taxonID	type	description	language	source
171087DBFFBD287D45D324AC223A9DF3.taxon	materials_examined	Type species. Abbrosoga errata Caldwell, 1951	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBD287D45D324AC223A9DF3.taxon	diagnosis	Diagnosis. Body pale with weakly patterned wings (known species with an entirely pale frons); head and thoracic carinae distinct, concolorous with body. Aedeagus bearing a ventral projection (unique among New World Delphacini); pygofer opening without ventral processes, anal tube (= segment X) without processes. Amended description. Elongate and slender forms. Head narrower than pronotum (Figs 1 A, 3 A), vertex weakly projected in front of eyes. Vertex and frons broad, fastigium rounded (Figs 1 C, 3 C). Medial facial carinae forking near fastigium (Figs 1 B, 3 B). Median carina of vertex weak. Lateral carinae of pronotum not clearly reaching posterior margin. Hind leg (Fig. 1 D) with 2 lateral spines, 1 near femoral-tibial joint, 1 near midlength; tibial apex with 5 spinules, arranged 3 + 2. Basitarsus with 7 apical spinules, arranged 5 + 2, second tarsomere with row of 4 spinules. Forewing (Figs 1 C, 3 C) elongate, mostly clear with diffuse patterning (in distal cubital area and less often in costal region), with an elongate dark spot on commissural margin at apex of claval veins. Forewing venation (Fig. 6) with Sc and RA unbranched, RP 3 branched, M unbranched and CuA 2 – 3 branched. Metatibial spur foliaceous and tectiform, row of many fine, black-tipped teeth on posterior margin. Male genitalia with pygofer subquadrate in lateral view (Figs 2 B, 3 E), distinctly taller than wide; opening in caudal view subcircular with rounded margins (Figs 2 A, 3 F), lacking projections. Gonostyli (= parameres) (Figs 4 C, 5 A) simple (unbranched), basal angle small; proximally diverging, apex medially converging in distal forth. Aedeagus (Figs 4 B, 5 B) strongly curved ventrally, subtended with slender process (es). Aedeagal base closely approximating anal tube (aedeagal base symmetrical), suspensorium O-shaped, surrounding base of aedeagus including subtending process, stem of suspensorium (between aedeagus and anal tube) short. Male anal tube unarmed.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBD287D45D324AC223A9DF3.taxon	discussion	Remarks. Superficially, Abbrosoga is similar to other pale Delphacini, such as Toya Distant, Metadelphax Wagner, Spartidelphax Bartlett & Webb, Syndelphax Fennah, Sogatella Fennah or perhaps Tagosodes Asche & Wilson. The dorsum of the mesonotum bears a narrow, weak median vitta, unlike the strong vitta of Sogatella and Tagosodes. Unlike Metadelphax, Toya, Spartidelphax and some Syndelphax, the frons is entirely pale, instead of bearing dark markings near the carinae. The wings are diffusely patterned unlike macropterous Syndelphax. The form of the terminalia, and the aedeagus in particular, contrasts with all these genera; the aedeagus of Abbrosoga bears a subtending process that is not found in any of these other genera. It is possible that Abbrosoga is phylogenetically most closely allied with Phrictopyga and Pygospina in that they are all pale taxa, with a weak median vitta (absent in A. multispinosa n. sp.), diffusely pale wings, and bearing (at least between Pygospina and Abbrosoga) similarities in the male genitalia (shape of pygofer, parameres and aedeagus). However, both Pygospina and Phrictopyga are slightly laterally compressed with the pygofer and anal tube bearing processes that are absent in Abbrosoga. Caldwell & Martorell (1951) compared Abbrosoga to Nilaparvata Distant and Columbisoga Muir. Nilaparvata has teeth on the basitarsus of the hind legs, a synapomorphy of the genus. Columbisoga may have similar coloration and also has a similar structure of the aedeagus (i. e., bearing a ventral projection), but is in the Tropidocephalini, and lacks teeth on the calcar (and the distinct suspensorium found in Delphacini). All available specimens of A. errata were collected at relatively high elevations, above 2000 ft (610 m) as