taxonID	type	description	language	source
173187E9DD12FFCF265A4E81FD9860D2.taxon	description	Although Physocephala species are among some of the most conspicuous and beautiful Conopidae the identifi- cation of species is very difficult due to high infraspecific variation and the lack of stable characters to divide the genus in groups. Key 1 presents a new attempt to distin- guish species-groups but we are very aware that this re- mains problematic. In many cases it will not be possible to identify single specimens of difficult species without comparative material. Much patience is necessary, and no little frustration encountered, when identifying Afro- tropical Physocephala! It should be noted that the species-groups given below are not intended to represent natural monophyletic groupings but are merely an attempt to provide a starting point for the identification of species within this large ge- nus. Where our species-groups are identical to those pre- viously identified by Camras (2001) we have retained the same species-group name, but where they are constituted or interpreted differently we have used a different name in order to minimise confusion in the future.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD15FFCB267F4B92FB8D6187.taxon	description	(Figs 1 – 5)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD15FFCB267F4B92FB8D6187.taxon	description	= Conops patelliformis Séguy, 1933	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD15FFCB267F4B92FB8D6187.taxon	materials_examined	Primary type material examined. 1 ♂ syntype of Physocephala fascipennis Brunetti, 1925: (1) “ Syn - / type ”; (2) “ 12. II. 11 / Caia / Zambesi / Dhawar / H. Swale. ”; (3) “ Pres. by / Impl. Bureau Ent. / 1915 - 164 ”; (4) “ Physo. / fascipennis / Brun. Type ♂ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249047 ”; coll. NHML. 1 ♀ syntype of Physocephala fascipennis Brunetti, 1925: (1) “ Syn - / type ”; (2) “ Nyasaland / Cholo / R. C. Wood ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physo. / fascipennis / Brun. Type ♀ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249048 ”; coll. NHML. Additional material. BENIN: 1 ♀, vi. 2002, Pénéssou- lou, forest area [09 ° 15 ’ 58.26 ″ N 01 ° 33 ’ 04.81 ″ E], leg. G. Goergen, coll. IITA; 1 ♂, 28. i. 2018, Togbota, leg. G. Go- ergen, coll. IITA; DEMOCRATIC REPUBLIC OF CON- GO: 1 ♂, iv. 1913, Congo da Lemba, det. as P. fascipennis by Brunetti 1925, leg. R. Mayné, coll. MRAC; 1 ♂, 1942, Bas-Congo, Mayidi, leg. P. Van Eyen, coll. ISNB; 1 ♀, 1945, ditto; ETHIOPIA: 1 ♀, iii. 1969, Bahir Dar, leg. Schäuffele, coll. SMNS; 2 ♂♂, 30. – 31. v. 2015, Sof Umer [6 ° 54 ′ N 40 ° 51 ′ E], 1200 m, leg. J. Halada, coll. CULSP; KENYA: 1 ♂, xi. 1948, Garissa-Bura, Tana River, det. as D. abdominalis by Camras 1999, leg. van Someren, coll. NHML [NHMUK 010922084]; 1 ♂, 12. x. 1998, Morigat, leg. F. Kaplan, A. Freidberg, coll. TAUI; 1 ♀, 12. – 26. ii. 2005, Nyanza Province, Ungoye Field Station [0 ° 36.91 ′ S 34 ° 05.52 ′ E], 1147 m, leg. R. Copeland, coll. NMKE; MALAWI: 1 ♂, 11. v. 1916, Nyasaland, Ruo Val- ley, 1000 – 2000 ft, des. as syntype of P. fascipennis, leg. R. C. Wood, coll. NHML [NHMUK 010922076]; MO- ZAMBIQUE: 1 ♀, 19. – 30. iv. 2015, Sofala pr., Gorongosa Park, small lake [18 ° 56 ′ 39 ″ E 34 ° 26 ′ 35 ″ E], 30 m, Mal- aise trap, leg. M. Hauser, A. Rung, coll. PMHA; SOUTH AFRICA: 1 ♂, i. 2000, Mondl Forest, Hilton, KZN, leg. Lyawb, coll. PMHA; 1 specimen, i. – iii. 1927, Natal, Weenen, det. as D. abdominalis by Kröber 1930, leg. H. P. Thommaset, coll. NHML [NHMUK 010922090]; 1 ♀, xii. 1923, Natal, Weenen, det. as D. fascipennis by Brunetti 1924, leg. H. P. Thommaset, coll. NHML [NHMUK 010922093]; 1 specimen, 2. x. 1915, Umbilo, Durban, Natal, des. as syntype of P. fascipennis, [col- lector unknown], coll. NHML [NHMUK 010922077]; 1 ♂, iv. 1955, Cape Province, Grahamstown, det. as D. fascipennis by Smith 1957, leg. Allison, coll. NHML [NHMUK 010922088]; 1 ♂, 26. xi. 1958, Cape Province, Grahamstown, leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, 2. xii. 1958, ditto; 1 ♀, 17. xi. 1961, Cape Province, Grahamstown, leg. E. McC Callan, coll. AMGS; 1 ♂, 19. xii. 1971, Cape Province, Grahamstown, det. as D. abdominalis by Camras 2000, leg. D. J. Greathead, coll. NHML [NHMUK 010922085]; 1 ♀, 26. ii. 1987, Cape Province, Grahamstown, leg. P. E. Hulley, coll. AMGS; 1 ♂, 7. vii. 2002, Cape Province, Grahamstown [33.17 ° S 26.31 ° E], leg. H. V. Lewis, coll. AMGS; 1 ♀, 15. ix. 2006, Cape Province, Grahamstown [33 ° 23 ′ S 26 ° 29 ′ E], leg. A. McClure, coll. AMGS; 1 ♂, 4. iv. 1981, Cape Province, Grahamstown [33 ° 19 ′ S 26 ° 31 ′ E], leg. G. T. Lloyd, coll. AMGS; 1 ♀, 10. iii. 1981, Cape Province, Grahamstown, 200 Dept. window [33.19 S 26.32 E], leg. E. Nieman, coll. AMGS; 2 ♂♂, 22. i. 1970, Cape Province, Grahamstown, Belmont Valley, on flowering Foeniculum vulgare Mill., leg. F. W. Gess, coll. AMGS; 2 ♂♂, 23. i. 1970, Cape Province, Grahamstown, Belmont Valley, on flowering Foeniculum vulgare Mill., leg. F. W. Gess, coll. AMGS; 2 ♂♂, 26. i. 1970, Cape Province, Grahamstown, Belmont Valley, on flowering Foeniculum vulgare Mill., leg. F. W. Gess, coll. AMGS; 1 ♂ 1 ♀, 25. i. 1972, Cape Province, Grahamstown, Belmont Valley, leg. F. W. Gess, coll. AMGS; 1 ♂, 6. i. 1977, Cape Province, Grahamstown, Hil- ton [- 33.310629 26.525595], on Acacia karroo flowers, leg. D. W. Gees, coll. AMGS; 1 ♂, 2. xii. 1979, Cape Prov- ince, Grahamstown, Hilton [- 33.310629 26.525595], leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 6. – 14. i. 1972, Cape Province, Grahamstown, Howison´s l′oort, leg. F. W. Gess, coll. AMGS; 1 ♀, 19. – 22. xi- 1971, ditto; 1 ♂, 8. – 9. iv. 1979, Cape Province, Salt Vlei, Port Alfred, leg. A. E. Mel. Collan, coll. AMGS; 1 ♂, 19. xi. 1999, East- ern Cape Province, 37 km nw of Steytlerville [33 ° 11 ′ S 24 ° 10 ′ E], 695 m, leg. M. Hauser, coll. PMHA; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as D. abdominalis by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922087]; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as D. abdominalis by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922089]; 1 ♀, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as D. abdominalis by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922092]; 1 ♀, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as D. abdominalis by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922094]; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as P. platycephala by Kröber, 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922157]; 1 ♂, 26. ix. 2010, Free State Province, Bloemfontein, 10 Jan Venter Street [29 ° 06 ′ 12 ″ S 26 ° 08 ′ 42 ″ E], leg. R. J. Nuttall, coll. BMSA; 1 ♀, 26. ix. 2010, Free State Prov- ince, Bloemfontein, 10 Jan Venter Street [29 ° 06 ′ 12 ″ S 26 ° 08 ′ 42 ″ E], on window, leg. R. J. Nuttall, coll. BMSA; 1 ♂, xii. 2009, Free State Province, Bloemfontein, Nation- al Museum Library [29 ° 6 ′ 54.35 ″ S 26 ° 13 ′ 9.72 ″ E], leg. L. Coetzee, coll. BMSA; 1 ♀, 13. xii. 2012, KwaZulu-Natal Province, Pietermaritzburg, Kwela Lodge, 940 m, leg. B. Lechner, coll. PASS; 1 ♂, 22. – 25. xi. 2003, Mpumalanga Province, 30 km ne Lydenbrug near Ohrig, leg. J. Hala- da, coll. CULSP; 1 ♀, no date, Natal Province, Howick, des. as syntype of P. fascipennis, leg. J. P. Gregor, coll. NHML [NHMUK 010922078]; 1 ♂, 14. i. 1953, Natal Province, St. Michaele, det. as D. fascipennis by Smith 1957, on flowers of Scutia myrtina, leg. E. McC Callan, coll. NHML [NHMUK 010922086]; 1 ♂, 28. iv. 2002, Western Cape Province, Nature Valley, 33 23 DC, leg. L. de Wet, coll. AMGS; TANZANIA: 1 ♂, 10. v. 1956, Old Shinyanga, Block 9, det. as P. caenoneura by Emden 1950, leg. E. Burtt., coll. NHML [NHMUK 010922143]; ZIMBABWE: 1 ♀, 13. iii. 1936, Salisbury [= Hara- re] [17.84 31.05], leg. W. L. Williams, coll. NHML [NHMUK 010922091]; 1 ♂, 24. vii. 1958, Salisbury [= Harare] [17.84 31.05], leg. A. D. Graham, coll. AMGS; 1 ♀, ii. 1987, Zambesi Valley, Rekometjie [16 ° 10 ′ S 29 ° 25 ′ E], leg. S. Gussmann, coll. ZFMK; UNKNOWN LOCATION: 1 ♀, 27. xii. 1937, [characters illegible], det. as D. abdominalis by Kröber 1938, leg. A. I. Bevis, coll. NHML [NHMUK 010922095].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD17FFC4267F4897FED7600B.taxon	description	(Fig. 6)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD17FFC4267F4897FED7600B.taxon	materials_examined	Material. CENTRAL AFRICAN REPUBLIC: 1 ♀, 17. xii. 2008, 45 km e Nola [03 ° 40 ′ N 16 ° 26 ′ E], 570 m, leg. J. Halada, coll. CULSP; DEMOCRATIC REPUB- LIC OF CONGO: 1 ♂, 1968, Tshuapa, Bamanya, leg. P. Hulstaert, coll. MRAC; ETHIOPIA: 1 ♀, 4. xi. 2018, Bahir Dar [11.597292 ° N 37.355696 ° E], leg. G. Goergen, coll. IITA; 1 ♀, 11. xii. 2014, Bonga, Strassensaum, 1955 m [07 ° 14 ′ 09 ″ N 036 ° 16 ′ 36 ″ E], leg. H. - J. Flügel, coll. PHJF; 1 ♂, 20. ix. 2012, Chencha, 1916 m [06 ° 09 ′ N 37 ° 34 ′ E], leg. A. Pauly, coll. ISNB; 1 ♂, 1912, Harar, leg. S. V. Kris- tensen, coll. ZMHB; 1 ♀, 18. x. 1957, near Axum, paratype of P. kaplanae, det. as D. abdominalis by Smith 1969, leg. D. J. Greathead, coll. NHML [NHMUK 010922075]; GABON: 1 ♂, 11. – 12. iii. 1990, Wolen-Ntem, Monts de Christal, Tchimbelé, 600 m [00 ° 37 ′ N 10 ° 24 ′ E], leg. J. J. Wieringa, coll. RMNH. It is possible that P. kaplanae is nothing more than a darker form of P. abdominalis, although we are not aware of any intermediates and therefore accept the validity of this species.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD18FFC2265A4E86FDE8611B.taxon	description	(Figs 7 – 10) Conops antiqua Wiedemann 1830 = Conops maculipes Bigot, 1887 syn. nov. = Conops interrupta Bezzi, 1901 = Conops erythraspis Bezzi, 1901	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD18FFC2265A4E86FDE8611B.taxon	description	= Physocephala brevistylata Kröber, 1931 = Physocephala minutissima Kröber, 1933 syn. nov. = Physocephala nigroscutellata Kröber, 1933	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD18FFC2265A4E86FDE8611B.taxon	materials_examined	Primary type material examined. ♀ holotype of Conops maculipes Bigot, 1887: (1) “ Holo - / type ”; (2) “ S. Africa: / Cape of Good Hope. / ex coll. J. Bigot / ex coll. G. H. Verrall / B. M. 1914 - 500 [strikethrough] ”; (3) “ C. maculigera ♀ / Cap. B. Hop. ”; coll. NHML. ♂ holotype of Physocephala rubicunda Kröber, 1915: (1) ” 3055 ”; (2) “ Type ”; (3) “ Physocephala ♂ / rubicunda Krb. / O. Kröber det. 1914 ”; coll. ZMHB. ♂ syntype of Physocephala decisa Brunetti 1925: (1) “ Syn- / type ”; (2) “ N. E. Rhodesia. / Upper Luangwa R. / 27 July 13 Aug. 1910. / S. A. Neave ”; (3) “ Physocephala / decisa / Brun. Type ♂ / Det. E. Brunetti 1924 ”; (4) “ BMNH (E) # / 249074 ”; coll. NHML. ♀ syntype of Physocephala decisa Brunetti 1925: (1) “ Syn- / type ”; (2) “ N. E. Rhodesia. / Upper Luangwa R. / 27 July 13 Aug. 1910. / S. A. Neave ”; (3) “ Physocephala / decisa / Brun. Type ♀ / Det. E. Brunetti 1924 ”; (4) “ BMNH (E) # / 249077 ”; coll. NHML. ♀ syntype of Physocephala decisa Brunetti 1925: (1) “ Syn- / type ”; (2) “ Rusapi / S. Rhodesia / 19.12.1920 / Rhodesi / Museum ”; (3) “ Pres. by. / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physocephala / decisa / Brun. / Type ♀ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249075 ”; coll. NHML. ♂ holotype of Physocephala minutissima Kröber, 1933: (1) “ 1 ” “ Holo- / type ”; (2) “ Type ”; (3) “ 8.6.16 / Ruo / Nyasaland / 200 ft / R. C. Wood / 601. ”; (4) “ Pres. by. / Imp. Inst. Ent. / Brit. Mus. / 1931 - 480 ”; (5) “ Physocephala / minutissima / ♂ Krb ”; (6) “ BMNH (E) # / 249071 ”; coll. NHML. ♀ holotype of Physocephala nigroscutellata Kröber 1933: (1) “ Holo- / type ”; (2) “ Type ”; (3) “ Lady Grey / 1 Feb. 1924 / R. I. Nel ”; (4) “ Pres. by / Imp. Inst. Ent. / Brit. Mus. / 1932 - 143. ”; (5) “ Phsocephala / nigroscutella [sic] / ♀ Krb / det. Kröber 1931 ”; (6) “ BMNH (E) # / 249072 ”; coll. NHML. Additional material. BENIN: 1 ♀, ix. 2006, Adohoun [06 ° 39 ’ 27.55 ″ N 01 ° 39 ’ 09.76 ″ E], light trap, leg. G. Goer- gen, coll. IITA; 1 ♂, v. 2007, Athiémé [06 ° 14 ‘ 20.00 ″ N 01 ° 40 ‘ 00.00 ″ E], leg. G. Goergen, coll. IITA; 1 ♂, viii. 2006, Athiémé [06 ° 14 ’ 20.00 ″ N 01 ° 40 ’ 00.00 ″ E], sweep netting, leg. G. Goergen, coll. IITA; 1 ♀, 14. vi. 2006, Calavi, sweep netting, leg. G. Goergen, coll. IITA; 1 ♂, 7. viii. 2006, Calavi, campus [06 ° 26 ’ 15.00 ″ N 02 ° 19 ’ 42.00 ″ E], sweep netting, leg. G. Goergen, coll. IITA; 1 ♀, 25. x. 2007, ditto; BURUNDI: 1 ♀, 24. viii. 1957, Kisenyi, leg. F. J. François, coll. ISNB; 1 ♀, 22. i. 1950, Rumonge, 780 m, det. as P. nigroscutellata by Janssen 1954, leg. F. J. François, coll. ISNB; 1 ♂, 9. iii. 1952, Bubanza Province, 7 km s de Gihanga, 850 m, det. as P. nigroscutellata by Janssen 1954, leg. F. J. François, coll. BMSA; CAMEROON: 1 ♀, 24. iv. 1914, Uamgebiet, Bossum, leg. G. Tessmann, coll. ZMHB; 1 ♀, 12. v. 1914, Uamgebiet, Bossum, det. as P. interrupta by Kröber, leg. G. Tessmann, coll. ZMHB; CENTRAL AFRICAN RE- PUBLIC: 1 ♀, 14. v. 2009, 45 km ssw Bamingui [07 ° 15 ′ N 20 ° 03 ′ E], leg. J. Halada, coll. CULSP; 1 ♂, 20. iv. 2010, Reserve Koukorou Bamingui [07 ° 15 ′ N 20 ° 03 ′ E], 440 m, leg. J. Halada, coll. CULSP; DEMOCRATIC REPUB- LIC OF CONGO: 1 ♀, i. – ii. 1913, Congo da Lemba, det. as P. decisa by Brunetti 1925, leg. R. Mayné, coll. MRAC; ETHIOPIA: 1 ♂, 24. v. 2015, 10 km nw Mega [04 ° 08 ′ N 38 ° 16 ′ E], 1670 m, leg. J. Halada, coll. CULSP; 8 ♂♂, 11. – 13. v. 2015, 20 km se Konsa [05 ° 15 ′ N 37 ° 32 ′ E], 850 m, leg. J. Halada, coll. CULSP; 1 ♂ 2 ♀♀, 14. v. 2015, 40 km w Konso [5 ° 19 ′ N 37 ° 04 ′ E], 600 m, leg. J. Halada, coll. CULSP; 1 ♂, 10. v. 2015, 45 km ne Arba Minch [6 ° 17 ′ N 37 ° 47 ′ E], 1200 m, leg. J. Halada, coll. CULSP; 1 ♀, 21. iv. 2016, Arsi [7 ° 49 ′ 06 ″ N 40 ° 31 ′ 52 ″ E], 886 m, leg. J. Halada, coll. CULSP; 1 ♀, 22. v. 2015, Wachile env. [04 ° 32 ′ N 39 ° 03 ′ E], 1070 m, leg. J. Halada, coll. CULSP; GAMBIA: 1 ♂, 4. xi. 1977, Bakau at tropic Bungalow [UTM 28 PCK 1790], swept in meadow rich in flowers, at the beach, leg. Cederholm, Daniellson, Hammarstedt, Hedqvist, Samuelsson, coll. MZLU; 1 ♀, 16. x. 1999, Ki- ang West National Park, headquarter, leg. W. Schacht, coll. ZSMC; IVORY COAST: 1 ♀, 9. xi. 1983, Badenous Mbingué [9.50 ° N 5.50 ° W], leg. R. Summkeller, coll. ZFMK; KENYA: 1 ♂, 27. iv. 2008, Nguni, n of Ngomeni, hand net, leg. M. Snížek, coll. PMME; 2 ♂♂, 29. iv. 1995, Tsavo East, leg. R. Copeland, coll. NMKE; 1 ♂, 10. vi. 1998, Tsavo East National Park, near Athi River [2 ° 37 ′ S 38 ° 22 ′ E], leg. R. Copeland, coll. NMKE; 1 ♂, 22. v. 2006, Coast Province, Tsavo East National Park, near Galana River [UTM UTM 37 m 503352 9665277], 246 m, hand net, leg. P. Cerretti, D. Avesani, G. Carpan- eto, G. Nardi, coll. PMME; 1 ♀, 26. – 29. vi. 1999, Eastern Province, at Athi river [2 ° 38.51 ′ S 38 ° 21.98 ′ E], Malaise trap, leg. R. Copeland, coll. IITA; 1 ♂, 17. – 31. viii. 2005, Nyanza Province, Ungoye, ICIPE Field Station [0.61325 ° S 34.08908 ° E], 1127 m, leg. R. Copeland, coll. NMKE; 1 ♂, 26. vi. – 10. vii. 2007, Rift Valley Province, Sumburu Nature Reserve, near Ewaso Ng′iro River [0.56797 ° N 37.53563 ° E], 874 m, Malaise trap, riverine forest next to headquarter, leg. R. Copeland, coll. NMKE; 1 ♂, 12. – 26. vi. 2006, ditto; 1 ♂, 10. iv. 1998, Samburu Dis- trict, Samburu Serena Lodge, leg. R. Copeland, coll. NMKE; LESOTHO: 1 ♂, 1. xi. 1949, Mamathes [- 29.136617 27.845796], leg. C. Jacot-Guillarmod, coll. AMGS; MADAGASCAR: 1 ♀, 8. xi. 2003, Anteninde, Sakahara [22 ° 53 ′ 57 ″ S 44 ° 28 ′ 12 ″ E], 395 m, leg. A. Ssy- mank, coll. PASS; 1 ♂, v. 1937, Bekily, leg. A. Seyrig, coll. MNHN; 1 ♂, iv. 1937, ditto; 1 ♂, v. 1937, ditto; 1 ♂ 1 ♀, iv. 1942, Bekily, det. as P. madagascariensis by Camras 1960, leg. A. Seyrig, coll. MRAC; 2 ♂♂, 9. iv. 1994, Berenty reserve 80 km w of Port-Dauphin [25 ° 00 ′ S 46 ° 18 ′ E], leg. M. Wasbauer, coll. UCDC, PMHA; 1 ♀, 6. xi. 2003, Ihasofotsy-Flussufer, ca 10 km wsw Ihosy [22 ° 25 ′ 16 ″ S 46 ° 00 ′ 14 ″ E], 1000 m, leg. A. Ssymank, coll. PASS; 1 ♂, 19. iii. 1994, Isalo, Analalava, leg. A. Pauly, coll. MRAC; 1 ♀, 25. ii. – 3. iii. 1968, Majun- ga s. l., det. as P. madagascariensis by Camras 2000, leg. K. M. G. & P. D., coll. NHML [NHMUK 010922098]; 1 ♂, i. 1992, Morarano-Chrome [17 ° 45 ′ S 47 ° 59 ′ E], leg. A. Pauly, coll. MRAC; 1 ♂, no date, Sambirano, leg. [?], coll. ZMHB; 1 ♀, no date, Tananarivo, det. as P. madagascariensis by Camras 1960, [collector unknown], coll. MRAC; 1 ♀ 1 specimen, i. 1952, Tzimbazaza-Tananarive, det. as P. madagascariensis by Camras 1960, leg. R. Benoist, coll. MRAC; 1 ♀, 28. vii. – vi. viii. 2001, Di- ego-Suarez (Antsiranana), Sakalava Beach, malaise across sandy trail in dwarf littoral forest [12 ° 15 ′ 46 ″ S 49 ° 23 ′ 51 ″ E], 10 m, leg. R. Harin′Hala, coll. CAS; 1 ♂, 4. i. 2007, Mahayanga Province, 20 km nw Borinziny [15 ° 27.07 ′ S 47 ° 36.85 ′ E], 37 m, hand netted in tropical dry forest on white sand, leg. M. E. Irwin, F. D. Parker, R. Harin′Hala, coll. CAS; 1 ♀, 1. xi. 1993, Tamatave (Toama- sina), Analmalotran Tamatave, leg. C. F. Kassebeer, coll. PCFK; 1 ♀, 12. xii. 1991, Tuléar (Toliara), Ampanihy, leg. A. Pauly, coll. MRAC; 1 ♀, 6. – 16. i. 2003, Tuléar (Toli- ara), Andohaela National Park, Parcel II, Tsimela, mal- aise trap in transitional forest, 175 m [24 ° 56.21 ′ S 46 ° 37.60 ′ E], leg. M. E. Irwin, F. D. Parker, R. Harin′Ha- la, coll. CAS; 1 ♂, 12. iv. 1968, Tuléar (Toliara), Bevilany, 300 m, det. as P. madagascariensis by Camras 2000, leg. K. M. G. & P. D., coll. NHML [NHMUK 010922097]; 1 ♀, 8. – 18. vi. 2002, Tuléar (Toliara), Beza Mahafaly Re- serve, malaise in dry deciduous gallery forest, 165 m [23 ° 41.19 ′ S 44 ° 35.46 ′ E], leg. R. Harin′Hala, M. E. Irwin, coll. CAS; MALI: 1 ♀, 20. viii. 1991, 10 km s Mopti, leg. M. Schwarz, coll. PMHA; 1 ♂, 20. viii. 1991, 10 km s Mopti, sweep netting, leg. M. Schwarz, coll. PMHA; 1 ♂, 31. vii. 1991, 40 km sw Segou, sweep netting, leg. M. Schwarz, coll. PMHA; MOZAMBIQUE: 2 ♂♂, 12. – 20. xii. 2003, Manica Province, 45 km nw Chimoio, leg. J. Halada, coll. CULSP; NAMIBIA: 1 ♀, 28. i. 1993, 100 km sw Rundu, leg. M. Schwarz, coll. PMHA; 2 ♀♀, 16. i. 1993, 125 km sw Rundu, leg. M. Schwarz, coll. PMHA; 2 ♀♀, 16. i. 1993, ditto; 2 ♂♂, 4. ii. 1993, 15 km e Swakopmund, sweep netting, leg. M. Schwarz, coll. PMHA; 2 ♂♂, 29. iii. 2000, 18 km se Stamprieton C 15 to Gochas [24.28 ° S 18.30 ° E], visiting pink flowers of Galenia, leg. F. W. Gees, S. K. Gees, coll. AMGS; 2 ♂♂, 4. ii. 1990, 30 km e Windhoek, sweep netting, leg. M. Schwarz, coll. PMHA; 1 ♂, 12. iii. 2014, 35 km w Gobabis [22 ° 23 ′ S 18 ° 39 ′ E], 1480 m, leg. J. Halada, coll. CULSP; 1 ♂, 16. ii. 1990, 40 km w Witvlei, leg. M. Schwarz, coll. PMHA; 1 ♂, 16. ii. 1990, 40 km w Witvlei, sweep netting, leg. M. Schwarz, coll. PMHA; 1 ♂, 11. iii. – 9. iv. 1985, 6 km n Arandis, Damaraland [22 ° 22 ′ S 14 ° 59 ′ E], leg. J. Irish, H. Rust, coll. SMWN; 1 ♀, 21. i. 1993, 60 km E Rundu, leg. M. Schwarz, coll. PMHA; 1 ♂, 13. ii. 1990, 73 km s Mariental, sweep netting, leg. M. Schwarz, coll. PMHA; 3 ♀♀, 25. ix. 1997, e Oranjemund 28 km from checkpoint to Sendelingsdrif [29.26 ° S 16.42 ° E], on yel- low flowers of Deverradenudata (Viv.) Pfisterer & Podl., leg. F. W. Gees, S. K. Gees, coll. AMGS; 2 ♂♂, 20. xii. 1995, Etosha National Park, 50 km ne Okaukuejo, “ Salvado ” [18.9 ° S 16.5 ° E], leg. C. Schmid-Egger, coll. PMHA; 1 ♀, 16. – 19. i. 2007, Gobabeb, Kuiseb-Tal, at light, leg. Mey, Ebert, coll. ZMHB; 1 ♀, 11. ii. 1993, Oka- handja, leg. M. Schwarz, coll. PMHA; 1 ♀, 7. iii. 1999, Karas Mountains, 6 km s on 201 from 26 [27.09 ° S 19.01 ° E], on yellow flowers of Vahlia capensis (L. f.) Thunb., leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 25. iv. 1972, Karasburg, Warmbad, SE 2818 Ba, [collector unknown], coll. SMWN; 1 ♂, 17. – 19. v. 1978, Khowarib, R., SE 1914 Ac, Kackoland, leg. S. Louw, M. - L. Penrith, coll. SMWN; 1 ♂, 12. – 14. iv. 1974, Mukorob 14, Namaland, SE 2918 Ac, [collector unknown], coll. SMWN; 1 ♂, 15. iii. 1997, near Aus on road to Helmering- hausen [26.37 ° S 19.20 ° E], visiting white flowers of Psilocaulon göareosum (Berger) Dinter & Schwantes, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 10. – 13. ii. 1972, Otjikoko-Sud 61, Omaruru, SE 2116 Ad, [collector un- known], coll. SMWN; 1 ♂, 4. – 5. iii. 1972, Plateau 38, Luderitz, SE 2616 Cb, [collector unknown], coll. SMWN; 1 ♂, 27. iv. 1972, Rotegab 95, Keetmanshoop, SE 2718 Ad, [collector unknown], coll. SMWN; 1 ♀, 11. iv. 1998, Swakop River bed on road to Goanikontes [22.41 ° S 14.35 ° E], on white flowers of Psilocaulon salicornioides (Pax) Schwantes, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 11. iv. 1998, Swakop River bed on road to Goanikontes [22.41 ° S 14.35 ° E], visiting deep pink flow- ers of Gelenia papulosa (Eckl. & Zeyh.) Sond., leg. F. W. Gees, S. K. Gees, coll. AMGS; 2 ♂♂, 11. iv. 1998, Swakop River bed on road to Goanikontes [22.41 ° S 14.35 ° E], visiting white flowers of Psilocaulon salicornioides (Pax) Schwantes, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 11. iv. 1998, Swakop River bed on road to Goanikontes [22.41 ° S 14.35 ° E], visiting yellow flowers of Zygophyllum simplex, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♀, 16. iii. 1999, Swakopmund, Swakop River at bridge [22.42 ° S 14 ° 32 ° E], on white flowers of Zygophyllum stapffii Schinz, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 13. iv. 2002, Walfisbay, Swakopmundvia Dune 7 [22.55 ° S 14.36 ° E], leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♀, 20. – 23. xii. 1974, Windhoek, Wasser- vallei [22 ° 55 ′ S 16 ° 22 ′ E], [collector unknown], coll. SMWN; 1 ♀, 6. – 8. ii. 2001, Gobabis District, Somerkoms 521 [22 ° 01 ′ 59 ″ S 19 ° 57 ′ 22 ″ E], leg. A. H. Kirk-Spriggs, coll. SMWN; 1 ♀, 20. iii. 2014, Harda Province, 15 km s Rehobot [23 ° 28 ′ S 17 ° 07 ′ E], 1400 m, leg. J. Halada, coll. CULSP; 2 ♂♂, 16. ii. 1990, Omaheke Region, Gobabis, 40 km w Witvlei, leg. M. Schwarz, coll. PMHA; 1 ♀, 25. – 27. iii. 2003, Rundu District, Mile 46 [18 ° 18 ′ 39 ″ S 19 ° 15 ′ 29 ″ E], leg. A. H. Kirk-Spriggs, coll. SMWN; SENEGAL: 1 ♂, 27. vi. 2004, 60 km s Velingara, sweep netting, leg. M. Halada, coll. CULSP; SOUTH AFRICA: 1 ♂ 2 ♀♀, 8. – 12. ii. 1982, Sandveld Nature Reserve, Hoop- stad, SE 2725 Da, leg. Entomology Department, coll. BMSA; 1 ♂, xii. 1956, [?] “ Snoot Rivien ”, leg. Martin, coll. AMGS; 1 ♂, 3. – 8. x. 1989, Cape Province, 15 km n of Nieuwoudtville on road to Loeriesfontein, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♀, 4. i. 2010, Cape Prov- ince, 25 km n Jansenville [32 ° 49 ′ S 24 ° 44 ′ E], 600 m, leg. J. Halada, coll. CULSP; 1 ♀, xii. 1922, Cape Province, Aliwal North, det. as P. brevistylatus by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922102]; 1 ♀, 3. xii. 1986, Cape Province, Bloutoring [33 ° 28 ′ 40 ″ S 20 ° 19 ′ 15 ″ E], leg. F. W. Gess, coll. AMGS; 1 ♀, 24. xii. 1960, Cape Province, Grahamstown, leg. E. McC Callan, coll. AMGS; 1 ♀, 27. xi. 1981, Cape Province, Grahamstown, [?] ” Olakwater ”, leg. F. W. Gess, coll. AMGS; 1 ♀, 13. i. 1986, Cape Province, Grahamstown, Clifton, leg. R. W. Gees, coll. AMGS; 1 ♀, 12. – 30. i. 1970, Cape Province, Grahamstown, Hilton [- 33.310629 26.525595], Malaise trap, leg. F. W. Gess, coll. AMGS; 1 ♂, 3. iii. 1978, Cape Province, Grahamstown, Hilton [- 33.310629 26.525595], on flowers of Melolobium candicans in sandpit, leg. F. W. Gess, coll. AMGS; 1 ♂, 3. iii. 1978, Cape Province, Grahamstown, Hilton [- 33.310629 26.525595], leg. S. K. Gess, coll. AMGS; 1 ♂, 2. xii. 1989, Cape Province, Grahamstown, Hilton [- 33.310629 26.525595], leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♀, 22. x. 1981, Cape Province, Graham- stown, Lynton, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 12. ii. 1967, Cape Province, Grahamstown, Strowahn, leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, 19. – 24. iii. 191 [?], Cape Province, Grahamstown, Table Farm, leg. F. W. Gess, coll. AMGS; 1 ♂ 1 ♀, 2. – 13. iii. 2008, Cape Province, Grahamstown, Three Chimneys Farm [33 ° 18.542 ′ S 26 ° 29.846 ′ E], leg. A. H. Kirk-Spriggs, coll. AMGS; 1 ♂, 11. x. 1994, Cape Province, Hamaqual and Kamieskroon / Sors Sors, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 17. – 21. & 24. ix. 1995, Cape Province, Richtersveld National Park, Koeroegabvlakte [28.11 ° S 17.03 ° E], visiting deep pink flowers of Hermbstaetia glauca (Wendl.) Reichb. ex Steud., leg. F. W. Gess, S. K. Gess., R. W. Gess, coll. AMGS; 1 ♂, 15. ix. 1996, Cape Province, Richtersveld, Pachtviel [28.33 ° S 16.34 ° E], leg. F. W. Gess, S. K. Gess., R. W. Gess, coll. AMGS; 1 ♂, 16. xi. 1994, Cape Province, Tierberg [33.10 ° S 22.16 ° E], visiting cream flowers of Asclepias buchenaviana Schinz, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 26. xi. – 5. xii. 1987, Cape Province, Tierberg, research station [33 ° 7 ′ 42 ″ S 22 ° 16 ′ 24 ″ E], leg. F. W. Gess, S. K. Gess., R. W. Gess, coll. AMGS; 1 ♀, 26. xi. – 25. xii. 1987, Cape Province, Tierberg, study site [33 ° 10 ′ S 22 ° 16 ′ 24 ″ E], on flowers of Asclepis buchenaviana Schinz, leg. F. W. Gess, S. K. Gess., R. W. Gess, coll. AMGS; 6 ♂♂, 26. xi. – 5. xii. 1987, Cape Province, Tierberg, study site [33 ° 10 ′ S 22 ° 16 ′ 24 ″ E], visiting flowers of Asclepias buchenaviana Schinz, leg. F. W. Gess, S. K. Gess., R. W. Gess, coll. AMGS; 1 ♀, 1. - 9 - xii. 1988, Cape Province, Vanwyksfontein, 8 km w of Norvalspont, leg. R. W. Gees, coll. AMGS; 1 ♀, 12. i. 2004, Eastern Cape Prov- ince, Grahamstown, [33 ° 16 ′ 00 ″ S 26 ° 28 ′ 60 ″ E], 629 m, leg. C. Hepburn, coll. AMGS; 1 ♂, 10. x. 1992, Eastern Cape Province, Grahamstown, Hoowisonspoort, 3326 BC, leg. T. Doubel, coll. AMGS; 1 ♂, 28. ii. 2004, Eastern Cape Province, Thomas Baines Nature Reserve [33 ° 23 ′ 02 ′ S 26 ° 29 ′ 01 ″ E], leg. C. C. Robertson, coll. AMGS; 1 ♀, no date, Eastern Cape Province, Willow- more, leg. Brauns, coll. PJHS; 1 ♂, xii. 1907, Eastern Cape Province, Willowmore, det. as P. interrupta, leg. H. Brauns, coll. MRAC; 1 ♀, no date, Eastern Cape Prov- ince, Willowmore, det. as P. interrupta, leg. H. Brauns, coll. MRAC; 1 ♂, 25. i. 1902, Eastern Cape Province, Wil- lowmore, leg. Brauns, coll. PJHS; 1 ♀, 25. – 29. x. 2010, Free State Province, Brandfort, Florisbad Res. Stat. [28 ° 46.039 ′ S 26 ° 04.234 ′ E], leg. A. H. Kirk-Spriggs, coll. BMSA; 2 ♀♀, 8. – 12. ii. 1982, Free State Province, Hoop- stad, Sandfeld Nature Reserve, SE 2725 Da, leg. Ento- mology Department, coll. BMSA; 1 ♀, 13. xii. 1995, Mpumalanga Province, Blyderevierspoort NP [24 ° 39 ′ S 30 ° 50 ′ E], leg. F. Koch, coll. ZMHB; 1 ♂, 1. x. 1997, Northern Cape, 60 km e Springbok [29.28 ° S 18.26 ° E], visiting cream flowers of Asclepias buchenaviana Schinz, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 10. x. 2000, Northern Cape, Pachtvlei, e Alexander Bay [28.33 ° S 16 ° 34 ′ E], visiting yellow flowers of Lebeckia, leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 20. xi. 1999, Western Cape Province, 7 km n of Avontuur [33 ° 40 ′ S 23 ° 09 ′ E], sweep netting, leg. M. Hauser, coll. PMHA; 1 ♀, 11. iv. 2006, Western Cape Province, Stellenbosch [33 ° 55 ′ S 18 ° 51 ′ E], leg. E. Mostert, coll. AMGS; SUDAN: 1 ♂, 14. vi. 1996, Blue Nile, Wad Medani, leg. G. G. M. Schul- ten, coll. RMNH; 1 ♂, 1935, Sangha près Bandiagara, leg. M. Griaule, coll. MNHN; TANZANIA: 1 ♀, 16. i. 2007, Pwani Province, 15 km e Utete [08 ° 03 ′ S 38 ° 53 ′ E], 75 m, leg. J. Halada, coll. CULSP; TOGO: 2 ♀♀, 7. iv. 2008, Mono riverside s Ahassomé [7 ° 11 ′ 14 ″ N 01 ° 28 ′ 52 ″ E], 90 m, leg. A. Ssymank, coll. PASS; ZAM- BIA: 1 ♀, 27. vii. – 13. viii. 1910, Upper Luangwa River, des. as syntype of P. decisa, leg. S. A. Neave, coll. NHML [NHMUK 010922107]; 5 ♂♂, 20. v. – 29. vii. 1988, Zambe- si Valley [16 ° 18 ′ S 30 ° 16 ′ E], leg. J. Weyrich, coll. ZFMK; ZIMBABWE: 1 ♂ 4 ♀♀, 18. – 19. xii. 2011, 60 km nnw Bu- lawayo [19 ° 41 ′ S 28 ° 21 ′ E], 1200 m, leg. J. Halada, coll. CULSP; 1 ♀, 14. xii. 1919, Bulawayo, des. as syntype of P. decisa, [collector unknown], coll. NHML [NHMUK 010922108]; UNKNOWN LOCATION: 1 ♀, 24. i. 1928, “ Rersolution ”, leg. A. Walton, coll. AMGS; 1 ♀, 1893, “ Shitlesia ”, leg., coll. AMGS; 1 ♂, no date, “ T 287 ”, [collector unknown], coll. ZFMK; 1 ♂, no date, “ T 426 ”, [collector unknown], coll. ZFMK. Stuke (2015 b) concluded that P. antiqua is an outstandingly variable species and as a result several new synonyms were recognised in the Palaearctic Region. While reviewing the large amount of material available it became evident that most of the commonly occurring Afrotropical Physocephala specimens also belong to the antiqua species-group. It cannot be excluded that factors such as DNA-sequencing, newly discovered morpholog- ical characters or the host ranges of larvae may even- tually necessitate the splitting of P. antiqua as defined herewith but at the present time the consistent separation of distinct species within this highly variable taxon is not possible using the morphological characters which are currently available, or which have been used in the past. We could not find any consistent differences in the male postabdomen or female theca within the various morphs, for example, and the Afrotropical material is highly vari- able in respect of characters such as wing pattern and microtrichiation, size and shape of the vertex, dusting and setulae on the pleura, body size, and colouration of the legs, abdomen and thorax. Dark-winged specimens with the basal cell almost completely covered with mi- crotrichia (although never completely covered!) appear remarkably distinct, but we also found intermediates. Such specimens could potentially be incorrectly assigned to the newly described P. guillarmodi (see below) although the dark brown subcosta of P. antiqua (light orange-brown or yellow in P. guillarmodi) and the more or less black scutellum (orange in P. guillarmodi) are good characters for separating these two species. Physocephala antiqua specimens with a shortened radial-medial crossvein appear very similar to species of the microvena species-group, but always lack a distinct shining in- dention at the hind margin of the eye and have a differ- ent male postabdomen. Specimens in which the pleural dusting stripe is indistinct, such as in old specimens or specimens which have been wetted, could very easily be misidentified as belonging to the vittata species-group.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFC225D8481BFC1E636E.taxon	description	(Fig. 11) Physocephala brevipennis Camras, 1962	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFC225D8481BFC1E636E.taxon	materials_examined	Primary type material examined. ♂ holotype of Physocephala brevipennis Camras, 1962: (1) “ Ngutu, / Zulu- land / III- 21 - 1951 / A. L. Capener ”; (2) “ Physocephala / brevipennis / Camras ”; (3) “ Holotype ♂ / Physocephala / brevipennis / Camras ”; (4) “ FMNHINS / 3130492 / Field Museum / pinned ”; coll. FMNH. The first impression when looking at this strange speci- men is that it is perhaps an aberrant example of Physocephala antiqua with exceedingly shortened wings. Until further material becomes available, however, we have re- tained this taxon.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFC2265A4A2FFA7D60BE.taxon	description	(Fig. 12) Physocephala claripennis Becker 1923	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFC2265A4A2FFA7D60BE.taxon	materials_examined	Primary type material examined. ♂ holotype of Physocephala claripennis Becker, 1923: (1) “ El Obeid / 13. – 14. III ”; (2) “ Aegypt. Sudan / Ebner, 1914 ”; (3) “ Ph. claripenis [sic] / Beck. / det. Becker ”; (4) “ Holotype / Physocephala / claripennis / Becker / det. Camras, 2000 ”; (5) “ J. Skevington / Specimen # / 45425 ”; coll. NMW. This may only be an extremely pale specimen of P. antiqua with completely hyaline wings, but until further material becomes available we have retained this taxon.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	description	(Figs 13 – 24) urn: lsid: zoobank. org: act: 8 F 911 A 66 - CF 42 - 4 A 60 - 837 F- 708 FA 78 B 1 E 9 C	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	materials_examined	Holotype ♀. (1) “ Mamathes / Basutoland / 28 - XII- 1959 / C. Jacot / Guillarmod ”; (2) “? Physocephala brevistylata / Kröber ♀ ”; (3) Holotypus / Physocephala guillarmodi / spec. nov. ♀ / 2018 ”. Holotype is deposited in AMGS. The specimen is pinned and in perfect condition. Paratypes. BURUNDI: 1 ♀, 18. xi. 1949, Bururi Prov- ince, Bururi, 1950 m, Orée dela forȇt, leg. F. François, coll. ISNB; LESOTHO: 1 ♂, 26. xii. 1946, Bokong, leg. A. Jacot Guillarmod, coll. AMGS; 1 ♀, 28. xii. 1947, Leribe, Hensley′s Dam, leg. A. Jacot-Guillarmod, coll. AMGS; 1 ♀, 16. xii. 1950, Mamathes, leg. C. Jacot-Guil- larmod, coll. AMGS; 1 ♂, 28. xii. 1959, ditto; SOUTH AFRICA: 1 ♂, 3. – 7. x. 1988, Cape Province, Clanwilliam Dam [32 ° 11 ′ 30 ″ S 18 ° 53 ′ 42 ″ E], on flowers of Aspalathus desertorum Bol., leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂, 5. xii. 1980, Cape Province, Grahamstown, Hilton, Senecio, leg. D. W. Gees, coll. AMGS; 1 ♀, 8. – 13. x. 1987, Cape Province, Klein Alexandershoek, Clan- william District [32 ° 20 ′ 20 ″ S 18 ° 46 ′ E], leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♂ 1 ♀, 28. ix. 1985, ditto; 1 ♂, 4. – 8. x. 1994, Cape Province, Koornplanskloof 10 km s Citrusdaal [32 ° 40 ′ S 19 ° 01 ′ E], 200 – 270 m, leg. R. Dan- ielsson, coll. MZLU.