama, Asakayama-cho, Kameyama-shi, 15. VII. 1997, K. Kumada leg. (NSMT-Ar 17649); 1 ♂, Kiwacho-Kogurusu, Kumano-shi, 26. V. 1996. T. Shiozaki leg. (TKPM-AR 3206); 1 ♀, same data as previous. (TKPM-AR 3207); 1 ♂, same locality as previous, 22. V. 1997, T. Shiozaki leg. (TKPM-AR 3208); 1 ♀, same data as previous (TKPM-AR 3209), 1 ♀, same data as previous (TKPM-AR 3210); 1 ♀, Osugi, Odai-cho, Taki-gun, 06. VII. 2002, T. Shiozaki leg. (TKPM-AR 3211); 1 ♀, Misugicho-Tarou, Tsu-shi, 08. VII. 2002, T. Shiozaki leg. (TKPM-AR 3212); 1 ♂, Mt. Obora-yama, Misugicho-Yachi, Tsu-shi, 14. VI. 2009, K. Kaihotsu leg. (TKPM-AR 3213); 1 ♀, Niu, Taki-cho, Taki-gun, 20. VI. 2021, K. Kaihotsu leg. (TKPM-AR 3214). Shiga Pref.: 1 ♂, 2 ♀, Yamakado-shicchi, Nishiazaichoyamakado, Nagahama-shi, 25. V. 2019, K. Kumada leg. (NSMT-Ar 18850). Nara Pref.: 1 ♂, Oono, Tsuchiyahara, Mitsue-mura, Uda-gun, 10. VI. 2002, K. Kumada leg. (NSMT-Ar 19504). Wakayama Pref.: 1 ♂, Myouhouzan, Minamihirano, Nachikatsuura-cho, Higashimuro-gun, 11. VII. 1990, Y. Chikuni leg. (NSMT-Ar 15756). Tokushima Pref.: 1 ♀, Mizunokuchi-Toge Pass, Oiya, Nishiiyayama-son, Miyoshi-shi, 23. VI. 2017, H. Bando leg. (TKPM-AR 3199); 3 juveniles, Hinomaruyama-Rindo, Oiya, Nishiiyayama-son, Miyoshi-shi, 30. IX. 2018, H. Bando leg. (TKPM-AR 3200). Fukuoka Pref.: 1 ♀, Mt. Tachibana-yama, Higashi-ku, Fukuoka-shi, 23. VIII. 1991, A. Tanikawa leg. (TKPM-AR 3278). Oita Pref.: 1 ♂, Sanko-usugi, Nakatsu-shi, 02. VI. 2021, A. Nakamura leg. (TKPM-AR 3418). Yakushima Island (Kagoshima Pref.): 1 ♂, Shiratani-sen, Yakushima-cho, Kumage-gun, 25. V. 2024, A. Tanikawa leg. (TKPM-AR 3279). REPUBLIC OF KOREA, Gangwon: 1 ♀, Guryongsa Course, Mt. Chiaksan, Socho-myeon, Wonju-si (35 ° 09 ’ 40 ’’ N, 128 ° 17 ’ 41 ’’ E), 24. VI. 2024, J. G. Lee leg. (KNU); 1 juvenile, Hangye-ri, Buk-myeon, Injegun (38 ° 08 ’ 47 ’’ N, 128 ° 15 ’ 48 ’’ E), 19. VI – 24. VII- 2022, Park & Nam leg., Malaise trap. (KNU). Gyeonggi-do: 1 juvenile, Tonggu-ri, Baekhak-myeon, Yeoncheon-gun (38 ° 01 ’ 43 ” N, 126 ° 55 ’ 17 ” E), 15. IV – 03. V. 2019, J. G. Lee leg., malaise trap (KNU). Gyeongsangbuk-do: 1 ♀, Gakhwasa Temple, Mt. Gakhwasan, Chunyang-myeon, Bonghwagun (36 ° 59 ’ 26 ’’ N, 128 ° 54 ’ 34 ’’ E), 08. VII. 2019, J. G. Lee leg. (KNU).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	diagnosis	Diagnosis. Males of Y. lyrica can be distinguished from congeners by the twisted, flame-like, distally sharper tip of conductor (Fig. 2 D – F) (vs in other Yunohamella species, tip of conductor is blunt or truncated, bifurcated or quadrifurcated) (blunt or truncated: e. g., Marusik & Tsellarius 1986, fig. 2; Esyunin & Efimik 1996, figs 1, 2; Yoshida 2003, fig. 238; Yoshida 2012, fig. 7; Zhong et al. 2025, fig. 4 C); (bifurcated: e. g., Gao & Li 2014, figs 107 – 109; Zhong et al. 2025, figs 2 C, 5 C); (quadrifurcated: e. g., Yoshida 2003, fig. 235). Females of Y. lyrica closely resemble those of Y. mneon stat. rev. in the shape of the epigyne, as