reported by Caldwell and Martorell (1951). In contrast, specimens of A. multispinosa n. sp. were collected at low elevation (the type locality is at approximately 40 m). No plant associations are available for Abbrosoga. At present the genus is only known from Puerto Rico; however, we have examined a single male specimen from Costa Rica (La Selva Biological Station, UDCC) that is substantially similar to Abbrosoga and may be placed in the genus once more material becomes available. We have also examined a specimen from Brazil (Santa Catarina, UDCC) that has substantially similar male terminalia, but is much larger and darkly patterned and may be an allied undescribed genus.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBD287D45D324AC223A9DF3.taxon	etymology	Etymology. The etymology of “ Abbrosoga ” was not specified in Caldwell & Martorell (1951), and the underlying meaning is not obvious to us. We speculate that the name is a truncation of the Latin word “ abbreviatus ” plus the Spanish word “ soga ” (rope), meaning ‘ short rope’ a possible reference to the suspensorium. Also, plausibly the name may be comprised of the prefix “ ab - ̏ (away, from) plus a truncation of “ brosis ” (Greek, food), plus “ - oga ” for euphony; or alternatively an arbitrary combination of letters. The name is treated as feminine in gender.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	description	(Figs 1, 2, 4)	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	materials_examined	Type locality. Toro Negro Mts., Puerto Rico.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	diagnosis	Amended diagnosis. Thorax with narrow pale median vitta, scutellum pale (Fig. 1 A). Forewing embrowned between apices of Sc and RA (Fig. 1 C). Frons relatively narrow (frons l: w ratio ~ 1: 2.15 – 2.37). Male gonostyli distally with ventral hook in lateral view; aedeagus with single ventral process (Figs 2 B, 4 A), anal tube dorsocaudally without projection in lateral view.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	description	Amended description. Color. General color stramineous to brownish-yellow, carinae concolorous to body; eyes dark (Figs 1 A, 1 C). Pro- and mesonotum with narrow pale median vitta, darker laterally; pronotum with small dark spot on either side of median carinae. Mesonotum pale medially, including scutellum, darker laterally. Macropterous wings (Fig. 1 C) clear with distal cubital area diffusely fuscous and a small spot on the commissural margin at apex of claval veins and a second on the leading margin between the apices of the Sc and RA. Hind wings hyaline. Structure. Length male 3.6 – 3.8 mm with wings (reported as 3.8 for male and 4.2 mm for female in Caldwell & Martorell, 1951); ~ 2.2 (n = 1) without wings. Head. Head (dorsal view, including eyes) just narrower than pronotum (Fig. 1 A); in lateral view, slightly projected anteriorly. Vertex with conspicuous carinae (median carina weak), length 0.30 mm, width 0.24 – 0.26 mm; posterior margin truncate. Frons (Fig. 1 B) widest between eyes (0.26 – 0.27 mm), weakly narrowed dorsally (0.22 mm) and ventrally (0.18 – 0.20 mm), length 0.56 – 0.64 mm; l: w ratio 1: 2.15 – 2.37; median carina forking near fastigium. Clypeus 0.29 – 0.30 mm, with median carina. Antennal scape small, longer than wide (length 0.10 – 0.11 mm), pedicel nearly 3 times longer than scape (0.27 – 0.29 mm) bearing rows of rhinaria; flagellum long and setose. Thorax. Pronotum shorter than vertex (length at midline 0.18 – 0.19 mm); lateral carinae of pronotum diverging, not reaching posterior margin; posterior margin straight (weakly notched medially). Mesonotum at midline more than 2 x length of pronotum (0.47 – 0.57 mm); lateral carinae weakly diverging, reaching posterior margin, median carina becoming obsolete on scutellum. Wings macropterous (Fig. 1 C), exceeding abdomen (forewing 3.07 – 3.08 mm); forewing venation with fork of RP at approximately same level as fork of CuA (inner and outer subapical cells subequal), fusion of anal veins (i. e., Pcu and A 1) much forks of RP