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	description	Description of holotype (female) Length 9.6 mm; Wing-length 7.6 mm; Head-height 2.9 mm. Head. Antenna orange-brown (Fig. 13). Arista very short stylus-like, with 2 aristomeres situated at tip of first flagellomere (Fig. 16). Both aristomeres minute, length of apical aristomere shorter than height. Scape about twice as long as maximum width, apically and lateral- ly with black setae. Pedicel about five times longer than maximum width, covered with black setae. Pedicel lack- ing any ridge at base, and expanded towards apex. First flagellomere long and conical, about two times as long as high, pointed, and ventrally with an indistinct membra- nous area. Lunule between base of antennae and ptilinal suture distinctly developed, shorter than width of scape. Eyes brown, lacking ommatrichia, facets all of about the same size. Posterior margin of eye lacking any shining indentation. Gena height / eye height (measurements tak- en from head in lateral view) = 0.3. No ocellar tubercle, no ocelli and no ocellar triangle evident. Frons (Fig. 15) yellow, somewhat broader than long, concave, slightly projecting above eyes posteriorly and lacking any setu- lae. Anterior margin of frons concave. No frontofacial spot. Frons with indistinct dusting all over, subshining laterally. Vertex as broad as frons, separated from latter by a more or less distinct ridge. Vertex shining, and in posterior half covered with black setulae. Vertex apical- ly with an indistinct depression. Ridge of vertex lacking longitudinal grooves. Face yellow, with brown facial keel. Face lacks dusting except for some very narrow silver dusting along eye margin. Gena yellow, lacking setae. Distinct facial grooves reaching mouth edge. Dis- tinct facial carina reaching from base of antennae to a distinctly broadened and outstanding frontoclypeal tu- bercle. Ptilinal suture extending well beneath antennal bases on either side. Oral cavity tapers dorsally. Postcra- nium not obviously invaginated, yellow to light brown. Whole postcranium lightly dusted, with dusting adjacent to posterior eye margin obviously denser. Occiput and postgena covered with long black setulae. Postgena not widened and not delimited from occiput. Bottom por- tion of postcranium clearly delimited and less setulose. Proboscis reddish-brown, labellum black. Frontoclypeal membrane long, light orange-brown and barely delim- ited from orange-brown clypeus. Palps absent. Labium distinctly longer than head-length, distinctly thickened basally, anterior section completely fused and tubular. Labrum not visible in type specimen. Labellum short, completely divided, hardly broader than adjacent haust- ellum, and covered with very short setulae only. Thorax mainly orange-brown, with mediotergite, katepisternum and meron mainly black. Scutum with large medial black spot and small lateral black spots (Fig. 17). Thorax grey dusted all over. Pleura as in Fig. 18. Distinct dusting stripe on pleura reaching from middle coxa to notopleuron. The dusting stripe depends on the viewing angle and is best seen in dorsal view – care is required as it may be hard to see with the wrong view- ing angle (Fig. 18). Anterior half of anepimeron shining to subshining. Presternum distinct, narrow. Basisternum broad, ventrally narrowed to a point, and with scattered setulae. Proepisternum lacks setae or setulae. Notopleuron with a few slightly stronger setae. Postalar callus with a few strong short setae flanked laterally by several curved setulae. Katepisternum with 13 – 15 setae postero- dorsally and no setae ventrally. Metakatepisternum lacks setae. Anepimeron with 8 – 10 small setulae. Mediotergite convex, with minute and hardly visible black setae. Sub- scutellum inconspicuous. Scutum covered with scattered small black setulae and with no outstanding setae. Wing as Fig. 21: Fore-margin tinged brown, with brown mem- brane between subcosta and media M. Cell r 4 + 5 hyaline apically and discal-medial cell dm brown basally. Veins brown, subcosta paler brown than radial veins. Wing mostly covered with microtrichia, but anal lobe partly lacking microtrichia. Radial-medial crossvein rm small but complete (Fig. 20). Basal-medial-cubital crossvein incomplete. Radius R 1 and R 2 + 3 terminate close together in costa, well beyond end of subcosta. Radius R 4 + 5 with shallow, even curve in distal section which is directed to- wards fore-edge of wing. Cell r 4 + 5 pedunculate, with vein R 4 + 5 + M 1 distinctly longer than radial-medial crossvein. Basal cell lacks any thickening. Cubital cell cup elongat- ed, distinctly longer than vein A 1 + CuA 2, and pointed dis- tally (i. e. cubitus CuA 2 and anal vein A 1 meet at an acute angle). Vein A 1 + CuA 2 not reaching hind margin of wing. Cubital veins CuA 1 and basal-medial-cubital crossvein distinctly separated. Upper and lower calypters yellow- ish-white to brown, margin of upper calypter with short white setulae. Alula slightly shorter than broad, lacking setulae on posterior margin. Venae spuriae pronounced in cell r 4 + 5, cubital cell cup and indistinctly in cubital cell cua 1. Haltere light yellowish-white with brown base and reddish knob. Knob of haltere with brown setulae. Legs orange with fore coxa mainly black. Legs with in- conspicuous silver dusting to shining, hind coxa densely silver-dusted. Posterior surfaces of fore and middle tib- iae with obvious silver-dusted fields distally. Legs with short, adpressed black setulae. Base of fore and middle femora lacking denser black setulae. Areas of dense black to brown setulae anteroventrally at tip of fore tibia, and ventrally and posteriorly at tip of hind tibia. Middle femur lacking a distinct row of regularly arranged setu- lae. Hind femur lacks outstanding setulae. No preapical setae dorsally and no setae ventrally on tibiae. Femora ventrally lacking rows of short black setae. Coxae with several setulae but lacking outstanding setae. Hind femur slightly thickened in basal half. All tibiae thickened in apical half. Each metatarsus with 2 stronger setae ven- trally at base. Pulvilli yellowish-white. Claws brown, with narrow black tips. Empodium light brown, about as long as pulvilli. Abdomen orange to dark brown, with diffuse dark markings dorsally on tergites 1 – 4. Abdomen with short black setulae all over, those on tergite 2 very small. ♀ abdomen completely silver-dusted, although the impression of dusting depends on viewing angle. Denser dusted hind margins of tergites are not obvious. Tergites 1 – 3 fused but remain distinct from each other. Maximum width of abdomen at segment 4. Length: maximum width of terg- ite 2 = 2.0; length: maximum width of tergite 3 = 1.9. Tergite 5 and sternite 5 not completely fused laterally. Shape of theca as shown in Fig. 19. Anterior surface of theca apically with a few long setulae. Posterior surface of theca almost completely covered with close-set long blunt spicules, arranged in about 12 horizontal lines. Tergite 7 bends distinctly ventrally, with an indistinct longitudinal gap, and with a minute protruding tooth at middle of posterior margin. Female characters added from one dissected paratype Sternites 1 – 2 fused. Sternites 3 – 4 not protruding ven- trally, posterior parts of sternites inconspicuous. Tergites 3 and 4 lack evaginations. Sternite 5 anteriorly lacking any elongation. Sternite 6 almost completely covered with long blunt spicules which are mainly arranged in close-set, broken horizontal lines. Sternite 7 longer than broad, anteriorly pointed, posterior margin with two patches of dense long black setae. Ventral sclerotisation fused with tergite 8 and therefore connecting at sides. Tooth on tergite 8 distinct, its base elongated anteriorly. Sternite 8 bulging posteriorly, covered with strong long black setae and with obvious long setulae on posterior margin. Paired cerci distinct. Sack-like ventral protrusion of vagina with distinct annular sclerotisation. Opposite the annular sclerotisation the ventral protrusion is hardly sclerotised and is covered with short, broad microtrichia giving the ventral protrusion a roughened surface. No ob- vious sclerotisation at base of either the accessory glands or the spermathecal ducts. Sack-like ventral protrusion of vagina lacks any additional sclerotisation. 2 pairs of spherical spermathecae, the spermathecal ducts fusing shortly after leaving the spermathecae. Spermathecal ducts are sclerotised at the spermathecae, this sclerotised part of the duct being sinuous. Description of male ♂ abdomen strongly dusted from posterior half of tergite 3 to tip (in anterior view) and almost shining laterally on tergites 1 – 3. No obvious dusting at hind margin of terg- ites. Tergite 1 with obvious black setulae laterally on bul- bous lateral projections. Tergite 2 distinctly elongated, about 5 times as long as width at posterior margin. Terg- ite 2 lacking lateral tufts of setulae. Tergite 3 obviously longer than broad and widened posteriorly, anteriorly about 2.8 times as wide as posteriorly. Sternites 1, 4 and 5 present, sternites 2 and 3 reduced. Tergite 5 and sternite 5 separate. Sternite 4 almost square, minute and not clear- ly delimited from membrane, with about 4 minute setu- lae. Sternite 5 apically with small field of thick setae and several long black setulae laterally. Protandrium broader than epandrium and therefore projecting over it. Sternite 8 delimited from protandrium. Line of minute black setu- lae at border of protandrium and sternite 8. Lateral edges of protandrium fused ventrally by a narrow sclerotised strip, which is not medially broadened. Paired cerci dis- tinct, completely sclerotised and covered with scattered black setulae. Epandrium as shown in Fig. 22, not fused behind cerci but with a slightly sclerotised connection on both sides of the epandrium. Within this connection there is an elongated sclerotisation. Posterior margin of epandrium with small black setulae, and an obvious short black tooth with a broad blackish base. Epandrium distinctly concave laterally. No hypoproct evident and no remains of surstyli recognised. No obvious strong black setae nor long black setulae which would mark the base of a surstylus. Subepandrial plate not sclerotised nor cov- ered with microtrichia, and therefore not distinct. Dorsal hypandrial bridge developed. No hypandrial lobe evi- dent. Hypandrial bars fused distally, hypandrium ending in a hypandrial arm. Hypandrial membrane almost ab- sent, lacking microtrichia. Phallus sheath fused dorsally, lacking any evagination or setulae. Postgonite distinct. Postgonite evagination not sclerotised, and not projecting above distiphallus. Lateral side of postgonite evagination with indistinct microtrichia, the medially directed surface with broad and overlapping microtrichia (Fig. 23). No plate at postgonite evagination. Ring sclerite between phallapodeme and distiphallus indistinct. No epiphallus recognised. Distiphallus shorter than epandrium, entirely covered with microtrichia, some of which are black and arranged in lines. Distiphallus dorsally with large light- ly sclerotized plate (Fig. 23) but lacking evaginations. Phallapodeme longer than hypandrium arm. Ejaculatory apodeme elongated, lacking distinct attachment to sperm sac. Variability Wing length 6.3 – 7.6 mm. Basal aristomere sometimes black apically. Face, gena, frons and postcranium some- times more or less black in melanistic specimens. Facial keel sometimes yellow. Extent of black marking on tho- rax varies somewhat, and scutum sometimes completely orange. Proepisternum may lack setulae. Wing some- times more extensively brown, with discal-medial cell dm almost completely brown and cubital cell cua 1 brown at base. Alula and cubital cell cup may partly lack mi- crotrichia. All coxae sometimes obviously blackish. Hind femur may have an obscure blackish or brown ring.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	diagnosis	Diagnosis Physocephala guillarmodi belongs to the antiqua species-group as defined in Key 1. It is easily overlooked amongst the more common and variable P. antiqua, but can be differentiated from the latter by the brown bas- al cell which is completely covered with microtrichia (Fig. 20). With this character P. guillarmodi can only be confused within the antiqua species-group with dark-winged specimens of P. antiqua which may atypically have the basal cell almost (but never completely) cov- ered with microtrichia, or with P. pusilla. Physocephala guillarmodi can be separated from both of these species by its light yellow to yellowish-brown subosta which is obviously different from the darker radial veins (Fig. 21) and by the characteristic light brown thorax with three black spots (which may be fused) on the scutum (Fig. 17). At a first glance this species resembles P. schmideggeri Stuke, 2017 which occurs in the Arabian Peninsula (Stuke, 2017 b) but P. guillarmodi is distinguished by the complete dusting band which starts immediately above the middle coxa and reaches up to the notopleuron with- out becoming narrower. In the event that P. brevipennis is eventually demonstrated to be an aberrant and misshaped antiqua specimen (see above) this will also be distinguished from P. guillarmodi by its dark brown subcosta which does not differ in colour from the radial veins.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	etymology	Etymology This species is named in honour of Charles Fréderic Jacot Guillarmod (1912 – 1979), who collected several speci- mens of this new species and whose collecting activities made a huge contribution to knowledge of the Diptera of South Africa.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD1EFFDF265A487FFBB56518.taxon	distribution	Distribution Physocephala guillarmodi is widely distributed in the Afrotropical Region (Fig. 24).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD03FFD8267F4C14FED16603.taxon	description	(Figs 25 – 29) Conops (Physocephala) larvatus Speiser 1911	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD03FFD8267F4C14FED16603.taxon	materials_examined	Material. DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, 30. xii. 1952, Nord-Kivu, Kinshasa, Vitshumbi, s Lake Edward, leg. J. Verbeke, coll. ISNB; TANZANIA: 1 ♂, 1911, Marienhof [= Murutunguru], Ukerewe, leg. Con- rads, coll. SMTD; UGANDA: 1 ♂, 12. viii. 1958, Uganda, det. as P. similis by Camras 2000, leg. J. Bowden, coll. NHML [NHMUK 010922104]. Physocephala similis was distinguished from P. larvata by Camras (2001) within his similis species-group, but the character which Camras used to identify similis (“ Face yellow ”) contradicts the original description of Kröber (1915). Kröber compared P. similis with P. larvata and stated that P. similis “ Ist P. larvata zum verwechseln ähnlich, hat aber ganz andere Flügelzeich- nung ” [Is confused with the similar P. larvata, but has quite a different wing pattern] although this difference is not obvious when comparing the original descriptions of P. similis and P. larvata. Both descriptions fit to the wing shown in Fig. 27 and there remains no other differ- ence between these two species. The depository of the holotype is unknown (Stuke 2017 a) and therefore Physocephala similis Kröber, 1915 is placed as a junior syn- onym of Physocephala [Conops] larvatus Speiser, 1911 (syn. nov.).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	description	(Figs 30 – 35) urn: lsid: zoobank. org: act: 7476796 B- 0 C 67 - 44 D 8 - AA 85 - 1313 BC 97 B 38 C	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	materials_examined	Holotype ♂. (1) “ 98 / 99 / 79 ”; (2) “ visiting white fls / Brownanthus kuntzei / (Schinz) Ihlenf. & Bittrich / Aizoe- ceae: Mesembryanthema ”; (3) “ Namibia / NW of Cape Cross / 21.44 S 13.59 E / 14. iii. 1999 / F. W. and S. K. Gees; (4) Holotypus / Physocephala ssymanki / spec. nov. / 2018 ”. Holotype is deposited in AMGS. The specimen is pinned and in perfect condition.