taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
1533B37EFFA2E14FD8E6F9C98169FE2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006663/files/figure.png	https://doi.org/10.5281/zenodo.17006663	Figure 2. Habitus of Z. affine (Simon, 1870). A–B, lectotype, male (MNHN-AR1521).C–D, paralectotype, female (MNHN-AR1523). A, C, dorsal view. B, D, ventral view. Scale bar = 0.4 mm.	Figure 2. Habitus of Z. affine (Simon, 1870). A–B, lectotype, male (MNHN-AR1521).C–D, paralectotype, female (MNHN-AR1523). A, C, dorsal view. B, D, ventral view. Scale bar = 0.4 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA2E14FD8E6F9C98169FE2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006723/files/figure.png	https://doi.org/10.5281/zenodo.17006723	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA2E14FD8E6F9C98169FE2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA2E14FD8E6F9C98169FE2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA2E14FD8E6F9C98169FE2B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA3E14CD8E9FB698050FCF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006665/files/figure.png	https://doi.org/10.5281/zenodo.17006665	Figure 3. Habitus of Z. beticum Denis, 1957. A–B, male (CMU). C–D, female (CJFM-18427, G7572.14658). A, C, dorsal view. B, D, ventral view. Scale bar = 0.4 mm.	Figure 3. Habitus of Z. beticum Denis, 1957. A–B, male (CMU). C–D, female (CJFM-18427, G7572.14658). A, C, dorsal view. B, D, ventral view. Scale bar = 0.4 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA3E14CD8E9FB698050FCF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA3E14CD8E9FB698050FCF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA3E14CD8E9FB698050FCF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA3E14CD8E9FB698050FCF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA0E14DD8F4FBED86AEFED0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006669/files/figure.png	https://doi.org/10.5281/zenodo.17006669	Figure 4. Habitus of Z. diatretum Denis, 1935. A–B, lectotype, male (MNHN-AR1513). C–D, paralectotype, female (MNHN-AR1513). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 4. Habitus of Z. diatretum Denis, 1935. A–B, lectotype, male (MNHN-AR1513). C–D, paralectotype, female (MNHN-AR1513). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA0E14DD8F4FBED86AEFED0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA0E14DD8F4FBED86AEFED0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA0E14DD8F4FBED86AEFED0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA1E14DD8F5FDC483D0F96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006671/files/figure.png	https://doi.org/10.5281/zenodo.17006671	Figure 5. Habitus of Z. elegans (Simon, 1873). A–B, specimen from type series, male (MNHN-AR2853). C–D, specimen from type series, female (MNHN-AR2853). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 5. Habitus of Z. elegans (Simon, 1873). A–B, specimen from type series, male (MNHN-AR2853). C–D, specimen from type series, female (MNHN-AR2853). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA1E14DD8F5FDC483D0F96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA1E14DD8F5FDC483D0F96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA1E14DD8F5FDC483D0F96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA1E14DD8F5FDC483D0F96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006675/files/figure.png	https://doi.org/10.5281/zenodo.17006675	Figure 6. Habitus of Z. maculatum (Simon, 1870). A–B, male (CMU). C–D, lectotype, female (MNHN- AR1512). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 6. Habitus of Z. maculatum (Simon, 1870). A–B, male (CMU). C–D, lectotype, female (MNHN- AR1512). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006679/files/figure.png	https://doi.org/10.5281/zenodo.17006679	Figure 7. Habitus of Z. maculatum (Simon, 1870), holotype of Z. tuber, male (ZMH-A00014214). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	Figure 7. Habitus of Z. maculatum (Simon, 1870), holotype of Z. tuber, male (ZMH-A00014214). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006729/files/figure.png	https://doi.org/10.5281/zenodo.17006729	Figure 22. Male palp of Z. maculatum (holotype of Z. tuber) (ZMH-A00014214). A, ventral view. B, retrolateral view. Scale bar = 0.2 mm.	Figure 22. Male palp of Z. maculatum (holotype of Z. tuber) (ZMH-A00014214). A, ventral view. B, retrolateral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAEE146D8CAFAC98085FB53.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006681/files/figure.png	https://doi.org/10.5281/zenodo.17006681	Figure 8. Habitus of Z. mallorca Bosmans, 1994. A–B, paratype, male (CJFM-4583, G7572.2929). C–D, paratype, female (CJFM-4583, G7572.2929). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 8. Habitus of Z. mallorca Bosmans, 1994. A–B, paratype, male (CJFM-4583, G7572.2929). C–D, paratype, female (CJFM-4583, G7572.2929). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006723/files/figure.png	https://doi.org/10.5281/zenodo.17006723	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006691/files/figure.png	https://doi.org/10.5281/zenodo.17006691	Figure 11. Habitus of Z. pseudoelegans Denis, 1934. A–B, lectotype, male (MNHN-AR9910). C–D, paralectotype, female (MNHN-AR9910). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 11. Habitus of Z. pseudoelegans Denis, 1934. A–B, lectotype, male (MNHN-AR9910). C–D, paralectotype, female (MNHN-AR9910). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFAAE147D8C2FAD583CBFD8B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFABE144D8C9FD1F8317FD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006685/files/figure.png	https://doi.org/10.5281/zenodo.17006685	Figure 9. Habitus of Z. marginiceps Simon, 1914. A–B, lectotype, male (MNHN-AR9900, 22549). C–D, paralectotype,female (MNHN-AR9900, 22549).A, C, dorsal view.B, D, ventral view. Scale bar = 0.2 mm.	Figure 9. Habitus of Z. marginiceps Simon, 1914. A–B, lectotype, male (MNHN-AR9900, 22549). C–D, paralectotype,female (MNHN-AR9900, 22549).A, C, dorsal view.B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFABE144D8C9FD1F8317FD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006723/files/figure.png	https://doi.org/10.5281/zenodo.17006723	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFABE144D8C9FD1F8317FD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFABE144D8C9FD1F8317FD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFABE144D8C9FD1F8317FD78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA8E145D8DEFCFE866AFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006687/files/figure.png	https://doi.org/10.5281/zenodo.17006687	Figure 10. Habitus of Z. murphyorum Bosmans, 1994. A–B, paratype, male (CJFM-18452, G7572.14697). C–D, paratype, female (CJFM-18452, G7572.14697). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 10. Habitus of Z. murphyorum Bosmans, 1994. A–B, paratype, male (CJFM-18452, G7572.14697). C–D, paratype, female (CJFM-18452, G7572.14697). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA8E145D8DEFCFE866AFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA8E145D8DEFCFE866AFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA8E145D8DEFCFE866AFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA8E145D8DEFCFE866AFDAF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA9E15AD8D4FD478317FC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006691/files/figure.png	https://doi.org/10.5281/zenodo.17006691	Figure 11. Habitus of Z. pseudoelegans Denis, 1934. A–B, lectotype, male (MNHN-AR9910). C–D, paralectotype, female (MNHN-AR9910). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 11. Habitus of Z. pseudoelegans Denis, 1934. A–B, lectotype, male (MNHN-AR9910). C–D, paralectotype, female (MNHN-AR9910). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA9E15AD8D4FD478317FC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006723/files/figure.png	https://doi.org/10.5281/zenodo.17006723	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA9E15AD8D4FD478317FC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA9E15AD8D4FD478317FC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFA9E15AD8D4FD478317FC45.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB6E15BD867FBC880A3FB67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006695/files/figure.png	https://doi.org/10.5281/zenodo.17006695	Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	Figure 12. Habitus of Z. sanchezi Shafaie and Pekár sp. n. A–B, holotype, male (NMP). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB6E15BD867FBC880A3FB67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006731/files/figure.png	https://doi.org/10.5281/zenodo.17006731	Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis.	Figure 23. Male palp of Z. sanchezi Shafaie and Pekár sp. n., holotype (NMP). A, D, prolateral view.B, E, ventral view. C, F, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; RTA = retrolateral tibial apophysis; Tm = mesal tubercle of the tibial apophysis; Tr = retrolateral tubercle of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB6E15BD867FBC880A3FB67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB6E15BD867FBC880A3FB67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006669/files/figure.png	https://doi.org/10.5281/zenodo.17006669	Figure 4. Habitus of Z. diatretum Denis, 1935. A–B, lectotype, male (MNHN-AR1513). C–D, paralectotype, female (MNHN-AR1513). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 4. Habitus of Z. diatretum Denis, 1935. A–B, lectotype, male (MNHN-AR1513). C–D, paralectotype, female (MNHN-AR1513). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB7E158D84EFAE083C1FC90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006697/files/figure.png	https://doi.org/10.5281/zenodo.17006697	Figure 13. Habitus of Z. valentii Bosmans, Loverre and Addante, 2019. A–B, male (CMU). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 13. Habitus of Z. valentii Bosmans, Loverre and Addante, 2019. A–B, male (CMU). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB7E158D84EFAE083C1FC90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006723/files/figure.png	https://doi.org/10.5281/zenodo.17006723	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	Figure 20. Comparison of palp shape among males of the elegans group. A–C, Z. affine (MNHN- AR1521). D–F, Z.valentii (CMU). G–I, Z. marginiceps (MNHN-AR9900, 22549). J–L, Z. mallorca (CJFM- 4583, G7572.2929). M–O, Z. pseudoelegans (MNHN-AR9910). A, D, G, J, M, prolateral view. B, E, H, K, N, ventral view. C, F, I, L, O, retrolateral view. Scale bar = 0.2 mm. Abbreviations: C = conductor; Em = embolus; Es = subterminal tooth of embolus; MA = median apophysis; Rt = retrolateral tooth of the tibial apophysis; RTA = retrolateral tibial apophysis; T = tegulum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB7E158D84EFAE083C1FC90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB7E158D84EFAE083C1FC90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB7E158D84EFAE083C1FC90.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB4E159D8C6FC9783CBFF67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006701/files/figure.png	https://doi.org/10.5281/zenodo.17006701	Figure 14. Habitus of Z. vanimpei Bosmans,1994. A–B, paratype, male (CJFM-18593, G7572.15281). C– D, female (CJFM-18557, G7572.15251). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 14. Habitus of Z. vanimpei Bosmans,1994. A–B, paratype, male (CJFM-18593, G7572.15281). C– D, female (CJFM-18557, G7572.15251). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB4E159D8C6FC9783CBFF67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006725/files/figure.png	https://doi.org/10.5281/zenodo.17006725	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	Figure 21. Comparison of palp shape among males of the elegans group. A–C, Z. beticum (CMU). D–F, Z. diatretum (MNHN-AR1513). G–I, Z. maculatum (CMU). J–L, Z. murphyorum (CJFM-18452, G7572.14697). M–O, Z. vanimpei (CJFM-18593, G7572.15281). P–R, Z. elegans (MNHN-AR2853). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviations: Bi = baso-lateral incision of the median apophysis; It = internal tooth of the tibial apophysis; TA = terminal apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB4E159D8C6FC9783CBFF67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB4E159D8C6FC9783CBFF67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB4E159D8C6FC9783CBFF67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB5E15FD8CCFCB183BAFA3C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006703/files/figure.png	https://doi.org/10.5281/zenodo.17006703	Figure 15. Habitus of Z. isabellinum (Simon, 1870). A–B, paralectotype, male (MNHN-AR1522). C–D, paralectotype, female (MNHN-AR1522). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 15. Habitus of Z. isabellinum (Simon, 1870). A–B, paralectotype, male (MNHN-AR1522). C–D, paralectotype, female (MNHN-AR1522). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB5E15FD8CCFCB183BAFA3C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006733/files/figure.png	https://doi.org/10.5281/zenodo.17006733	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB5E15FD8CCFCB183BAFA3C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB5E15FD8CCFCB183BAFA3C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB5E15FD8CCFCB183BAFA3C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB3E150D8D1FA2A8363FA67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006707/files/figure.png	https://doi.org/10.5281/zenodo.17006707	Figure 16. Habitus of Z. italicum (Canestrini, 1868). A–B, male (MNHN-AR2866). C–D, female (MNHN- AR2866). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 16. Habitus of Z. italicum (Canestrini, 1868). A–B, male (MNHN-AR2866). C–D, female (MNHN- AR2866). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB3E150D8D1FA2A8363FA67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006733/files/figure.png	https://doi.org/10.5281/zenodo.17006733	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB3E150D8D1FA2A8363FA67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB3E150D8D1FA2A8363FA67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFB3E150D8D1FA2A8363FA67.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFBCE16AD8C7F9E48009FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006709/files/figure.png	https://doi.org/10.5281/zenodo.17006709	Figure 17. Habitus of Z. modestum (Simon, 1870). A–B, lectotype, male (MNHN-AR1519). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 17. Habitus of Z. modestum (Simon, 1870). A–B, lectotype, male (MNHN-AR1519). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFBCE16AD8C7F9E48009FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006733/files/figure.png	https://doi.org/10.5281/zenodo.17006733	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFBCE16AD8C7F9E48009FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFBCE16AD8C7F9E48009FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFFBCE16AD8C7F9E48009FC6B.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF86E16ED8E8FBED83A2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006717/files/figure.png	https://doi.org/10.5281/zenodo.17006717	Figure 18. Habitus of Z. nesiotes Denis, 1965. A–B, holotype, female (MNHN-AR9949, 22556). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	Figure 18. Habitus of Z. nesiotes Denis, 1965. A–B, holotype, female (MNHN-AR9949, 22556). A, dorsal view. B, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF86E16ED8E8FBED83A2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006733/files/figure.png	https://doi.org/10.5281/zenodo.17006733	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF86E16ED8E8FBED83A2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF86E16ED8E8FBED83A2FB17.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF82E16DDB1AFB01838EF96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006719/files/figure.png	https://doi.org/10.5281/zenodo.17006719	Figure 19. Habitus of Z. cesari Pekár, A–B, male (CMU). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	Figure 19. Habitus of Z. cesari Pekár, A–B, male (CMU). C–D, female (CMU). A, C, dorsal view. B, D, ventral view. Scale bar = 0.2 mm.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF82E16DDB1AFB01838EF96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006733/files/figure.png	https://doi.org/10.5281/zenodo.17006733	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	Figure 24. Comparison of palp shape among males belonging to the italicum and pusio groups. A–O, italicum group members. A–C, Z. isabellinum (MNHN-AR1522). D–F, Z. italicum (MNHN-AR2866). G–I, Z. modestum (MNHN-AR1519). J–L, Z. nesiotes (CMU). M–O, pusio group member, Z. cesari (CMU). A, D, G, J, M, P, prolateral view. B, E, H, K, N, Q, ventral view. C, F, I, L, O, R, retrolateral view. Scale bar = 0.2 mm. Abbreviation: B = base of embolus. Arrow indicates the tooth at the base of the tibial apophysis.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF82E16DDB1AFB01838EF96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006735/files/figure.png	https://doi.org/10.5281/zenodo.17006735	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	Figure 25. Comparison of epigyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: Ep = Epigynal plate; Ms = median septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF82E16DDB1AFB01838EF96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006739/files/figure.png	https://doi.org/10.5281/zenodo.17006739	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	Figure 26. Comparison of endogyne shape among females of the elegans, italicum, and pusio groups in ventral view. A–K elegans group members. A, Z. affine (MNHN-AR1523). B, Z. valentii (CMU). C, Z. marginiceps (MNHN-AR9900, 22549). D, Z. mallorca (CJFM-4583, G7572.2929). E, Z. pseudoelegans (MNHN-AR9910). F, Z. beticum (MNHN-AR9950). G, Z. diatretum (MNHN-AR1513). H, Z. maculatum (MNHN-AR1512). I, Z. murphyorum (CJFM-18452, G7572.14697). J, Z. vanimpei (CJFM-18557, G7572.15251). K, Z. elegans (MNHN-AR2853). L–O italicum group members. L, Z. isabellinum (MNHN- AR1522). M, Z. italicum (MNHN-AR2866). N, Z. modestum (CMU). O, Z. nesiotes (MNHN-AR9949, 22556). P, pusio group member, Z. cesari (CMU). Scale bar = 0.2 mm. Abbreviations: A = atrium; Sp = spermatheca.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
1533B37EFF82E16DDB1AFB01838EF96C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/17006741/files/figure.png	https://doi.org/10.5281/zenodo.17006741	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	Figure 27. Scanning electron microscopy (SEM) images of the epigyne among females of the elegans, italicum, and pusio groups, in ventral view. A–H elegans group members. A, Z. affine (CMU). B, Z. valentii (CMU). C, Z. marginiceps (CMU).D, Z. pseudoelegans (CMU).E, Z. beticum (CMU).F, Z.maculatum (CMU). G, Z. murphyorum (CMU). H, Z. elegans (MNHN-AR2853). I–K italicum group members. I, Z. isabellinum (CMU). J, Z. italicum (CMU). K, Z. modestum (CMU). L, pusio group member, Z. cesari (CMU). Scale bar = 40 µm. Abbreviations: CO = copulatory opening; Ei = incision of the epigynal plate; Ep = epigynal plate; Ml = midline suture of the septum; P = pit; Pb = posterior borders of the copulatory openings; Se = septum; Si = incision of the septum.	2025-06-05	Shafaie, Sepideh;Pekár, Stano;Ortiz, David		Zenodo	biologists	Shafaie, Sepideh;Pekár, Stano;Ortiz, David			