and CuA; Sc and RA unbranched, RP 3 branched, M unbranched, CuA 2 branched. Metatibial spur large (Fig. 1 D, 0.31 mm), tectiform and smooth, bearing 24 – 32 fine, black-tipped teeth on trailing edge. Basitarsus with 7 apical spinules, arranged 5 + 2, second tarsomere with row of 4 spinules. Abdomen. Male terminalia (Figs 2, 4). Pygofer in lateral view (Figs 2 B, 4 A) roughly quadrate, narrower dorsally than ventrally; anterior and caudal margin truncate, caudal margin without teeth or processes. In caudal view (Fig. 2 A), pygofer opening with smoothly rounded margins; diaphragm developed but not strongly sclerotized; opening for gonostyli small, approximately oval; armature projection parallel sided, rectangular and weekly scoop-shaped. Aedeagus (Figs 2 B, 4 B) tubular, gradually tapering distally, curved ventrally at midlength, with an acuminate process dorsally originating just past apex of curve; gonopore appearing apical. Subtending the aedeagus is a conspicuous process, ~ 2 / 3 length of aedeagus; sinuate, widest at base narrowed to an acuminate apex. Gonostyli weakly flattened and rather parallel sided (Fig. 4 C); basal angles small, diverging basally, parallel near midlength and abruptly converging near apex to broad apices, weakly convex in caudal view; in lateral view (Figs 2 B, 4 A) apices appearing hooked ventrally. Anal segment small, rather truncate in lateral view, processes absent, dorsocaudal angle (in lateral view) not projecting); anal column nearly as elongate as height of anal segment.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	discussion	Remarks. Caldwell & Martorell (1951) remarked that all the specimens of this species were collected at above 2000 ’ elevation. Few specimens of this species are reported and none of these with host plant associations.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBC287B45D3226222569852.taxon	materials_examined	Type material examined. “ Toro Negro Mts / 11 - 14 - 47 P. R. // HOLOTYPE // Abbrosoga / errata / Cald. / [barcode label UDCC _ TCN 00094754] ” (1 male, dissected, USNM); “ Maricao / P. R. / July 27, 1914 // PARATYPE / Columbisoga / errata n. sp. / Caldwell / [barcode label] UDCC _ TCN 00016869 ” (1 male, AMNH). Other material examined. Puerto Rico, Jayuya, Puntita, July 1962, J. Maldonado C (1 male, dissected, USNM); HWY 120, km 14, Maricao St [ate]. For [est]., VIII- 8 - 1999, C. W. O’Brien, P. W. Kovarik (1 male, dissected, UDCC).	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	description	(Figs 3, 5, 6)	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	materials_examined	Type locality. Puerto Rico, Mayagüez, 18 ° 13. 11 N, 67 ° 08.86 W.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	diagnosis	Diagnosis. This species can be separated from A. errata by having its thorax without median vitta and scutellum dark (Fig. 3 A). Forewing clear between apices of Sc and RA (Fig. 3 C). Frons broad (Fig. 3 B, frons l: w ratio ~ 1: 1.4 – 1.5). Male gonostyli distally without ventral hook in lateral view (Fig. 3 E, distally acute in caudal view); aedeagus (Fig. 5 B) latitudinally subdivided into 3 processes plus subtending process; anal segment with dorsocaudal projection in lateral view.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	description	Description. Color. General color light brown (paler ventrally), eyes dark reddish (Fig. 1 A). Pronotum with small dark spot on each side of median carinae, scutellum darker than scutum. Macropterous forewings (Fig. 3 C) clear with cubital area distally diffusely fuscous, with an elongated spot at the apex of the clavus, hind wings clear. Structure. Body length 3.39 – 3.63 mm including wings (~ 2.23 mm without wings). Head. Head (dorsal view, including eyes) narrower than pronotum (Fig. 3 A), in lateral view (Fig. 3 C) slightly projected anteriorly. Vertex with conspicuous carinae (median carina weak but evident), vertex subquadrate, length 0.23 – 0.26 mm, width 0.22 – 0.25 mm posterior margin truncate. Frons broad (Fig. 3 B), widest near ventral margin of eyes; length 