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	description	Description of holotype (male) Length 11.7 mm; Wing-length 7.0 mm; Head-height 2.6 mm. Head. Antenna black, tip of pedicel and base of first flagellomere brown. Arista stylus-like, with 2 aristomeres situated at tip of first flagellomere (Fig. 32). Both ari- stomeres minute, length of apical aristomere shorter than height. Scape about twice as long as maximum width, apically with black setae. Pedicel about six times longer than maximum width, mainly dorsally and apically cov- ered with black setae. Pedicel lacking any ridge at base, and expanded towards apex. First flagellomere long and conical, about 2.5 times as long as high, pointed, ventral- ly lacking a membranous area. Lunule between base of antennae and ptilinal suture distinct, shorter than width of scape. Eye brown, lacking ommatrichia, with facets all of about the same size. Posterior margin of eye lack- ing any shining indentation. Gena height / eye height (measurements taken from head in lateral view) = 0.2. No ocellar tubercle, no ocelli and no ocellar triangle evi- dent. Frons (Fig. 31) broader than long, slightly concave, slightly projecting above eyes posteriorly, and lacking setulae. Anterior margin concave. Frons yellow, with no frontofacial spot and with indistinct dusting all over. Ver- tex as broad as frons, separated from latter by a more or less distinct ridge. Vertex shining, and in posterior half covered with black setulae; apically with an indistinct depression. Ridge of vertex with minute longitudinal grooves. Face yellow, lacking dusting except for some silver dusting along eye margin. Gena yellow, lacking setae. Distinct facial grooves reaching mouth edge, and facial carina reaching from base of antennae to a distinct- ly broadened and outstanding frontoclypeal tubercle. Ptilinal suture extending well beneath antennal bases on either side. Oral cavity tapers dorsally. Postcranium not obviously invaginated, black to dark brown. Whole post- cranium slightly dusted, with dusting adjacent to posteri- or margin of eye obviously denser. Occiput and postgena covered with black setulae. Postgena not widened and not delimited from occiput. Bottom portion of postcra- nium not delimited and not setulose. Proboscis black to reddish-brown, labellum black. Frontoclypeal membrane long, light orange-brown and hardly delimited from or- ange-brown clypeus. Palps absent. Labium longer than head-length in lateral view, distinctly thickened basally, and with anterior section completely fused into a tube. Labrum not visible in the holotype. Labellum short, com- pletely divided, hardly broader than adjacent haustellum, and covered with very short setulae only. Thorax mainly black, with pleura black to brown. Post- pronotum orange-brown. Thorax grey-dusted all over, with distinct dust-stripe reaching from middle coxa to notopleuron and becoming wider dorsally. Anterior half of anepisternum and posterior half of anepimeron shining (Fig. 34). Presternum distinct, broad. Basisternum broad, ventrally narrowed to a point, lacking setae or setulae. Proepisternum lacking setae or setulae. Scutum covered with small black setae. Notopleuron with several stronger setae. Postalar callus with a few strong, short black setae above several curved setulae. Katepisternum with 2 setae posterodorsally and no setae ventrally. Metakatepis- ternum lacks setae. Anepimeron with 12 long setulae about as long as maximum width of tibiae. Mediotergite convex, with minute and barely visible black setae. Sub- scutellum inconspicuous. Scutum covered with scattered small black setae, and with no outstanding setae. Wing as Fig. 35: Fore margin of wing tinged brown, with brown membrane between subcosta and radius R 4 + 5. Cell r 2 + 3 hy- aline apically and cell r 4 + 5 brown basally. Veins brown, subcosta paler brown than radial veins. Wing mostly covered with microtrichia, but with basal cell, basal me- 230 Jens-Hermann Stuke & David K. Clements dial cell, base of discal-medial cell and base of cubital cell cup lacking microtrichia. Radial-medial crossvein small but complete. Basal-medial-cubital crossvein also complete. Radius R 1 and R 2 + 3 terminate close together in costa, well beyond insertion point of subcosta in costa. Radius R 4 + 5 with shallow and even curve in distal section directed towards fore-edge of wing. Cell r 4 + 5 pedunculate, with vein R 4 + 5 + M 1 well expressed and distinctly longer than radial-medial crossvein. Basal cell lacks any thick- ening. Cubital cell cup elongated, distinctly longer than New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 231 vein A 1 + CuA 2, and pointed distally (i. e. cubitus CuA 2 and anal vein A 1 meet at an acute angle). Vein A 1 + CuA 2 not reaching hind margin of wing. Cubital veins CuA 1 and crossvein distinctly separated. Upper and lower calypters yellowish-white to brown, margin of upper calypter with white setulae. Alula almost square, about as long as broad, lacking setulae on posterior margin. Venae spuriae pronounced in cell r 4 + 5 and cubital cell cup. Haltere white, with light brown base. Knob of haltere with brown set- ulae. Legs orange, with hind femur and hind tibia black in apical half. Legs with inconspicuous silver dusting or shining, hind coxa densely silver-dusted. Posterior sur- faces of fore and middle tibiae with obvious silver-dusted fields distally. Legs generally with short, adpressed black setulae. Base of fore and middle femora basally with denser black setulae. Areas of dense black to brown set- ulae anteroventrally on tip of fore tibia and ventrally and posteriorly at tip of hind tibia. Middle femur lacking any distinct row of regularly arranged setulae. Hind femur lacking outstanding setulae. No preapical setae dorsally, and no setae ventrally on tibiae. Femora ventrally lacking rows of short black setae. Coxae with several setae, but lacking any outstandingly long ones. Hind femur slight- ly thickened in basal half. All tibiae thickened in apical half. Each metatarsus with 0 – 1 stronger seta ventrally at base. Pulvilli yellowish-white. Claws brown, with broad black tips. Empodium light brown, and about as long as pulvilli. Abdomen orange to dark brown, with diffuse paler markings. Abdomen with short black setulae all over. ♂ abdomen strongly dusted from posterior half of tergite 3 to tip (in anterior view) and almost shining laterally on tergites 1 – 3. No obvious dusting at hind margin of tergites. Tergites 1 – 3 fused but remain distinct. Abdomen of holotype not dissected. Tergite 1 with obvious black setulae on bulbous lateral projections. Tergite 2 elongat- ed, about four times as long as smallest width, lacking obvious lateral tufts of setulae. Tergite 3 slightly widened posteriorly, about twice as wide posteriorly than ante- riorly. Tergite 5 and sternite 5 separate. Sternite 5 with scattered long black setulae and denser small black setulae forming an indistinct field. Protandrium broader than epandrium and projecting over it. Sternite 8 distinctly de- limited from protandrium. Indistinct line of small black setulae at border between sternite 8 and protandrium. Paired cerci distinct, completely sclerotised and covered with scattered setulae. Epandrium not fused behind cerci, but with a slightly sclerotised connection at both sides of epandrium.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	diagnosis	Diagnosis Physocephala ssymanki belongs to the P. antiqua species-group and can easily be recognised from the anepi- meron, which has several characteristically long setulae which are about as long as the maximum width of the tibiae. The dusting stripe on the pleura obviously con- trasts with the shining anepisternum and anepimeron, and becomes wider dorsally (Fig. 34). The hind tibia and femur are black in the apical two-thirds, contrasting with the remaining orange-brown legs (Fig. 30).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	etymology	Etymology This species is dedicated to Axel Ssymank (Bonn) who generously loaned numerous Conopidae specimens for this project.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD04FFDB25D84F13FB6A630D.taxon	distribution	Distribution To date P. ssymanki is only known from the locus typicus on the coast of southern Namibia.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD07FFDB267F4A0EFA996187.taxon	description	(Fig. 36) Conops pusilla Meigen 1804	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD07FFDB267F4A0EFA996187.taxon	materials_examined	Material. CENTRAL AFRICAN REPUBLIC: 1 ♂ 2 ♀♀, 4. xii. 2010, 60 km w Banibo [3 ° 50 ′ N 16 ° 44 ′ E], 630 m, leg. J. Halada, coll. CULSP. Afrotropical specimens of this otherwise Palaearctic spe- cies have previously caused confusion, and for a long time were suspected of belonging to a different species. Somewhat surprisingly, however, their morphological characters fall within the variability of P. pusilla as recently defined by Stuke (2016). Physocephala pusilla had not previously been recorded from the Afrotropi- cal Region and was assumed to be a Palaearctic species (Stuke 2017 a). Together with P. antiqua, P. vittata and Conops elegans Meigen, 1804, this is the fourth example of a very variable and mainly Palaearctic species with a distribution which extends into the Afrotropical Region.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD07FFD4267F4E6EFDDF64FB.taxon	description	(Figs 37 – 41) Physocephala bimarginipennis Karsch 1887 = Physocephala gigantea Kröber, 1936	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD07FFD4267F4E6EFDDF64FB.taxon	materials_examined	Primary type material examined. ♀ holotype of Physocephala bimarginipennis Karsch, 1887: (1) “ Pungo An- dongo / A. v. Homayer ”; (2) “ 11031 ”; (3) “ Physocephala / bimarginipennis / n. sp. ”; (4) “ Holotypus ”; coll. ZMHB. ♂ syntype of Physocephala gigantea Kröber, 1936: (1) “ Ph. Type / gigantea ”; (2) “ Musée du Congo / Elisabeth- ville / XI- 1927 / Dr. M. Bequaert “; (3) “ Physocephala / gigantea / Krb. / det. Kröber 1935 ”; (4) “ R. DÈT / V / 2992 ”; (5) “ Typus ”; (6) “ RMCA ENT / 000012173 ”; coll. MRAC. ♂ syntype of Physocephala gigantea Kröber, 1936: (1) “ Paratype / Ph. gigantea ”; (2) “ Musée du Congo / Kivu: Musingiro / 8 - IX- 1927 / Ch. Seydel ”; (3) “ Musinglro [si] / 8 Sept. 1927 / Ch. Seydel ”; (4) “ Physocephala / gigantea Krb. / det. Kröber 1935 ”; (5) “ R. DÈT / W / 2992 ”; coll. MRAC. Additional material. DEMOCRATIC REPUBLIC OF CONGO: 1 ♀, 8. vii. 1937, Bambesa, det. as P. gigantea by Janssens 1954, leg. J. Vrydagh, coll. MRAC; 1 ♀, vi. 1932, Eala, det. as P. bimarginipennis by Camras 1962, leg. A. Corbisier, coll. MRAC; 1 ♀, 5. iii. 1954, North Kivu Province, Lac Vert, forêt route Goma – Sake, leg. J. Verbeke, coll. ISNB; ETHIOPIA: 1 ♀, 4. xii. 2014, Bonga Waldsaum, 1628 m [07 ° 11.87 ′ N 036 ° 15.13 ′ E], Bothriocline schimperi, leg. H. - J. Flügel, coll. PHJF; 1 ♂, viii. 2019, Bonga, Kafa Development Association Guest- house [7 ° 14 ’ 59.14 ’’ N 36 ° 15 ’ 12.26 ’’ E], leg. T. Kirschey, coll. PJHS; KENYA: 1 ♀, 16. xi. 1967, Nairobi, ex Xylocopa flavorufa found dead 23. ix. 1967 Green Office Colo- ny, [collector unknown], coll. NHML; 1 ♀, 9. – 23. v. 2006, Coast Province, Shimba Hills National Park, near artifi- cial Ponds [4.22752 ° S 39.43197 ° E], 335 m, mixed grassshrub-land, Malaise trap, leg. R. Copeland, coll. NMKE; SOUTH AFRICA: 1 ♀, 1. – 3. iv. 2001, Mpumalanga Prov- ince, Blyderevierspoort NP [24 ° 39 ′ S 30 ° 50 ′ E], leg. D. J. Greathead, coll. ZMHB; TOGO: 1 ♂, ii. 2017, Kloto, for- est area [6 ° 57 ’ 31.66 ’’ N 0 ° 34 ’ 29.75 ’’ E], leg. G. Goergen, coll. IITA; 1 ♀, xii. 2016, ditto; 1 ♀, x. 2001, ditto; UGAN- DA: 1 ♂, 22. vii. 1966, Kwanda, ex ♂ Xylocapa rufa, leg. D. J. Greathead, coll. NHML; 1 ♀, 12. iii. 2012, Western Uganda, n Fort Portal Kijura, forest edge [N 302540 E 004731], leg. Oehlke, coll. PASS.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD08FFD425D84DBAFB1867F3.taxon	description	Females of the caenoneura species-group are easily recognised by the normally developed, usually ad- pressed theca and short tergites 5 and 6. From outside the caenoneura group, only P. microvena and P. pseudomicrovena share this combination of characters but are easily recognised as belonging to the Physocephala microvena species-group by the lack of a vena spuria in cell r 4 + 5, and typically also by the reduced or absent radi- al-medial crossvein. Males of caenoneura group can be more difficult to assign, however, the elongated basal ari- stomere and less modified wing venation being the only characters which reliably distinguish them from males of the microvena species-group. All members of this group are restricted to the Afrotropical Region.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD625D84F50FDD16703.taxon	description	(Figs 42 – 45)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD625D84F50FDD16703.taxon	materials_examined	Material. SOUTH AFRICA: 1 ♀, 20. – 24. xi. 2014, Kwa- Zulu-Natal Province, Royal Natal National Park, Gudu Bush falls [28 ° 40.925 ′ S 28 ° 55.778 ′ E], 1628 m, Malaise trap in indigenous afromontane forest, leg. A. H. Kirk-Spriggs & E. Letsobe, coll. BMSA.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD625D84E11FE6665E5.taxon	description	(Figs 46 – 49) Physocephala braunsi Kröber 1931	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD625D84E11FE6665E5.taxon	materials_examined	Material. SOUTH AFRICA: 1 ♂, 15. iv. 1948, Marieps Mountain, det. as P. braunsi by Camras 2000, leg. G. van Son., coll. FMNH; 1 ♂, xi. 1933, Cape Province, Swel- lendam, det. as P. platycephala by Kröber 1938, det. as P. braunsi by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922151]; ZIMBABWE: 1 ♀, 7. xii. 1976, Salisbury [= Harare] [17.84 31.05], leg. P. E. Hulley, coll. AMGS. We are not convinced that the characters used to dis- tinguish P. braunsi from the very similar P. caenoneura merit separate species status, but the few specimens which are available can be segregated using the characters given in Key 4.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD6265A4B92FAE566C0.taxon	description	(Figs 50 – 54)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD6265A4B92FAE566C0.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala caenoneura Kröber 1939: (1) “ Type ”; (2) “ Nyassa-See / Langenburg / 26. VII. – 8. VIII. 98 / Fül- leborn S. ”; (3) “ Typus ”; (4) “ Pseudophysocephala / stylata Kb. / det. Kröber 1938 ”; (5) “ Zool. Mus. / Berlin ”; coll. ZMHB. 1 ♂ holotype of Pseudophysocephala stylata Kröber 1939: (1) “ Type ”; (2) “ Nyassa-See / Langenburg / 1. – 26. VII. 98 / Fülleborn S. ”; (3) “ Typus ”; (4) “ Pseudophysocephala / stylata Krb. ”; (5) “ Zool. Mus. / Berlin ”; (6) “ Pseudophysocephala / caenoneura Kröber / det. Camras 1999 ”; coll. ZMHB. Additional material. KENYA: 1 ♀, 2. iii. 1993, Il- rad, w Nairobi, 1900 m, leg. B. Merz, coll. TAUI; 1 ♀, 10. viii. 2004, near Karura Forest, Nairobi [1 ° 14.829 ’ S 36 ° 48.953 ’ E], leg. R. Copeland, coll. NMKE; TAN- ZANIA: 1 ♂ 1 ♀, 4. ii. 1977, Amani, East Usambara, 1000 m, det. as P. caenoneura by Camras 1999, leg. H. Enghoff, O. Lomholdt, O. Martin, coll. FMNH; 1 ♂, 10. iii. 1963, Kilimajaru, Weruweru, det. as P. caenoneura by Camras 1999, leg. D. J. Greathead, coll. NHML [NHMUK 010922144]; 1 ♂, 20. iv. 1966, plains e Arusha, det. as P. caenoneura by Camras 1999, leg. D. J. Great- head, coll. NHML [NHMUK 010922145] 6.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD6265A4FCDFC2164DF.taxon	description	(Figs 55 – 59) Physocephala capensis Kröber 1931	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD6265A4FCDFC2164DF.taxon	materials_examined	Material. SOUTH AFRICA: 1 ♂, 25. ii. 2016, Barberton, Mpumalanga [25 ° 10 ′ 18 ″ S 30 ° 56 ′ 45 ″ E], 1400 m, leg. A. Ssymank, coll. PASS; 1 ♀, 4. – 7. xi. 1931, Cape Prov- ince, George, det. as P. capensis by Kröber, leg. H. P. Thommaset, coll. NHML [NHMUK 010922135]; 1 ♀, 23. ix. 2007, Eastern Cape Province, Zuurberg [33 ° 21 ′ S 25 ° 44 ′ E], leg. C. Turnbull, coll. AMGS; 1 ♀, 15. xii. 2012, KwaZulu-Natal Province, Kwela Lodge, Pietermaritz- burg [29 ° 29 ′ 34 ″ S 30 ° 21 ′ 40 ″ E], 930 m, leg. G. Ståhls, coll. PASS; 1 ♂, 15. x. 2003, Western Cape Province, Hu- mansdorp [34 ° 1 ′ 60 S 24 ° 46 ′ 0 E], 114 m, leg. C. Hepburn, coll. AMGS.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD1265A4DDFFE64668F.taxon	description	(Figs 60 – 62)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0AFFD1265A4DDFFE64668F.taxon	materials_examined	Primary type material examined. 1 ♂ syntype of Pseudophysocephala fenestralis Kröber 1939: (1) “ Type ”; (2) “ Nyassa-See / Langenburg / 14. V. 1999 / Fülleborn S. ”; (3) “ Typus ”; (4) “ Pseudophysocepha- 236 Jens-Hermann Stuke & David K. Clements la / fenestralis Krb. / det. Kröber 1938 ”; coll. ZMHB. 1 ♀ syntype of Pseudophysocephala fenestralis Kröber 1939: (1) “ Syn- / type ”; (2) “ George, Cape Province, / 27. VI. – 1. VII. 1920. ”; (3) “ S. Africa / R. E. Turner. / 1920 - New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 237 318. ”; (4) “ Pseudophysocephala / fenestralis, Krb. / ex- amined & det. / O. Krober, 1938. ”; (5) “ Pseudophysocephala / fenestralis / Krb ”; coll. NHML. Additional material. TANZANIA: 1 ♀, 20. viii. 1980, Mt. Rungwe, sw, 1900 m, det. as P. fenestralis by Cam- ras 2000, leg. M. Stoltze, N. Scharff, coll. FMNH; 1 ♀, 16. ii. 1952, Njombe, 6000 – 6500 ft, det. as P. fenes- 238 Jens-Hermann Stuke & David K. Clements tralis by Camras 1999, leg. W. Peters, coll. NHML [NHMUK 010922142].