both have a pair of semicircular sclerotized epigynal processes and a broad atrium, but can be distinguished by the uniformly reddish-brown carapace; legs with dark brown annulations on both the terminal and middle regions of each segment; a dark reddish-brown abdomen bearing a diamond-shaped whitish marking with pink spots; a pair of large blackish markings on the anterior side of the abdomen; and copulatory ducts that are relatively short compared to the width of the spermathecae and less convoluted (Figs 1 E, 3 D – F, 4, I, J).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	description	Description. See Yoshida (1987, 2001, 2003) for details.	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8EFFB8B7D4F9B7FD92F882.taxon	distribution	Distribution. China, Korea (All provinces), Japan (Hokkaido to Kyushu and Yakushima Island), and North America (World Spider Catalog 2025).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	vernacular_names	[Japanese name: Sato-himegumo; Korean name: A-dam-son-kko-ma-geo-mi]	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	description	Fig. 8	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	materials_examined	Type-material: Theridion adamsoni: ♀ syntypes from Hitiaa, Tahiti, in the B. P. Bishop Museum, Honolulu (BPBM); not examined. Material examined. JAPAN, Tokushima Pref.: 1 ♂, Kaminakasuji, Hachiman-cho, Tokushima-shi, 12. VII. 2024, Y. Suzuki leg. (TKPM-AR 3261); 1 ♀, Ichibacho-Kamigirai, Awa-shi, 18. VIII. 2024, Y. Suzuki leg. (TKPM-AR 3262). REPUBLIC OF KOREA, Jeollanam-do: 1 ♀, Gageodo Is., Heuksan-myeon, Sinan-gun (34 ° 03 ' 13 " N, 125 ° 07 ' 42 " E), 24. VIII. 2024, J. G. Lee leg. (KNU).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	diagnosis	Diagnosis & Description. See Levi (1957) (under T. hobbsi Gertsch & Archer, 1942) and Levi (1967).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	distribution	Distribution. USA, Caribbean, South America (including Brazil, Castanheira et al. 2016). Introduced to St. Helena, Ascension Is., Ghana, South Africa, Seychelles, China, Korea, Japan, Pacific Is.	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
057387FDFF8AFFBEB7D4FF0AFE4BFE8E.taxon	discussion	Remarks. Platnickina adamsoni is commonly found in tropical America, and widespread across the world including Asia (Yoshida 2003; World Spider Catalog 2025). Although the general appearance of P. adamsoni resembles Y. mneon, they can clearly be distinguished by the genital morphology (Figs 2 A – F, 3 A – F, 4 A – J vs 8 C – E). Moreover, P. adamsoni usually display synanthropic habits, inhabiting walls of buildings, gardens and yards (Fig. 8 A, B; Yoshida 2003).	en	Suzuki, Yuya, Lee, Jun-Gi, Kumada, Ken-Ichi (2025): Taxonomic notes on a comb-footed spider, Yunohamella mneon (Bösenberg & Strand, 1906), stat. rev. (Araneae: Theridiidae) in East Asia. Zootaxa 5691 (2): 359-375, DOI: 10.11646/zootaxa.5691.2.10, URL: https://doi.org/10.11646/zootaxa.5691.2.10