0.46 mm, width (at widest 0.31 – 0.32; l: w ratio 1: 1.4 – 1.5), narrowed both dorsally and toward frontoclypeal suture; median carina forking near fastigium. Clypeus as ¾ as long as frons, bearing median carina. Antennal scape small about as long (0.08 – 0.10 mm) as wide; pedicel about 2 x length of scape (0.19 – 0.21 mm), bearing rows of rhinaria and small setae, flagellum elongate and bristle-like, about 2 x length of pedicel. Thorax. Pronotum shorter than vertex (length at midline 0.17 – 0.20 mm); median carina distinct, lateral carinae diverging, nearly reaching posterior margin; posterior margin weakly concave. Mesonotum at midline about 3 x length of pronotum (0.59 – 0.60 mm); lateral carinae slightly sinuate and weakly diverging, reaching posterior margin, median carina reaching posterior margin of scutellum. Wings macropterous, exceeding abdomen (forewing 2.67 – 3.01 mm); forewing venation (Fig. 6) very similar to A. errata except CuA 3 - branched. Nodal line appearing obsolete near CuA (not reaching wing margin). Metatibial spur large (0.33 – 0.34 mm), tectiform and smooth, bearing 27 – 30 fine, black-tipped teeth on trailing edge. Abdomen. Male terminalia (Figs 3 E, F, 5). Pygofer, in lateral view, roughly quadrate, much narrower dorsally than ventrally; anterior margin weakly concave, caudal margin weakly convex, without teeth or processes. In caudal view, pygofer opening with smoothly rounded margins; diaphragm developed, not strongly sclerotized; opening for gonostyli small, approximately oval. Diaphragm armature caudally projected with paired processes dorsally and midventral carina extending to opening at gonostyli. Aedeagus (Figs 3 E, 5 B), in lateral view, downcurved, with 2 elongate dorsal processes and a single elongate sinuate ventral process about 2 / 3 length of aedeagus; gonopore apparently at apex of aedeagus. Gonostyli, in caudal view (Figs 3 F, 5 A), weakly flattened, broad at base with basal angle blunt and distinct, distally rather forceps-like, converging to acute apex (in widest view), without ventral hook in lateral view. Anal tube small, rather truncate in lateral view, ventrocaudal processes absent, dorsocaudal angle (in lateral view) elongately projected; anal column about 2 / 3 as elongate as height of anal tube.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	discussion	Remarks. Abbrosoga multispinosa externally is strongly similar to A. errata, although it is slightly more robust, with a broader vertex and frons. We note that the scutellum is dark in A. multispinosa (thorax without a pale median vitta), and pale in A. errata (thorax bearing a narrow pale median vitta), and the forewing in A. errata is infuscated near between the apices of the Sc and the RA, but not in A. multispinosa, at least among the specimens we have examined to date. The most definitive diagnostic features is the multiple-subdivided aedeagus of A. multispinosa as opposed to the simple aedeagus of A. errata.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	etymology	Etymology. The species name is derived from the Latin word “ multus ” (much, more), truncated and joined with an - i to the Latin word “ spinosus ” (thorny), given a feminine termination to match Abbrosoga. The name is a reference to the form of the aedeagus.	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
171087DBFFBA287645D3228226AB9EFD.taxon	materials_examined	Type material examined. “ PUERTO RICO / Mayagüez / 18 ° 13.11 N 67 ° 08.86 W / 5 - 6. X. 2011 / Ex: Blacklight / Coll. A. Segarra // [red paper] HOLOTYPE / Abbrosoga / multispinosa Otero & Bartlett 2018 // UDCC _ TCN 00058470 ” (1 male, dissected, USNM). Paratype: same as holotype, except ‘ paratype’ indicated and barcode label UDCC _ TCN 0 0 0 58469. ...... continued on the next page	en	Otero, Miriel, Bartlett, Charles R. (2019): A New Species of Abbrosoga (Hemiptera: Fulgoroidea: Delphacidae), An Endemic Puerto Rican Planthopper Genus, with an Updated Checklist of the Delphacidae of Puerto Rico. Zootaxa 4563 (2): 372-386, DOI: 10.11646/zootaxa.4563.2.10