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0DFFD125F24F8FFB1564FD.taxon	description	(Figs 63 – 65) Pseudophysocephala rufitarsis Camras 1962	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0DFFD125F24F8FFB1564FD.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala rufitarsis Camras 1962: (1) “ Holo- / type ”; (2) “ S. Africa. / R. E. Turner / Brit. Mus. / 1933 - 108. ”; (3) “ Holotype ♀ / Pseudophysocephala / rufitar- sis / Camras ”; (4) “ Pseudophysocephala / platycephala Loew / examined & det. / O. Kröber. 1938 ”; (5) “ E. Cape Prov. / Katberg. / 15 – 30. i. 1933. ”; (6) “ BMNH (E) # / 249149 ”; coll. NHML Additional material. SOUTH AFRICA: 1 ♂, i. 1953, Na- tal, Hilton Road, det. as P. rufitarsis by Camras 2000, leg. P. Graham, coll. NHML [NHMUK 010922159]; 1 ♂, xi. 1930, Cape Province, Somerset East, det. as P. platycephala by Kröber 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922162]; 2 ♂♂, 19. x. 1994, Cape Province, Tsitsikama National Park, Storms River Pass [33 ° 59 ′ S 23 ° 55 ′ E], leg. R. Daniels- son, coll. MZLU; 1 ♀, 1. – 10.1932, Eastern Cape Prov- ince, Katberg, det. as P. platycephala by Kröber 1938; det. as P. rufitarsis by Camras, 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922189]; 1 ♀, 1. – 10.1932, Eastern Cape Province, Katberg, det. as P. platycephala by Kröber 1938; det. as P. rufitarsis by Camras, 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922190]; 1 ♂, 11. – 18. ii. 1933, Eastern Cape Province, Katberg, det. as P. platycephala by Kröber, 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922158]; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as P. platycephala by Kröber 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922160]; 1 ♂, 1. – 12. iii. 1933, Eastern Cape Province, Katberg, det. as P. platycephala by Kröber 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922161]; 1 ♀, 15. – 30. i. 1933, Eastern Cape Province, Ratberg, det. as P. platycephala by Kröber 1938, det. as P. rufitarsis by Camras 2000, leg. R. E. Turner, coll. FMNH.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0DFFD2267F4DB9FED6649C.taxon	description	New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 239 halteres. Members of the Physocephala halterata species-group can be identified using Key 5 below. All mem- bers of this species-group are restricted to the Afrotrop- ical Region.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0FFFD325F24E6EFAD76729.taxon	materials_examined	Primary type material examined. ♂ holotype of Conops acroschista Speiser, 1911: (1) “ Holo- / TYPE ”; (2) “ Musée du Congo / Kasaï: Kondué / E. Luja ”; (3) “ R. DÉT / B / 1477 ”; (4) “ Type! / Conops (Physocephala) / acroschistus m. / P. Speiser det. ” /; (5) “ RMCA ENT / 000012172 ”; coll. MRAC. The holotype reported here fits in all aspects to the origi- nal description and there is no reason to assume that it is not the type specimen, although according to the original description this should be in the Museum d’Histoire Nat- uralle du Grand-Duchy de Luxembourg (MGDL). Cam- ras (2001) reported concerning a specimen in the MGDL as follows: “ The unidentified holotype of acroschista was finally found by J. M. Guinet and he informed me that Luja was the collector. ” The specimen reported by Camras (2001) was a female, however, whereas in the original description it was explicitly stated that the type was a male, and a male is described. Evidently there are more than one specimen bearing the same labels, which led to the misinterpretation of the female specimen in the MGDL. Camras placed the species in his Pseudophysocephala vitripennis species-group, having presumably misinterpreted the species due to this wrongly labelled specimen. The synonymy of P. intermedia Kröber, 1936 and P. acroschista which was introduced by Camras (2001) is therefore invalid and P. acroschista should be treated as valid species (status rev.: valid species).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0FFFEC267F4EE5FB9764DE.taxon	description	(Fig. 66) Physocephala halterata Brun. var. congoënsis Kröber 1936	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD0FFFEC267F4EE5FB9764DE.taxon	materials_examined	Primary type material examined. ♂ syntype of Physocephala halterata Brun. var. congoënsis Kröber, 1936: (1) “ Ph. Type ♂ / halterata var / congoensis ”; (2) “ Musée du Congo / Bambesa / 15 - IX- 1933 / H. J. Bredo ”; (3) “ Physocephala ♂ / halterata Brun. / var. congoensis / Krb / det. Kröber 1935 ”; (4) “ R. DÉT ” / P / 2992 ”; (5) “ Typus ”; (6) “ RMCA ENT / 000012174 ”; coll. MRAC. ♀ syntype of Physocephala halterata Brun. var. congoënsis Kröber, 1936: (1) “ Ph. Type ♀ / halterata var / congoensis ”; (2) “ Musée du Congo / Bambesa / 25 - IX- 1933 / H. J. Bredo ”; (3) “ Physocephala ♀ / halterata Brun. / v. congoensis Krb / det. Kröber 1935 ”; (4) “ R. DÉT ” / Q / 2992 ”; (5) “ Typus ”; coll. MRAC. New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 241 Additional material. BURUNDI: 1 ♂ 1 ♀, 15. i. 1950, Bururi, 1900 m, leg. F. J. François, coll. ISNB; 1 ♀, 26. v. 1949, Bururi Province, Bururi, 1950 m, leg. F. J. François, coll. ISNB; 1 ♂, 15. v. 1949, Bururi Province, Bururi, 2050 m, leg. F. J. François, coll. ISNB; DEM- OCRATIC REPUBLIC OF CONGO: 1 ♀, 21. – 27. viii. 1931, Tshibinda [- 0.287 28.776], det. as P. congoensis by Camras 2000, leg. J. Ogilvie, coll. NHML [NHMUK 010922105]; 1 ♂, 5. ii. 1949, North Kivu Province, Gishari, Territory Masisi, 2300 m, leg. F. J. François, coll. ISNB; EQUATORIAL GUINEA: 1 ♀, 15. – 31. i. 1907, Uelleburg, Benitogebiet, det. as P. congoensis by Camras 2000, leg. G. Tessmann, coll. ZMHB. Physocephala congoensis is very similar to P. halterata and P. acroschista, and all three species could be inter- preted as colour morphs of a single species. As Camras (2001) pointed out before, however, the few specimens which are available can be consistently segregated using the characters given in Key 5.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD30FFEF265A4DDFFD7960F1.taxon	description	(Figs 67 – 70) Physocephala halterata Brunetti 1925	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD30FFEF265A4DDFFD7960F1.taxon	materials_examined	Primary type material examined. ♀ holotype of Physocephala halterata Brunetti 1925: (1) “ Holo- / type ”; (2) “ S. Africa. ”; (3) “ Umbilo / 17. X. 15 ”; (4) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306. ”; (5) “ Durb / Mus ”; (6) 242 Jens-Hermann Stuke & David K. Clements “ Physocephala / halterata / Brun Type ♀ ”; (7) BMN- H (E) # / 249078; coll. NHML	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD30FFEF265A4DDFFD7960F1.taxon	description	Additional Material. BURUNDI: 1 ♀, 23. iv. 1955, Gitega [= Kitega], 1700 m, leg. F. J. François, coll. ISNB; DEMOCRATIC REPUBLIC OF CONGO: 1 ♀, 6. iii. 1948, Katanga Province, Mukana, Lusinga, 1810 m, leg. G. F. de Witte, coll. ISNB; 1 ♀, 15. iii. 1948, ditto; ETHIOPIA: 1 ♀, 4. xii. 2014, Bonga Waldsaum, 1628 m [07 ° 11.87 ′ N 036 ° 15.13 ′ E], Bothriocline schimperi, leg. H. - J. Flügel, coll. PHJF; KENYA: 1 ♀, 25. ix. 2005, Ker- icho junction [0 ° 23 ′ S 35 ° 16 ′ E], 1530 m, leg. L. Fried- man, coll. TAUI; 1 ♂, 23. v. 1910, Muguga, det. as P. halterata by Camras 2000, [collector unknown], coll. NHML [NHMUK 010922109]; 1 ♀, vii. 1937, Nairobi, det. as P. halterata by Camras 2000, leg. V. G. L. van Someren, coll. NHML [NHMUK 010922121]; 1 ♂, vii. 1937, Nairo- bi, leg. van Someren, coll. NHML [NHMUK 010922111]; LESOTHO: 1 ♂, 30. xii. 1950, Mamathes [- 29.136617 27.845796], leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♀, 4. xi. 1951, ditto; 2 ♂♂, 9. xi. 1951, ditto; 1 ♂, 11. xi. 1951, ditto; 2 ♀♀, 3. ii. 1952, ditto; 1 ♂, 15. ii. 1952, ditto; 1 ♀, 11. xi. 1952, ditto; 2 ♂♂, 23. xi. 1952, ditto; 1 ♂, 14. xi. 1954, ditto; SOUTH AFRICA: 1 ♂, 1906, Transvaal, leg. A. J. Cholmley, coll. NHML [NHMUK 010922113]; 1 ♀, 6. – 7. i. 1993, Cape Province, Cape of Good Hope Nat. Res., leg. F. Koch, coll. ZMHB; 1 ♂, 12. xi. 1952, Cape Province, Grahamstown, leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, 23. i. 1970, Cape Province, Grahamstown, Belmont Valley, on flowering Foeniculum vulgare Mill., leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, i. 1922, Cape Prov- ince, Mossel Bay, det. as P. halterata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922114]; 1 ♀, iv. 1933, Cape Province, Mossel Bay, det. as hal- terata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922117]; 1 ♀, 1. – 14. xi. 1921, Cape Prov- ince, Mossel Bay, det. as P. halterata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922120]; 1 ♀, 22. iii. 1939, Cape Province, Mossel Bay, det. as P. halterata by Camras 1962, leg. R. E. Turner, coll. NHML [NHMUK 010922124]; 1 ♂, 29. i. – 5. ii. 1924, Eastern Cape Province, Port St. John, Bondoland, det. as P. halterata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922115]; 1 ♀, 6. – 25. ii. 1924, East- ern Cape Province, Port St. John, Bondoland, det. as P. halterata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922119]; 1 ♀, ii. 1927, Free State Province, Harrismith, det. as P. halterata by Kröber 1938, leg. R. E. Turner, coll. NHML [NHMUK 010922118]; 1 ♀, 10. x. 2004, Gauteng Province, Benoni [26 ° 08 ′ S 28 ° 23 ′ E], leg. C. Midgley, coll. AMGS; 1 ♂, ii. 1897, Na- tal Province, Karkloof, [collector unknown], coll. NHML [NHMUK 010922110]; 1 ♀, 25. – 31. iii. 2001, Northern Province, Lekgalameetse Reserve [24 ° 12 ′ S 30 ° 20 ′ E], leg. F. Koch, coll. ZMHB; TANZANIA: 1 ♂, end of xii. 1898 to end of i 1899, Nyasasee, Langenburg [? = Tukuyu], det. as P. congoensis by Camras 2000, leg. S. Fülleborn, coll. ZMHB; 1 ♀, 24. xi. 1963, Iringa Region, Ulete, det. as halterata by Camras 2000, leg. D. J. Great- head, coll. NHML [NHMUK 010922122]; UGANDA: 1 ♂, 13. ii. 1966, Lake Nabugabo, leg. D. J. Greathead, coll. NHML [NHMUK 010922112]; ZIMBABWE: 1 ♂, 10.1926, Chirinda Forest, det. as P. halterata by Bry- ant, leg. G. Arnold, coll. NHML [NHMUK 010922116]; 1 ♀, 16. – 18. iii. 2001, Vumba Mountain, Botanical Gar- den [19 ° 07 ′ S 32 ° 47 ′ E], leg. F. Koch, coll. ZMHB; UN- KNOWN LOCATION: 1 ♀, 25. v. 1901, [characters il- legible], det. as P. halterata by Kröber 1938, [collector unknown], coll. NHML [NHMUK 010922123].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEA25D84964FC5260F1.taxon	description	(Figs 71 – 74)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEA25D84964FC5260F1.taxon	materials_examined	Primary type material examined. ♀ holotype of Physocephala longitheca Camras, 2001: (1) “ Angola (A 37) / 5 mls. NE. Negola / 25. iii. 1972 ”; (2) “ Southern / African Exp. / B. M. 1972 – 1 ”; (3) “ Holotype ♀ / Physocephala / longitheca / Camras ”; coll. NHML. ♂ holotype of Physocephala atronata Camras, 2001: (1) “ W. Uganda / Kibale Forest / 12. xii. 1971 – 9. i. 1972 / R. L. Mason ”; (2) “ Holotype ♂ / Pseudophysocephala / atronata / Camras ”; coll. NHML. Additional material. DEMOCRATIC REPUBLIC OF CONGO: 2 ♀♀, 1. – 20. i. 1919, Albertville [= Kalemi], 780 m, det. as P. “? abyssinica ” by Brunetti 1925, leg. R. Mayné, coll. MRAC; 1 ♂, x. 1932, Lulua, Kapan- ga, det. as P. “? abyssinica Krb ” by Kröber 1935, leg. F. G. Overlaet, coll. MRAC; MALAWI: 1 ♂, 4. x. 1919, Nyasaland, Cholo. [= Thyolo District], det. as P. “? abyssinica ” by Brunetti 1924, leg. R. C. Wood, coll. NHML [NHMUK 010922100]; UGANDA: 1 ♀, 21. i. 1913, West Mengo District, Entebbe, det. as P. “? abyssinica ” by Brunetti 1924, leg. C. C. Gowdey, coll. NHML [NHMUK 010922101]; ZIMBABWE: 1 ♂, 2. i. 1966, n Vumba [= Bvumba] [- 19.093432 32.740631], paratype of P. fumivena, leg. D. Cookson, coll. FMNH.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEA265A49BDFB0267C8.taxon	description	(Fig. 75) Pseudophysocephala brevipetiola Camras, 1962	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEA265A49BDFB0267C8.taxon	materials_examined	Primary type material examined. ♂ holotype of Pseudophysocephala brevipetiola Camras, 1962: (1) “ ♂ / Ho- lotypus ”; (2) “ Musé du Congo / Eala / III- 1932 / H. J. Bredo ”; (3) “ Holotype ♂ / Pseudophysocephala / brevipetiola / Camras ”; (4) “ RMCA ENT / 000012177 ”; coll. MRAC. Additional Material. GABON: 1 ♂, viii. 1892, Libre- ville, determined as a Physocephala nov. spec. by Kröber 1921, [collector unknown], coll. MLUH.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEA265A49BDFB0267C8.taxon	description	Physocephala brevipetiola is a rarely recorded species. Only the two males mentioned above are known to date. These are obviously well characterised by the unique ab- domen shape, having a very narrow base of the abdomen (“ petiole ” sensu Camras (1962), cf Fig. 75) which is ob- viously much shorter than remaining bulbous abdomen (“ abdominal club ” sensu Camras 1962).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEB265A4EA7FB2665C6.taxon	description	(Figs 76 – 83)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD36FFEB265A4EA7FB2665C6.taxon	materials_examined	Primary type material examined. ♀ lectotype of Physocephala constricta Kröber 1915 designated by Camras (2001): (1) “ 3053 ”; (2) “ Type ”; (3) “ Physocephala ♀ / constricta Krb. / O. Kröber det. 1914 ”; coll. ZMHB. ♂ holotype of Physocephala ruficoxa Kröber 1933: (1) “ Holo - / type ”; (2) “ Lady Grey / 3 Feb. 1925. / R. I. Nel ”; (3) “ Pres. By / Imp. Inst. Ent. / Brit. Mus. / 1932 - 143 ”; (4) “ Physocephala / ruficoxa Kröb / det. Kröber 1930 ”; (5) “ Type ”; (6) “ Pseudophysocephala / constricta Kröber / compared to type / det. Camras, 2000 ”; coll. NHML. 248 Jens-Hermann Stuke & David K. Clements Additional material. LESOTHO: 1 ♂, 13. xi. 1949, Mamathes [- 29.136617 27.845796], leg. C. Jacot-Guil- larmod, coll. AMGS; 1 ♂, 7. xi. 1951, Mamathes [- 29.136617 27.845796], leg. A. Jacot-Guillarmod, coll. AMGS; 2 ♂♂, 3. ii. 1952, Mamathes [- 29.136617 27.845796], leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, 11. ii. 1952, ditto; 1 ♂, 23. xi. 1952, ditto; 1 ♂, 15. iii. 1951, Quthing, det. as P. spec., interrupta group by Smith 1966, leg. Brink, Rudebeck, coll. MZLU; 1 ♂, 12. iii. 1951, Se- balabala, leg. C. Jacot-Guillarmod, coll. AMGS; 1 ♂, 8. – 14. i. 1963, Maseru district, Bushmans Pass, Maloti Mountains, 2125 – 2250 m, det. as P. lineifrons by Cam- ras 1963, leg. B. R. Stuckenberg, P. J. Stuckenberg, coll. FMNH; SOUTH AFRICA: 1 ♂, “ 3053 ” [? Cape], paralectotype of P. constricta, leg., coll. ZMHB; 1 ♀, iii. 1959, Cathedral Peak, Forestry reserve, Natal Drak- ensberg Little Berg summits, themeda Grassland, 5500 – 6000 ft, det. as P. constricta by Camras 1962, leg. B. R. Stuckenberg, P. J. Stuckenberg, coll. PJHS; 1 ♂, 10. – 12. xi. 2009, Cape Province, Harrismith Scotland farm [27 ° 58 ′ 59.5 ″ S 29 ° 37 ′ 09.8 ″ E], Malaise trap, dense Leucosedea dominated scrub, leg. A. H. Kirk-Spriggs, coll. BMSA; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Kat- berg, det. as P. constricta by Kröber 1938, det. as P. constricta by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922174]; 1 ♂, 1. - 10. ii. 1933, Eastern Cape Province, Katberg, det. as P. constricta by Kröber 1938, det. as P. constricta by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922175]; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as P. constricta by Kröber 1938, det. as P. constricta by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922177]; 1 ♂, 1. – 10. ii. 1933, Eastern Cape Province, Katberg, det. as P. constricta by Kröber 1938, det. as P. constricta by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922178]; 1 ♀, 19. – 26. ii. 1933, Eastern Cape Province, Katberg, det. as P. ruficoxa by Kröber 1938; det. as P. constricta by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922173]; 1 ♂, 6. i. 1925, East- ern Cape Province, Lady Grey, paratype of P. ruficoxa, leg. R. I. Nel, coll. NHML [NHMUK 010922180]; 1 ♂, 26. i. 1960, Eastern Cape Province, Molteno, leg. M. D. Anderson, coll. AMGS; 1 ♂, 4. – 5. ii. 1992, Eastern Cape Province, Rhode, village area [30 ° 48 ′ S 27 ° 58 ′ E], 1820 m, det. as P. constricta by Camras 2000, leg. Natal Muse- um Expedition, coll. FMNH; 1 ♂, 22. i. 1965, Free State Province, Senekal, leg. D. J. Brother, coll. AMGS; 1 ♂ 1 ♀, 12. ii. 2016, KwaZulu-Natal Province, Garden Castle N. R., Sleeping Bea [29 ° 44 ′ 56 ′ S 29 ° 10 ′ 35 ″ E], 2190 m, leg. RSA-team Ståhls, coll. PASS; 1 ♂, 4. xii. 2012, Kwa- Zulu-Natal Province, Royal Natal National Park, day visitor Ca [28 ° 41 ′ 25.4 ″ S 28 ° 56 ′ 53.9 ″ E], 1410 m, Para- type, leg. A. Ssymank, coll. PASS; 1 ♀, 20. – 21. xi. 2003, Mpumalanga Province, 20 km sw Lydenbrug, leg. J. Halada, coll. CULSP; 1 ♂, iii. 1933, Natal Province, Na- tal, Nationalpark, det. as P. constricta by Camras 2000, leg. J. Ogilvie, coll. NHML [NHMUK 010922179]; 1 ♂, 13. iv. 1963, Northwest Reg., Retiefs Kloof, Rustenburg, leg. H. N. Empey, coll. PMHA; UNKNOWN LOCA- TION: 1 ♂ ,, “ 3054 ”, det. as P. platycephala by Kröber 1914, wrongly labelled as allotype of P. platycephala, leg., coll. ZMHB; 2 ♀♀ ,, “ AcP 5470 ”, leg., coll. NHML; 1 ♀, no date, “ AcP 5470 ”, [collector unknown], coll. NHML; 1 ♂, 13. xi. 1954, [characters illegible], det. as P. constricta by Camras 2000, leg. K. G. V. Smith, coll. NHML [NHMUK 010922176]. Females of P. constricta are easily identified by the unique shape of the abdomen and theca as described in Key 6 (Figs 76, 77). Males are extremely difficult to di- agnose, however, and can be confused with members of the abdominalis, microvena, caenoneura and antiqua species-groups. Males of P. constricta should be compared very carefully, bearing in mind that not all speci- mens can be reliably assigned to species.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD38FFE725D84C35FEFB670A.taxon	description	(Figs 84 – 90) Conops (Physocephala) digitatus Speiser 1909	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD38FFE725D84C35FEFB670A.taxon	description	= Physocephala ethiopica Camras, 1962 (syn. nov.)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD38FFE725D84C35FEFB670A.taxon	materials_examined	Primary type material examined. ♂ holotype of Pseudophysocephala ugandae Kröber 1939: (1) “? Psilocephala [sic] / ugandae, Krb. / examined & det. / O. Kröber, 1938. ”; (2) “ Psilocephala [sic] / ugandae / Krb ”; (3) “ Uganda, / Kampala, / 6. IX. 1918. / C C. Gowdey. ”; (4) “ Kampala, / Uganda / 6. IX. 1918 / No. 6545 ”; (5) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1921 - 153 ”; (6) “ Type ”; coll. NHML. ♂ holotype of Physocephala nigricoxa Brunetti, 1925: (1) “ Syn- type ”; (2) “ Kampala, / Uganda / 4. IX. 1918 / C. C. / No. 5392 Gowdey ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306. ”; (4) “ Physo. / nigrocoxae [sic] / Brun Type ♂ / Det. E. Brunetti 1924 ”; coll. NHML. There is a second damaged specimen lacking both head and abdomen that is designated as “ Syntype ”. This can- not be identified but in any event, in the original descrip- tion the specimen listed above as the holotype is explicit- ly identified as the “ Type ”. ♀ holotype of Physocephala ethiopica Camras, 1962, pinned together with ♂ paratype: (1) “ Holo- / type ”; (2) “ Allo- / type ”; (3) “ Ethiopia: / Dilla, / (Sidamo). / iv. 1948. ”; (5) “ Holotype ♀ / Physocephala / ethiopica / Camras ”; (6) “ Allotype ♂ ”; (7) “ K. M. Guichard / B. M. 1948 - 200. ”; (8) “ [characters illegible] ”; (9) “ BMNH (E) # / 249079 / BMNH (E) # / 249080 ”; coll. NHML. Additional material. BURUNDI: 1 ♀, V. 1956, Mugera, leg. F. J. François, coll. ISNB; CAMEROON: 1 ♀, 30. vii. 1920, Lolodorf, det. as P. simplex by Camras 1962, leg. A. I. Good, coll. FMNH; 1 ♂, 8. – 10. viii. 2013, Far- North Reg., Mayo-Tsanga, Mogode-Cha [10 ° 34.111 ′ N 13 ° 36.422 ′ E], 1001 m, Malaise trap, degrated savanna forest, leg. A. H. Kirk-Spriggs, coll. BMSA; DEMO- CRATIC REPUBLIC OF CONGO: 2 ♂♂, 15. ix. 1933, Bambesa, det. as P. ugandae by Kröber 1935, leg. J. V. Leroy, coll. MRAC; 1 ♀, 16. iii. 1948, Basoko, Yamabuki, Div. 153, det. as P. simplex by Camras 1962, leg. P. L. G. Benoit, coll. MRAC; 1 ♂, 27. iv. 1914, Beni [0 ° 30 ′ N 29 ° 30 ′ E], det. as P. bequaertorum by Camras 2000, leg. J. Bequaert, coll. FMNH; 1 ♀, 23. i. 1927, Elisabethville [Lubumbashi] [- 11.664232 27.482626], det. as P. ugandae by Kröber 1935, leg. M. Bequaert, coll. MRAC; 1 ♂, 4. iii. 1936, Kamogobe, Sud Masisi, det. as P. nigricoxa by Camras 1962, leg. L. Lippens, coll. MRAC; 1 ♂, 18. vii. 1935, Kivu, Nyongera, près Rutshuru, 1218 m, leg. G. F. de Witte, coll. ISNB; 1 ♂, 10. vii. 1935, Kivu, Ritshu- ru, riv. Musugereza, 1100 m, leg. G. F. de Witte, coll. ISNB; 1 ♂, 10. vii. 1935, Kivu, Ritshuru, riv. Musugere- za, 1100 m, det. as P. nigrocoxalis by Vanschuytbroeck 1950, leg. G. F. de Witte, coll. MRAC; 1 ♀, 15. x. 1957, Massif Ruwenzori, Bomboka près., Kyandolire, 1650 m, leg. P. Vanschuytbroeck & J. Kekenbosch, coll. MRAC; 1 ♀, 18. xii. 1957, Massif Ruwenzori, riv. Lume, moy- enne, 1800 m, leg. P. Vanschuytbroeck, coll. MRAC; 1 ♀, xi. 1937, Rutshuru, det. as P. digitata by Camras 2000, leg. J. Ghesquière, coll. FMNH; 1 ♀, no date, Ville Malela, Chief Casende [5,40 S 23,45 E], det. as P. simplex by Camras, leg. J. Bequaert, coll. FMNH; RWANDA: 1 ♂, 6. ii. 1935, Ruhengeri, riv. Mugara-Kigombe, 1800 – 1825 m, det. as P. nigrocoxalis by Vanschuytbroeck 1950, leg. G. F. de Witte, coll. MRAC; TOGO: 1 ♂, viii. 2015, Kloto, forest area [6 ° 57 ’ 31.66 ’’ N 0 ° 34 ’ 29.75 ’’ E], leg. G. Goergen, coll. IITA; 1 ♀, ii. 2002, ditto; 1 ♀, iii. 2017, ditto; 1 ♂, viii. 2008, ditto; 1 ♂, vi. 2016, ditto; UGAN- DA: 1 ♂, 22. iv. 1966, 20 miles e Mubende, det. as P. nigricoxa by Camras 2000; det. as P. digitata by Camras 2001, at Chickweed flowers, leg. D. J. Greathead, coll. NHML [NHMUK 010922194]; 1 ♂, 14. v. 1958, Bugisu Bugusege, det. as P. nigricoxa by Camras 2000; det. as P. digitata by Camras 2001, at Chickweed flowers, leg. J. Bowden, coll. NHML [NHMUK 010922195]; 1 ♂, 1. – 5. xii. 1911, Bugoma Forest, Unyoro, 3700 ft, det. as P. nigricoxa by Camras 2000; det. as P. digitata by Cam- ras 2001, at Chickweed flowers, leg. S. A. Neave, coll. NHML [NHMUK 010922192]; 1 ♂, 21. iv. 1927, Dwoli, det. as P. nigricoxa by Camras 2000; det. as P. digitata by Camras 2001, leg. H. Hargreaves, coll. NHML [NHMUK 010922193]; 1 ♂, 15. – 17. iii. 1912, Uganda West, Kibale Region, surrounding Bigodi [N 0029209 E 3020022], 1400 m, leg. Oehlke, coll. PASS; 1 ♂, 15. – 17. iii. 2012, ditto; UGANDA / KENYA: 1 ♂, v., Elgon, 1700 m, leg. Lindblom, coll. NHRS; ZAMBIA: 7 ♂♂ 1 ♀, 12. – 15. i. 2003, 45 km se Kitwe, leg. J. Halada, coll. CULSP. Physocephala digitata is a species which shows great variation in colouration of the face (with or without dark marking); frons (more or less uniformly yellow to light brown, the latter especially in old specimens, and with distinct black midline or almost completely black); coxae (completely light brown to almost black); scutum (completely black to orange-brown with black centre only); and wing (e. g. cell r 2 + 3 completely brown or hy- aline apically, subcosta light orange to dark brown). In all of these variations we find different intermediates and no one of these characters, or combination of characters, can be used to consistently segregate species. Camras (2001) did not mention any character in his key for the simplex species-group which is not variable in P. digitata. Out of the eleven species he included in his simplex species-group key, only three species can be recognised consistently (see Key 6, couplets 8 – 9) and these three are still difficult to identify convincingly. The original description of P. simplex falls within the variation of P. digitata as do the specimens reported by Camras (2001) as P. simplex. The holotype of P. simplex is stored in the Museo Civico di Storia Naturale “ Giaco- mo Doria ” (Stuke 2017) and was not available for exam- ination. Physocephala simplex Kröber, 1915 is, however, placed as a junior synonym of Physocephala [Conops] digitata (Speiser, 1909) (syn. nov.). Most specimens of P. ugandae are easily recognised by the wing pattern (Fig. 89) with cell r 2 + 3 hyaline apical- ly, and the frons yellow with a distinct black T-marking (Fig. 84). There are, however, less commonly encoun- tered forms typified by a completely dark cell r 2 + 3 and / or either a completely dark or completely pale frons, togeth- er with intermediates. Since this variation does not allow P. ugandae to be distinguished from P. digitata with any consistency, Physocephala ugandae Kröber, 1915 is herewith placed as a junior synonym of Physocephala [Conops] digitata (Speiser, 1909) (syn. nov.). The original description and all of the key characters used by Camras (1962 b) to identify P. bequaertorum fall within the variation of P. digitata. One available headless specimen in FMNH identified by Camras as P. bequaertorum also fits to P. digitata. Physocephala bequaertorum Camras, 1962 is therefore placed as a junior syn- onym of Physocephala [Conops] digitata (Speiser, 1909) (syn. nov.). The type material of P. ethiopica falls within the varia- tion of P. digitata. Physocephala ethiopica Camras, 1962 is therefore placed as a junior synonym of Physocephala [Conops] digitata (Speiser, 1909) (syn. nov.). The original description of P. lineifrons compares it with P. bequaertorum, stating that the two differ only in the colouration of the gena and frons. The holotype is deposited in the Natal Museum (KwaZulu-Natal, Pi- etermaritzburg) and was not available for examination, but the original description of P. lineifrons falls within the variation of P. digitata and therefore Physocephala lineifrons Camras, 1962 is also placed as a junior syn- onym of Physocephala [Conops] digitata (Speiser, 1909) (syn. nov.).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE725F24DBEFC9566C1.taxon	description	(Figs 91 – 93)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE725F24DBEFC9566C1.taxon	materials_examined	Primary type material examined. ♂ holotype of Pseudophysocephala annulipes Kröber 1939: (1) “ Holo- / type ”; (2) “ Kinangop / below bamboo / forest 8500 ft ”; (3) “ Kenya: / Aberdare Range / x. 1934. / B. M. E. Afr. Exp. / B. M. 1935 - 203 ”; (4) “ Pseudophysocephala / annulipes. Krb / examined & det. / O. Kröber 1938 ”; (5) “ Pseodophysoce / phala annuli - / pes Krb ”; (6) [blank red label]; coll. NHML. Additional material. KENYA: 1 ♂, 22. ix. 1990, Mount Kenya, ≈ 10.000 ft, sweep netting, leg. R. Copeland, coll. IITA; 1 ♀, i. – ii. 1946, N. W. Mau Forest, 8000 – 10000 ft, det. as P. annulipes by Camras 1962, leg. H. P. Thomma- set, coll. NHML [NHMUK 010922136].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE7267F4FAFFC9564BB.taxon	materials_examined	Primary type material examined. ♂ holotype of Physocephala maculifacies Camras, 2001: (1) “ Angola (A 36) / Chianga / 21. – 24. iii. 1972 ”; (2) “ Southern / African Exp. / B. M. 1972 - 1 ”; (3) “ Holotype ♂ / Physocephala / maculifacies / Camras ”; coll. NHML. Additional material. ANGOLA: 1 ♀, 21. – 24. iii. 1972, Chianga, paratype of P. maculifacies, leg. British Mu- seum Expedition, coll. NHML [NHMUK 010922096]; TANZANIA: 1 ♂, 10. iii. 1963, Kilimajaru, Weruweru, det. as P. abyssinica by Camras 2000, leg. D. J. Great- head, coll. NHML [NHMUK 010922099]. A difficult species for which more material is necessary to test the stability of the identification characters given in Key 6.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE725F24EE5FDD964FE.taxon	materials_examined	Material. UGANDA: 1 ♂, vii. 1945, Bwamba Valley, det. as P. discalis by Camras 2000, leg. van Someren, coll. NHML [NHMUK 010922150]. The interpretation of this species is based only upon the above specimen in the NHML which was identified by Camras as P. discalis following his comparison with the holotype of that species at the USNM.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE3267F4C7BFDC06472.taxon	description	(Figs 94 – 101) Physocephala microvena Brunetti 1925	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3BFFE3267F4C7BFDC06472.taxon	materials_examined	New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 253 Primary type material examined. 1 ♀ syntype of Physocephala microvena Brunetti 1925: (1) “ Syn- / type ”; (2) “ H. S. Stannus / Zomba. / Nyasaland. ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physo. / microvena / Brun Type ♀ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249157 ”; coll. NHML. 1 ♂ syntype of Physocephala microvena Brunetti 1925: (1) “ Syn- / type ”; (2) “ H. S. Stannus / Zomba. / Nyasa- land. ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physo. / microvena / Brun / Type ♂ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249156 ”; coll. NHML. 1 ♂ syntype of Physocephala microvena Brunetti 1925: (1) “ Syn- / type ”; (2) “ H. S. Stannus / Zomba. / Nyasa- land. ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physo. / microvena / Brun / Cotype ♂ / Det. E. Brunetti 1924 ”; (5) “ BMNH (E) # / 249158 ”; coll. NHML. 1 ♀ syntype of Physocephala microvena Brunetti 1925: (1) “ Syn- / type ”; (2) “ H. S. Stannus / Zomba. / Nyas- aland. ”; (3) “ Physo. / microvena / Brun / Cotype ♂ / Det. E. Brunetti 1924 ”; (4) “ BMNH (E) # / 249158 ”; coll. NHML. 1 ♀ syntype of Physocephala microvena Brunetti 1925: (1) “ Syn- / type ”; (2) “ H. S. Stannus / Zomba. / Nyasa- land. ”; (3) “ Pres. by / Imp. Bur. Ent. / Brit. Mus. / 1924 - 306 ”; (4) “ Physocephala / microvena Brun. / ♂. Cotype. / (Det. E. Brunetti 1924) ”; (5) “ BMNH (E) # / 249160 ”; coll. NHML. ♀ holotype of Pseudophysocephala nigritarsis Kröber 1939: (1) “ Holo - / type ”; (2) “ Type ”; (3) “ Fort Portal / Nyakasura / 24. i. 1935 / J. W. Edwards ”; (4) “ Uganda: / Ruwenzori Range / xii. 1934 – i. 1935. / B. M. E. Afr. Exp. / B. M. 1935 - 203 ”; (5) “ Pseudophysocephala / nigritarsis, Krb. / examined & det. / O. Krober, 1938. ”; (6) “ Pseodophysoce - / phala nigri - / tarsis Krb ”; (7) “ BMNH (E) # / 249147 ”; coll. NHML. Additional material. BENIN: 1 ♀, 1. xii. 2011, Lokoli, swamp forest [07 ° 03 ’ 40.00 ″ N 02 ° 15 ’ 50.00 ″ E], leg. G. Goergen, coll. IITA; BURUNDI: 2 ♀♀, 5. – 12. iii. 1953, Bururi, 1800 – 2000 m [- 3.949 29.623], det. as P. nigritarsis by Camras 1962, leg. P. Basilewsky, coll. FMNH, MRAC; 1 ♀, 4. vi. 1952, Colline Muramba, Bugoni, Terr. de Muhinga [- 2.930 30.357], 1400 m, det. as P. nigritarsis by Janssen 1955, leg. F. J. François, coll. ISNB; CAM- EROON: 1 ♂, no date, Neu Kamerun, det. as P. nigritarsis by Camras 2000, leg. G. Tessmann, coll. ZMHB; DEMOCRATIC REPUBLIC OF CONGO: 1 ♀, no date, Elisabethville [Lubumbashi] [- 11.664232 27.482626], det. as P. microvena by Camras 1962, leg. M. Bequaert, coll. FMNH; 1 ♀, 10. xi. 1928, Ituri, Blukwar, syntype of P. curta but not conspecific with designated lectotype, leg. A. Collart, coll. MRAC; 1 ♀, 12. iv. 1953, Kivu, Kiten- ga, près riv. Ruzizi [- 6.896 25.975], leg. J. Verbeke, coll. ISNB; 1 ♀, 9. iv. 1945, Rumangabo, riv. Bugombwa, det. as P. gigantea by Janssens 1950, leg. G. F. de Witte, coll. MRAC; 1 ♀, 29. iii. 1954, Secteur Tshiaberimu, riv. Kaliv- ina, affl. Talia Nord, 2350 m [- 0.13 29.42], 2350 m, leg. P. Vanschuytbroeck & H. Synave, coll. MRAC; 1 ♂, 21. – 27. viii. 1931, Tshibinda [- 0.287 28.776], det. as P. nigritarsis by Camras 1962, leg. J. Ogilvie, coll. FMNH; KENYA: 1 ♂, 12. iii. 1993, 5 km e Kaimosi, se Kakamega [0.125 34.892], 1750 m, leg. B. Merz, coll. TAUI; 1 ♂, 10. x. 1998, Bungoma [0.5645 34.558], leg. F. Kaplan, A. Freidberg, coll. TAUI; 2 ♂♂, 12. – 13. i. 1996, Bungo- ma [0.5645 34.558], leg. I. Yarom, A. Freidberg, coll. TAUI; 1 ♀, 8. x. 1998, Gilgil [- 0.500043 36.326122], leg. F. Kaplan, A. Freidberg, coll. TAUI; 3 ♂♂ 1 ♀, 10. x. 1998, Kakamega [0.281 34.753], leg. F. Kaplan, A. Freidberg, coll. TAUI; 1 ♂, 7. – 8. ii. 2011, Kakamega Forest, 1586 m [0 ° 13 ′ 37.2 ″ N 34 ° 52 ′ 49.8 ″ E], 1586 m, leg. M. Mei, P. Ceretti, D. Whitmore, coll. PMME; 1 ♂, 1. – 10. ii. 2002, Kakamega Isecheno Nature Reserve [0.24 ′ N 34.86 ′ E], 1800 m, leg. Okeka, coll. PMHA; 1 ♂, 11. – 20. i. 2003, ditto; 1 ♂, iv. 1969, Karen nr Nairobi, leg. van Someren, coll. NHML [NHMUK 010922149]; 1 ♀, 9. x. 1998, Keri- cho [- 0.368726 35.281219], leg. F. Kaplan, A. Freidberg, coll. TAUI; 1 ♂, 11. – 12. iv. 1957, Molo, Mau Escarpment, 2150 – 2200 m, det. as P. nigritarsis by Camras 1962, leg. P. Basilewsky, N. Leleup, coll. MRAC; 1 ♂, ix. 1939, Naivasha, det. as P. nigritarsis by Camras 1999, leg. H. J. A. Turner, coll. NHML [NHMUK 010922138]; 1 ♀, ix. 1939, Naivasha, det. as P. nigritarsis by Camras 2000, leg. H. J. A. Turner, coll. NHML [NHMUK 010922141]; 1 ♂, 12. iii. 1993, near Kaimosi, se Kakamega [0.125606 34.844837], 1750 m, leg. Muhangani, coll. TAUI; 1 ♀, 16. xi. – 22. xii. 1998, Rift Valley Province, Mpala research Station [00 ° 19 ′ N 36 ° 53 ′ E], Malaise trap, leg. S. Miller, coll. NMKE; 1 ♂, 2. – 9. i. 2000, Western Province, Ka- kamega Forest, Malaise trap [0 ° 14.13 ′ N 34 ° 51.87 ′ E], leg. R. Copeland, coll. NMKE; UGANDA: 1 ♂, 22. iv. 1966, 20 miles e Mubende, leg. D. J. Greathead, coll. NHML [NHMUK 010922147]; 1 ♂, 19. x. 1958, Kawanda, det. as P. nigritarsis by Camras 2000, [collector unknown], coll. NHML [NHMUK 010922137]; 1 specimen, xii. 1934 – i. 1935, Ruwenzori Range, det. as P. nigritarsis by Camras 2000, leg. B. M. E. Africa Expedition, coll. NHML [NHMUK 010922140]; 1 ♂, 20. iii. 2012, Rwenzo- ri Gebirge, Nyakalengila [N 002100 E 300149], 1705 m, leg. Oehlke, coll. PASS; 1 ♂, 16. xi. 1949, Shangugu [= Cyangugu] [- 2.483333 28.896667], 1460 m, det. as P. platycephala by Janssen 1954, leg. F. J. François, coll. ISNB; ZIMBABWE: 1 ♀, 2. – 6. xii. 2015, Chirinda Forest, Mt. Selinda [20 ° 25 ′ S 32 ° 43 ′ E], 1000 m, leg. J. Halada, coll. CULSP; 1 ♂, 18. vi. 1964, n Vumba [= Bvumba] [- 19.093432 32.740631], det. as P. microvena Camras 2000, leg. D. Cookson, coll. FMNH; 2 ♂♂, 4. vii. 1964, n Vumba [= Bvumba] [- 19.093432 32.740631], det. as P. microvena Camras 2000, leg. D. Cookson, coll. FMNH; 1 ♀, 18. vii. 1964, n Vumba [= Bvumba] [- 19.093432 32.740631], det. as P. microvena by Camras 2000, leg. D. Cookson, coll. FMNH; 1 ♀, 8. x. 1964, n Vumba [= Bvum- ba] [- 19.093432 32.740631], det. as P. microvena by Cam- ras 2000, leg. D. Cookson, coll. FMNH; 1 ♂, 20. ii. 1965, n Vumba [= Bvumba] [- 19.093432 32.740631], det. as P. microvena Camras 2000, leg. D. Cookson, coll. FMNH; 1 ♀, 8. v. 1965, n Vumba [= Bvumba] [- 19.093432 32.740631], det. as P. microvena by Camras 1999, leg. D. Cookson, coll. FMNH; 1 ♀, 8. v. 1965, n Vumba [= Bvum- ba] [- 19.093432 32.740631], det. as P. microvena by Camras 2000, leg. D. Cookson, coll. FMNH; 1 ♀, v. 1932, Xmas Pas, Umtali [= Christmas Pass near Mutare], leg. J. Ogilvie, coll. NHML [NHMUK 010922146]; 1 ♂, v. 1932, ditto [NHMUK 010922148]. Most P. microvena specimens characteristically have veins M and R 4 + 5 fused, thus lacking any radial-medial crossvein (Figs 98, 100). A minority of specimens do have a very short radial-medial crossvein present, how- ever, and a reduced crossvein rm may also occur occa- sionally in specimens of other species, causing complica- tions in identification. Females of P. microvena are easily identified by the short, broad theca (Fig. 97) and short abdominal segments (Fig. 96), with confusion only really likely with P. pseudomicrovena, which may be conspe- cific. Conversely, males of P. microvena are extremely difficult to identify reliably and could be easily confused with males of other species-groups and also with males of P. constricta where these also have a reduced radial-medial crossvein. Where present, the distance between 256 Jens-Hermann Stuke & David K. Clements the two short dark submedial stripes in the microtrichial dusting on the anterior scutum may be the best character for separating P. constricta and P. microvena, these being separated by a wider medial dusted stripe in the former (Fig. 81) than in the latter (Fig. 99). As mentioned by Camras (2001) there are several inter- mediates between typical P. microvena and P. nigritarsis. Therefore, Pseudophysocephala nigritarsis Kröber, 1939 is herewith placed as a junior synonym of Physocephala microvena Brunetti, 1925 (syn. nov.).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3FFFE325F24D1DFA9B6411.taxon	description	(Figs 102 – 105)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3FFFE325F24D1DFA9B6411.taxon	materials_examined	Material. SOUTH AFRICA: 1 ♂, 3. – 7. x. 1988, Cape Province, Clanwilliam Dam [32 ° 11 ′ 30 ″ S 18 ° 53 ′ 42 ″ E], on flowers of Aspalathus desertorum Bol., leg. F. W. Gees, S. K. Gees, coll. AMGS; 1 ♀, ix. 1928, Cape Prov- ince, Worcester, det. as Physocephala spec. by Kröber 1938, det. as P. platycephala by Camras 2000, leg. R. E. Turner, coll. NHML [NHMUK 010922130]. While the female of P. platycephala is easily recognised by the obvious shape of the postabdomen and theca (Figs 102, 104), males are more problematic. We doubt that males can be reliably identified using the colour pattern of the scutum as proposed by Camras (2001) and as de- scribed in Key 6. Figs 103 & 105 show that the black spots on the scutum are less distinctly separated in males (Fig. 105) compared to females (Fig. 103), and may per- haps sometimes become fused. Perhaps males of this rare species – only two female specimens are known – have therefore previously been misidentified as other species.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3FFFFC267F4D1DFBDB6527.taxon	description	(Figs 106 – 107)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD3FFFFC267F4D1DFBDB6527.taxon	materials_examined	Primary type material examined. ♂ lectotype of Physocephala pseudomicrovena Kröber, 1939 designated by Camras (1962): (1) “ ♂ / Holotypus ”; (2) “ Musée du Congo / Tanganyka-Moero: / Nyunzu - I - II - 1934 / De Saeger ”; (3) “ R. DÉT / O / 2992 ”; (4) “ Physocephala ♂ / microvena Brun. / var. / det. Kröber 1935 ”; (5) “ RMCA New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 257 ENT / 000012181 ”; (7) “ Type ♂ / Physocephala / pseudomicrovena / Kröber ”; (7) “ designated by / Camras, 1962 ”; coll. MRAC. Additional material. BENIN: 1 ♂, viii. 2006, Athiémé [06 ° 14 ’ 20.00 ″ N 01 ° 40 ’ 00.00 ″ E], leg. G. Goergen, coll. IITA; 2 ♀♀, viii. 2006, Athiémé [06 ° 14 ’ 20.00 ″ N 01 ° 40 ’ 00.00 ″ E], sweep netting, leg. G. Goergen, coll. IITA; 1 ♀, v. 2006, Lokossa, sweep netting, leg. G. Goer- gen, coll. IITA. The lectotype is in poor condition, with the antennae missing and a broken abdomen almost completely cov- ered with glue. All of the characters which can be seen fit well to the concept of the species as given in Key 6, how- ever. Neverthess, this species is only poorly differentiat- ed from P. microvena, and it may easily be no more than a dark morph of that species. On the other hand, all of the specimens available to us can be distinguished with the characters given in Key 6 and therefore the species is here accepted as valid.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD21FFFD267F4E86FBAC649C.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala basilewskyi Camras, 1962: (1) “ ♀ / Ho- lotypus ”; (2) “ Coll. Mus Congo / Kib. - Ituri: Irumu / 18 / XII- 1952 / P. Basilewsky ”; (3) “ Holotype ♀ / Pseudophysocephala / basilewskyi / Camras ”; (4) “ RMCA ENT / 000012176 ”; coll. MRAC.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD21FFFF267F4D99FC3F62EE.taxon	description	(Figs 108 – 113)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD21FFFF267F4D99FC3F62EE.taxon	materials_examined	Primary type material examined. ♂ holotype of Conops nitidus Kröber 1915 f: (1) “ Togo / Bismarckburg / 2. – New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 259 18. vi. 93 / L. Conradt S. ”; (2) “ Conops ♂ / nitidus Kröb / O. Kröber det. 1914 ”; (3) “ Cotype ”; (4) “ Zool. Mus. Ber- lin ”; (5) “ Pseudophysocephala / nitidus / Kröber / det. Camras, 2000 ”; coll. ZMHB. Additional material. BENIN: 1 ♂, x. 2008, Ahozon, for- est area [06 ° 22 ’ 57.59 ″ N 02 ° 9 ’ 15.92 ″ E], leg. G. Goergen, coll. IITA; 1 ♀, 26. iii. 2013, Dangbo, sacred forest, leg. G. Goergen, coll. IITA; KENYA: 1 ♂, 12. – 26. ii. 2005, Nyanza Province, Ungoye Field Station [0 ° 36.91 ′ S 34 ° 05.52 ′ E], 1147 m, leg. R. Copeland, coll. NMKE; 1 ♂, 27. ii. – 6. iii. 2005, ditto; TANZANIA: 1 ♀, 1. vi. – 4. vii. 1893, Bismarckburg [= Kasanga], det. as P. nitida by Kröber 1914 and Camras 2000, wrongly interpreted as female syntype, leg. L. Conradt S., coll. ZMHB; TOGO: 1 ♂, 26. i. 2016, Dzobégan, monastery [7 ° 14 ’ 22.07 ″ N 0 ° 41 › 58.64 ″ E], leg. G. Goergen, coll. IITA; 1 ♂, ix. 2006, Kloto, forest area [6 ° 57 ’ 31.66 ″ N 0 ° 34 › 29.75 ″ E], leg. G. Goergen, coll. IITA; 1 ♂, viii. 2008, ditto; 1 ♂, i. 2005, dit- to; 1 ♀, vi. 2008, ditto; 1 ♀, x. 2016, ditto. 260 Jens-Hermann Stuke & David K. Clements	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD23FFF825F24D61FD5E66AD.taxon	description	(Figs 114 – 118) Physocephala pilitarsis Kröber 1936	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD23FFF825F24D61FD5E66AD.taxon	materials_examined	Additional material. CENTRAL AFRICAN REPUB- LIC: 1 ♂, 14. vi. 2009, 150 km nww Mbaiki [04 ° 05 ′ N 17 ° 02 ′ E], 620 m, leg. J. Halada, coll. CULSP; 1 ♀, 2. v. 2010, 20 km nne Mbaiki [03 ° 04 ′ N 18 ° 00 ′ E], 450 m, leg. J. Halada, coll. CULSP; 1 ♂, 9. xi. 2012, 40 km e of Bambio [3 ° 60 ′ N 17 ° 12 ′ E], 500 m, leg. J. Halada, coll. CULSP; 1 ♂, 9. xii. 2008, 50 km ne Bambio [03 ° 59 ′ N 17 ° 11 ′ E], 450 m, leg. J. Halada, coll. CULSP; 3 ♂♂, 4. xii. 2010, 60 km w Banibo [3 ° 50 ′ N 16 ° 44 ′ E], 630 m, leg. J. Halada, coll. CULSP; 1 ♂, 24. – 28. xi. 2010, 70 km nne Bangui [04 ° 57 ′ N 18 ° 46 ′ E], 445 m, leg. J. Halada, coll. CULSP; DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, xii. 1923, Katompe, Katanga, det. as “? abyssinica ” by Brunetti 1925, det. as P. pilitarsis by Camras 1962, leg. Bequaert, coll. MRAC; 1 ♂, viii. 1932, Lulua, Kapan- ga, det. as P. “ abyssinica Krb var. ” by Kröber 1935, leg. F. G. Overlaet, coll. MRAC; 1 ♂, xi. 1932, Lulua, Kapan- ga, det. as P. “ abyssinica Krb var. ” by Kröber 1935, leg. F. G. Overlaet, coll. MRAC; 1 ♂, 1. iv. 1937, Terr. Lisala, det. as P. pilitarsis by Camras 1962, leg. Leontovich, coll. MRAC; EQUATORIAL GUINEA: 1 ♀, 16. – 31. viii. 1906, Alén, Benitogebiet, det. as P. pilitarsis by Camras 2000, leg. G. Tessmann, coll. ZMHB; 1 ♀, 1. – 15. xi. 1906, Alén, Benitogebiet, det. as P. pilitarsis by Camras 2000, leg. G. Tessmann, coll. ZMHB; 2 ♂♂, vi. – viii. 1908, Uelleburg, det. as P. pilitarsis by Camras 2000, leg. G. Tessmann, coll. ZMHB; KENYA: 1 ♂, 13. – 27. viii. 2006, Western Province, Kakamega Forest near Rondo Guest House [0.22767 ° N 34.88533 ° E], 1630 m, Malaise trap set across small permanent stream, leg. R. Copeland, coll. NMKE; 1 ♀, 3. – 17. xii. 2006, ditto; 1 ♀, 13. – 27. viii. 2006, ditto; 1 ♂, 12. – 19. iii. 2000, Western Province, Kakame- ga Forest, Malaise trap [0 ° 14.13 ′ N 34 ° 51.87 ′ E], leg. R. Copeland, coll. NMKE; 1 ♂, 28. viii. – 4. ix. 1999, ditto; 1 ♂, 2. – 9. i. 2000, ditto; 1 ♀, 20. – 27. xi. 1999, ditto; 2 ♀♀, 17. – 24. vii. 1999, ditto; 1 ♀, 13. – 20. ii. 2000, ditto; 2 ♀♀, 11. – 18. ix. 1999, ditto; 2 ♀♀, 3. – 10. vii. 1999, ditto; 1 ♀, 21. – 28. v. 2000, ditto; UGANDA: 1 ♂, vi. 1912, Kawanda, det. as P. pilitarsis by Camras 2000, leg. D. J. Greathead, coll. NHML [NHMUK 010922154]; 1 ♂, 2. viii. 1962, Ma- bira Forest, det. as P. pilitarsis by Camras 2000, leg. D. J. Greathead, coll. NHML [NHMUK 010922155].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD24FFF825D84E86FE1165E5.taxon	description	(Figs 119 – 121)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD24FFF825D84E86FE1165E5.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala spinipes Camras 2001: (1) “ Tanganyika / Amani / 1957 / J. G. Halcrow / in thick bush ”; (2) “ C. I. E. Coll. / NO 15485 ”; (3) “ Pseudophysocephala / sp.? / Det. R. L. Coe. 1958 ”; (4) “ Pres. by / Com Inst Ent / B M 1969 - 3 ”; (5) “ Holotype ♀ / Pseudophysocephala / spinipes / Camras ”; coll. NHML. Additional material. KENYA: 1 ♀, 23. ix. 2005, Ka- kamega Forest, Yala River [0 ° 13 ′ N 34 ° 53 ′ E], 1450 m, leg. A. Freidberg, coll. TAUI.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD24FFF8265A4A6CFAE061F8.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala tetratarsata Camras 2001: (1) “ Yaoundé Cam. / Nkolbisson / Nonvll. X 1974 ”; (2) “ parasitizing 6 ”; (3) “ 61 a ”; (4) “ C. I. E. Coll. / A. 7811 ”; (5) “ Pres by / Comm Inst Ent / B M 1975 - 1 ”; (6) “ Holotype ♀ / Pseudophysocephala / tetratarsata / Camras ”; (7) “ Pseudophysocephala / tetratarsata ” and on underside of this label “ Pseudophyso. / BM. Mus. From. / Yaounde Cam ”; coll. NHML. Below the holotype is a large label with additional infor- mation: “ Following and ovipositing on bees which were collecting sap flowing from Scolyid entrance holes in felled trees. Each was following a bee flying immediate- ly behind it. When a Physocephala managed to approach closely a bee the fly deposited an egg on its body with an extreme rapidity and I could establish later that the egg remained attached to the tegument of the bee. Dr. Gui- do Nonveiller in letter 6.4.76 to K. M. Harris. C. I. E. ” [Commonwealth Institute of Entomology].	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD24FFFA265A48B4FD4367F6.taxon	description	(Figs 122 – 125) Physocephala vitripennis Curran 1928 = Physocephala intermedia Kröber 1936 (syn. nov.) = Conops bouvieri Séguy 1936 (syn. nov.)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD24FFFA265A48B4FD4367F6.taxon	materials_examined	Primary type material examined. ♂ holotype of Physocephala intermedia Kröber, 1936: (1) “ Ph. Type ♂ / intermedia ”; (2) “ Musée du Congo / Ubangi: Nzali / 3 / 4 - II- 1932 / H. J. Brédo ”; (3) “ R. DÉT / Z / 2992 ”; (4) “ Physocephala ♂ / intermedia Krb. / det. Kröber 1935 ”; (5) “ Type ”; (6) “ RMCA ENT / 000012179 ”; coll. MRAC. ♀ holotype, ♂ paratype (pinned on one needle) of Pseudophysocephala meii Camras 2001: (1) “ Repub- lique de Guinea PNHN 5 / 10 ° 16 ’ 43 ’’ N 10 ° 26 ’ 02 ’’ W / Faranen, Sidakoro, mangor, / 29. XII. 1995, leg. M. Mei ”; (2) “ Holotype ♀ and ♂ / Pseudophysocephala / meii / Camras ”; (3) [glued male abdomen]; (4) “ Pseudophysocephala / cf vitripennis Curran / M. Mei det. 1997 ”; (5) “ Pseudophysoc / meii ”; coll. FMNH. Additional material. BENIN: 1 ♂, 20. iii. 2011, Niaou- li, forest area [06 ° 44 ’ 3.08 ″ N 02 ° 8 ’ 2.59 ″ E], leg. G. Go- ergen, coll. IITA; 1 ♂, 2. ii. 2014, Niaouli, forest area [06 ° 44 ’ 3.08 ″ N 02 ° 8 ’ 2.59 ″ E], sweepnetting, leg. G. Goergen, coll. IITA; 2 ♂♂, xi. 2016, Sérou, forest area [09 ° 40 ’ 03.00 ″ N 01 ° 41 ’ 50.00 ″ E], leg. G. Goergen, coll. IITA; DEMOCRATIC REPUBLIC OF CONGO: 1 spec- imen, 18. x. 1945, Tshuapa, Flandria, det. as P. intermedia 262 Jens-Hermann Stuke & David K. Clements by Camras 1962, leg. P. Hulstaert, coll. MRAC; 2 ♂♂, no date, Ville Malela, Chief Casende [5,40 S 23,45 E], det. as P. acroschista by Camras 2000, det. as P. intermedia by Camras 1962, leg. J. Bequaert, coll. FMNH; GHA- NA: 1 ♂, 9. ix. 1947, Kumasi, paratype of P. meii, leg. J. Bowden, coll. NHML [NHMUK 010922153]; NIGE- RIA: 2 ♂♂, xii. 1998, Ibadan, IITA, sweeping net in for- est, leg. G. Goergen, coll. IITA; 1 ♀, 30. i. 1988, Ibadan, IITA, leg. G. G. M. Schulten, coll. RMNH; 1 ♀, 3. ii. 2000, Ibadan, IITA, on mud at forest margin, leg. G. Goergen, coll. IITA; SIERRA LEONE: 1 ♂ 1 ♀, 28. xi. 1993, 5 km w Kabala, swept along road [09 ° 35 ′ N 11 ° 35 ′ W], leg. L. Cederholm, R. Danielsson & R. Hall, coll. MZLU; UGANDA: 1 ♀, 16. i. 1966, nr. Entebbe, paratype of P. meii, attacking Apis, leg. D. J. Greathead, coll. NHML [NHMUK 010922152]. Camras (1962 b, 2001) was the only author to distinguish four very similar species of the pilitarsis group: P. vitripennis (Curran, 1928), P. intermedia (Kröber, 1936) (as P. acroschista sensu Camras (2001) nec Speiser, 1911), P. bouvieri (Séguy, 1936) and P. meii Camras, 2001. He used characters of wing colouration, relative length of tarsomeres and extent of dusting on the postpronotum to segregate these, but all of these characters are variable and the four taxa intergrade. There is little consistent dif- ference in the female theca or tarsi between these taxa. The holotypes of P. intermedia and P. meii were avail- New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 263 able to us but the holotype of P. bouvieri, which should be in Muséum National d’Histoire Naturelle, Paris, but which was not included in the list of material available there (GBIF 2019), could not be investigated. These four species are herewith treated as synonyms: Physocephala vitripennis Curran, 1928 = Physocephala intermedia Kröber, 1936 (syn. nov.) = Physocephala [Conops] bouvieri Séguy, 1936 (syn. nov.) = Physocephala [Pseudophysocephala] meii Camras, 2001 (syn. nov.).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFFB267F4B92FA9E617D.taxon	description	(Figs 126 – 130)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFFB267F4B92FA9E617D.taxon	materials_examined	Material. SOUTH AFRICA: 1 ♂, 10. – 22. xii. 1930, Cape Province, Somerset East, paratype of P. barbata, det. as P. constricta by Kröber 1938, leg. R. E. Turn- er, coll. NHML [NHMUK 010922128]; ZIMBABWE: 2 ♂♂, 28. x. 1965, n Vumba [= Bvumba] [- 19.093432 32.740631], paratype of P. barbata, leg. D. Cookson, coll. FMNH. Physocephala barbata looks like P. hirta but with shorter setulae. Only the female holotype of P. hirta is known whilst only males are known of P. barbata, leading to the suspicion that they may be conspecific. It would be use- ful to find the opposite sex of one or other of these spe- cies to exclude the possibility of sexual dimorphism such as that observed in other species of the pubescens group.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFFB267F4839FC2C67B3.taxon	description	(Figs 131 – 133) Physocephala caenostylatus Kröber 1936	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFFB267F4839FC2C67B3.taxon	materials_examined	Primary type material examined. ♀ holotype of Physocephala caenostylatus Kröber, 1936: (1) “ Ph. Type ♂ / caenostylus [sic] ”; (2) “ Musée du Congo / Kibali-Ituri: Kilo / V- 1930 / G. du Soleil ”; (3) “ R. DÉT / U / 2992 ”; (4) “ Physocephala ♂ / caenostylus [sic] / Krb / det. Kröber 1935 ”; (5) “ Typus ”; (6) “ RMCA ENT / 000012178 ”; coll. MRAC. Additional material. KENYA: 1 ♀, 7. x. 2013, S. Masai Reserve, des. as syntype of P. pubescens, leg. T. J. An- derson, coll. NHML [NHMUK 010922163]; TOGO: 1 ♂, vi. 2008, Kloto, forest area [6 ° 57 ’ 31.66 ″ N 0 ° 34 › 29.75 ″ E], leg. G. Goergen, coll. IITA. Whilst identification of female P. caenostylata is straightforward males are very difficult. We are not convinced that all specimens can be safely distinguished from those of P. pubescens.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	description	(Figs 134 – 142) urn: lsid: zoobank. org: act: A 4 F 67 D 5 A- 00 A 2 - 451 D- 8 B 10 - 59 F 1 CA 272 EC 0	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	materials_examined	Holotype ♀. (1) “ Togo / Kloto / forest area / Feb. 2018 / Col.: G. Goergen ”; (2) “ Holotypus / Physocephala goergeni / spec. nov. ♀ / det. Stuke 2019 ”. Holotype is depos- ited in ZMHB. The specimen is pinned and in very good condition.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	description	Description of holotype (female) Length 10.8 mm; Wing-length 7.0 mm; Head-height 2.8 mm. 266 Jens-Hermann Stuke & David K. Clements Head. Antenna orange-brown, pedicel and tip of first flagellomere blackish-brown. Arista stylus-like, with 2 aristomeres situated at tip of first flagellomere (Fig. 137). Basal aristomere with projection about as long as api- cal aristomere. Scape about four times longer than max- imum width, apically and ventrally with setae. Pedicel about six times longer than maximum width, apically and most of dorsal surface covered with black setulae. Ped- icel lacking any ridge at base, expanded towards apex. First flagellomere long and conical, about three times as long as high, pointed, ventrally with indistinct membra- nous area. Lunule between base of antennae and ptilinal suture distinct, slightly longer than width of scape. Eye brown, lacking ommatrichia, facets all of about the same size. Posterior margin of eye with distinct shining inden- tation. Gena-height / eye-height (measurements taken from head in lateral view) = 0.1. No ocellar tubercle, no ocelli and no ocellar triangle evident. Frons (Fig. 135) longer than broad, slightly concave, not projecting above eyes posteriorly, lacking any setulae. Anterior margin of frons slightly concave. Frons yellow with broad brown midstripe. No frontofacial spot. Frons indistinctly dusted all over. Vertex as broad as frons, separated from latter by laterally distinct and medially indistinct ridge. Dense cover of black setulae on vertex forming a more or less level-topped pile in lateral view (Fig. 136). Vertex api- cally with triangular field which lacks setulae or any distinct depression. Face yellow with central brown to blackish spot. Facial grooves with indistinct dusting, nar- rowly dusted along eye margin. Gena yellow, generally lacking setae, although with several setae below postcra- nium. Distinct facial grooves reaching mouth edge, fa- cial carina reaching from base of antennae to a distinctly broadened and outstanding frontoclypeal tubercle. Ptili- nal suture stretching on either side well beneath antennal bases. Oral cavity tapers dorsally. Postcranium not obvi- ously invaginated, black dorsally and yellow ventrally. Whole postcranium slightly dusted, with no obviously denser dusting adjacent to posterior margin of eye. Oc- ciput and postgena covered with black setulae. Postgena not widened and not delimited from occiput. Bottom portion of postcranium distinctly separated-off and not setulose. Proboscis reddish-brown basally and black api- cally, labellum blackish-brown. Frontoclypeal membrane long, light orange-brown and hardly delimited from or- ange-brown clypeus. Palps absent. Labium longer than head-length in lateral view, thickened basally, anterior section completely fused into a tube. Labrum as long as labium but very narrow. Labellum short, completely di- vided, hardly broader than adjacent haustellum, and cov- ered with very short setulae. Thorax mainly black, with pleura black to brown. Postpronotum orange-brown. Thorax evenly grey-dusted all over, lacking any distinctly denser dusting or shining areas. Presternum distinct, broad. Basisternum broad, not narrowed ventrallyh to a point, lacking setae or setulae. Proepisternum also lacking setae or setulae. Scutum covered with black setae. Notopleuron and postalarcallus with several stronger setae. Postalarcallus lacking any curved setulae beneath the black setae. Katepisternum with 10 – 15 setae posterodorsally, 1 – 2 setae medially, and no setae ventrally. Metakatepisternum, anepisternum and anepimeron lacking setae or setulae. Mediotergite convex, covered with strong black setae which can be as long as maximum diameter of hind femur. Subscute- llum inconspicuous. Scutellum densely covered with black setae and with 3 outstanding larger setae on posterior margin. Wing as Fig. 142: Fore-margin of wing tinged brown, with brownish membrane between costa and media, brown basal-medial cell and slightly brown- ish hind-margin of discal-medial cell. Hyaline membrane between media and vena spuria in cell r 4 + 5. Veins brown to black. Wing completely covered with microtrichia. Radial-medial crossvein short but complete. Basal-me- New conopid records from the Afrotropical Region (Diptera) – Part 3: Physocephalini 267 dial-cubital crossvein incomplete. Radius R 1 and R 2 + 3 terminate close together in costa, well beyond end of subcosta. Radius R 4 + 5 with shallow, even curve in dis- tal section directed towards fore-edge of wing. Cell r 4 + 5 pedunculate, vein R 4 + 5 + M 1 well distinctly longer than radial-medial crossvein. Cubital cell cup elongated, lon- ger than vein A 1 + CuA 2, and pointed distally (i. e. cubitus CuA 2 and anal vein A 1 meet at an acute angle). Cubital veins CuA 1 and Basal-medial-cubital crossvein separat- ed. Upper and lower calypters yellowish-white to brown, upper calypter with black setulae on margin. Alula broad (distinctly broader than long), lacking setulae on posteri- 268 Jens-Hermann Stuke & David K. Clements or margin. Venae spuriae pronounced in cell r 4 + 5, cubital cells cup and cua 1, and discal-medial cell. Haltere white, with light brown base. Knob of haltere with black setulae. Legs orange with black to dark brown tarsi. Legs with in- conspicuous silver dusting, coxae densely silver-dusted. Posterior surfaces of fore and middle tibiae with obvious silver-dusted fields distally. Legs generally with short, adpressed black setulae. Base of fore and middle femo- ra lacking denser black setulae basally. Areas with dense black to brown setulae anteroventrally on tip of fore tibia, and ventrally and posteriorly on tip of hind tibia. Middle femur lacking a distinct row of regularly arranged setu- lae. Hind femur ventrally with several long thin setulae. No preapical setae dorsally on tibiae. No setae ventral- ly on tibiae but all tibiae dorsally with distinct line of densely-arranged small black setulae (Fig. 140). Femo- ra ventrally lacking rows of short black setae, but with indistinct lines of setulae. Coxae with several setae but none outstandingly long. Hind femur slightly thickened in basal half. Each metatarsus with 1 – 2 stronger seta ven- trally on base. Pulvilli yellowish-white. Claws brown, with broad black tips. Empodium light brown, about as long as pulvilli. Abdomen dark brown to black, with theca and later- al margins of tergites 2 – 3 orange-brown. Abdomen with dense black setulae all over, those on tergite 2 less dense and obviously longer. Abdomen entirely somewhat sil- ver, brown or golden-dusted, more strongly so at hind margins of tergites 1 – 3. Tergite 8 shining. Tergites 1 – 3 fused and hardly distinguishable from each other. Max- imum width of abdomen at segment 3. Sternites cannot be seen due the ventrally overlapping tergites. Tergite 5 and sternite 5 not completely fused laterally. Shape of theca as Figs 138 & 139. Anterior surface of the theca with long black setulae. Posterior surface apically with crescentric field of long, blunt, close-set black spicules which stand very close together, not arranged in rows.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	diagnosis	Diagnosis Physocephala goergeni is easily identified as member of the Physocephala pubescens species-group by the long setae on the mediotergite (Fig. 141). Within this group it belongs among those species which lack any setulae on the frons. The shape of the field of thick black setae on the theca distinguishes ♀ P. goergeni immediately from P. nigrita (Fig. 139 vs. Fig. 146). Additional important characters include the level-topped pile of setulae on the vertex (Fig. 136) and the shorter basal aristomere (cf Fig. 137 vs. Fig. 145).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	etymology	Etymology This species is dedicated to Georg Goergen (Ibadan, Ni- geria) who collected the holotype, and who appears to be the first entomologist to collect Conopidae in Benin and Togo.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD27FFF0267F4D40FCFD66EF.taxon	distribution	Distribution To date only the locus typicus in Togo is known. The sampling site is situated at 6 ° 57 ’ 31.66 ″ N 0 ° 34 › 29.75 ″ E.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2CFFF025D84D02FC1E61A6.taxon	description	(Figs 143 – 146) Pseudophysocephala nigrita Camras 1962	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2CFFF025D84D02FC1E61A6.taxon	materials_examined	Material. CAMEROON: 1 ♀, 17. – 19. viii. 2003, Northwest Reg., Mezam, Bafut village [06 ° 05.026 ′ N 10 ° 07.442 ′ E], 1060 m, leg. A. H. Kirk-Spriggs, coll. BMSA; KENYA: 2 ♀, 4. – 11. xii. 1999, Western Province, Kakamega Forest, Malaise trap [0 ° 14.13 ′ N 34 ° 51.87 ′ E], leg. R. Copeland, coll. NMKE; 1 ♀, 16. – 23. viii. 2000, dit- to; TOGO: 1 ♂, ii. 2017, Kloto, forest area [6 ° 57 ’ 31.66 ″ N 0 ° 34 › 29.75 ″ E], leg. G. Goergen, coll. IITA; 1 ♀, viii. 2015, ditto; UGANDA: 1 ♂, vii. – viii. 1946, Bwamba, leg. van Someren, coll. NHML [NHMUK 010922185]; 1 ♀, vii. – viii. 1946, ditto [NHMUK 010922187]; 1 ♂, 16. xii. 1934, Kampala, flying against bees, leg. T. W. Chorley, coll. NHML [NHMUK 010922191]; 1 ♂, 18. ii. 1966, Lake Nabugabo, det. as P. nigrita by Camras 2000, leg. D. J. Greathead, coll. NHML [NHMUK 010922156]. The main character given to distinguish P. brevivertex in the original description of Camras (2001) was a “ short vertex ”. This character is difficult to assess, however, be- cause the vertex may be tightly curved and of variable shape in the material to hand. Apically the vertex has an area which is not covered with setulae and which may have a depression, but the shape, size and extent of any depression can vary considerably, giving a different im- pression of the vertex shape. There is no other character available to distinguish P. brevivertex from P. nigrita, and therefore Physocephala [Pseudophysocephala] brevivertex Camras, 2001 is herewith placed as a junior synonym of Physocephala [Pseudophysocephala] nigrita Camras, 1962 (syn. nov.).	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2CFFF025D84F8EFEB96472.taxon	materials_examined	Primary type material examined. ♀ holotype of Pseudophysocephala hirta Kröber 1939: (1) “ Holo- / type ”; (2) “ Type ”; (3) “ E. Cape Prov. / Katberg. / 1. – 10. ii. 1933. ”; (4) “ S. Africa. / R. E. Turner. / Brit. Mus. / 1933 - 139. ”; (5) “ Pseudophysocephala / hirta Kröber / examined & det. / O. Kröber, 1938. ”; coll. NHML See comment under Physocephala barbata (Camras, 2001), above.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2CFFF1265A4F51FC4565C6.taxon	description	(Figs 147 – 151)	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2CFFF1265A4F51FC4565C6.taxon	materials_examined	Primary type material examined. ♂ lectotype of Physocephala pubescens Brunetti 1925 herewith designated: (1) “ Syn - / type ”; (2) “ 23.9.16 / Limbe / Nyasaland. / 4000 ′ (R. C. W.) / R. b. Wood / 684. ”; (3) “ Physo. / pubescens / Brun Type ♂ / Det. E. Brunetti 1924 ”; (4) “ Lec- totypus / Physocephala / pubescens ♂ / Brunetti, 1925 / des. Stuke 2019 ”; NHML. ♀ lectotype of Physocephala curta Kröber, 1936 here- with designated: (1) “ Type / Ph. curta ♀ ”; (2) “ Musée du Congo / Elisabethville / 1927 / Dr. M. Bequaert ”; (3) “ Physocephala / curta Krb. / det. Kröber 1935 ”; (4) “ R. DÉT / L / 2992 ”; (5) “ Type ”; (6) “ RMCA ENT / 000012182 ”; coll. MRAC. Additional material. DEMOCRATIC REPUBLIC OF CONGO: 1 ♂, no date, Elisabethville [Lubumbashi] [- 11.664232 27.482626], det. as P. pubescens by Cam- ras 1962, leg. M. Bequaert, coll. FMNH; KENYA: 1 ♀, ii. 1932, Kijabe, det. as P. pubescens by Camras 1962, leg. van Someren, coll. NHML [NHMUK 010922169]; 1 ♀, 20. iii. 1949, Nairobi, det. as P. pubescens by Camras 1962, leg. G. Salt, coll. NHML [NHMUK 010922168]; 1 ♂, 9. iv. 1949, Nairobi, det. as P. pubescens by Camras, 2000, leg. G. Salt, coll. NHML [NHMUK 010922188]; 1 ♀, 3. vii. 1994, Nairobi, leg. R. Copeland, coll. NMKE; 270 Jens-Hermann Stuke & David K. Clements 1 ♂, 12. viii. 1966, Nairobi, 5400 ft, det. as P. pubescens by Smith, 1967, leg. G. R. C. van Someren, coll. NHML [NHMUK 010922186]; 1 ♂, 2. i. 1997, Nairobi, Kebete, 1800 m, leg. T. Romig, coll. PMHA; MALA- WI: 1 ♂, 17. iii. 1913, Nyasaland, Mlanje [= Mulanje], des. as syntype of P. pubescens, leg. S. A. Neave, coll. NHML [NHMUK 010922166]; 1 ♂, 6. xi. 1912, Nyas- aland, Mt. Mlanje [= Mulanje Massif], des. as syn- type of P. pubescens, leg. S. A. Neave, coll. NHML [NHMUK 010922165]; 1 specimen, 1. x. 1913, Nyas- aland, Mt. Mlanje [= Mulanje Massif], des. as syn- type of P. pubescens, leg. S. A. Neave, coll. NHML [NHMUK 010922164]; UNKNOWN LOCATION: 1 ♀, iv. 1941, “ Mtowambo ”, det. as P. pubescens by Camras 1962, leg., coll. FMNH; 1 ♀, viii. 1943, [characters illeg- ible], det. as P. pubescens by Camras 1962, leg. H. J. A. Turner, coll. NHML [NHMUK 010922167]. The material held under ‘ Physocephala pubescens ‘ at the NHML includes four syntypes of P. pubescens and two syntypes of P. curta. The former comprises a mixture of P. pubescens and P. caenostylata Kröber, 1936. It is therefore necessary to designate a lectotype for P. pubescens and we herewith select a male syntype from Limbe which agrees with the species concept as here defined in order to prevent any change in the past usage of this name. Stuke (2017 a) incorrectly stated that Smith (1980) had synonymized P. curta with P. pubescens when in fact this was done by Camras (1962). Re-examination of the two P. curta syntypes held at NHML concluded that these also comprise multiple species. The female syntype from Elisabethville is identical with P. pubescens, and in order to prevent further confusion we herewith designate this specimen as the lectotype of P. curta. The second syn- type belongs to P. microvena Brunetti, 1925 and bears the labels: (1) “ Paratype / Ph. curta ♀ ”; (2) “ Musée du Congo / Ituri: Blukwa / 10 - XI - 1928 / A. Collart ”; (3) “ Cotype ”; (4) “ Physocephala ♀ / curta Krb. / det. Kröber 1935 ”; (5) “ R. DÉT / M / 2992 ”; coll. MRAC. This spec- imen has, unusually, a distinctly developed radial-medial crossvein and could therefore easily be confused with other species.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
173187E9DD2FFFF325F24A6DFD6463AA.taxon	materials_examined	Material. ERITREA: 1 ♀, 28. ix. 1957, Taramna, det. as P. maculigera by Camras 2000, leg. D. J. Greathead, coll. NHML [NHMUK 010922106]; ETHIOPIA: 1 ♀, 3. iv. 2016, SNNPS State, Arba-Minch, Dorze [06 ° 10 ′ N 37 ° 35 ′ E], 2340 m, leg. J. Halada, coll. CULSP.	en	Stuke, Jens-Hermann, Clements, David K. (2019): New conopid records from the Afrotropical Region (Diptera) - Part 3: Physocephalini. Bonn zoological Bulletin 68 (2): 209-273, DOI: 10.20363/BZB-2019.68.2.209, URL: https://www.mendeley.com/catalogue/2d30fdc3-47e4-355c-b575-0cbcb9209acd/
