taxonID	type	description	language	source
277E1331FFAFFFA5FF4326F9FA2C1750.taxon	type_taxon	Type species. Mutilla erronea André, 1900, ♂, by original designation (junior subjective synonym of Mutilla curtiventris André, 1901 according to Pagliano & Strumia (2007: 69), resurrected to valid species by Lelej & Yildirim (2009: 15).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAFFFA5FF4326F9FA2C1750.taxon	diagnosis	Diagnosis. MALE. Inner eye margin with weak notch. Mandible strongly widened apically, quadridentate, rarely tridentate, beneath with large subbasal tooth; preapical inner teeth equal to apical one or larger than it. Clypeus concave, anterior border usually with protruding medial part. Stigmatic cell 1.4 × distance between base of stigmatic cell and origin of base RS on Sc. T 2 with long lateral felt lines, S 2 with short (reduced) lateral felt lines. FEMALE. Head not widened posterad, posteriorly rounded. Mandible tridentate. Meso- and metafemur with two row of well developed setae. Pronotum with protruding humeral part, distinctly wider than propodeum. Scutellar scale more or less developed. T 2 with one or three basal spots of pale setae, rarely with basal band of pale setae. Pygidial area widely triangular, carinated laterally, surface with divergent striae.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAFFFA5FF4326F9FA2C1750.taxon	description	Sex association. The female of the type species was associated and identified (as Mutilla curtiventris André, 1901: 269) by Pagliano & Strumia 2007: 69; and collected in copula in Turkey (Lelej & Yildirim 2009). Species-groups included. ErrОПеа, dichroa and saharica species-groups are proposed here. For diagnosis see the key to the species below.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAFFFA5FF4326F9FA2C1750.taxon	discussion	Remarks. All but two of the species are dull brown in color and apparently nocturnal in their behavior. These nocturnal species are somewhat commonly encountered in arid habitats from North Africa east to India (Lelej 2002, 2005). The more colorful diurnal species occur mainly in the Mediterranean (Lelej 2002). Lelej’s (1985) key includes about half of the currently recognized species.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAAFFA0FF432146FB031233.taxon	description	(Figs 1 – 10, 16 – 19)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAAFFA0FF432146FB031233.taxon	diagnosis	Diagnosis. MALE. Ocelli small, OOD 6 – 7 × diameter of lateral ocellus. Head black. Wings dark. Mesosoma ferruginous-red with blackish mesopleuron beneath. Metasoma with pale band on T 3 – 4 and pale apical fringe on T 1 – 2. T 7 apically rounded. Body length 9.0 – 14.0 mm. FEMALE. T 2 with one median subbasal spot of pale setae. Mesosoma ferruginous to brownish red, with sparse golden or argent setae dorsally. Pygidial area much narrow. Longitudinal eye diameter less than 2.0 × distance from posterior edge of eye to posterior edge of vertex. Body length 7.0 – 9.0 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAAFFA0FF432146FB031233.taxon	materials_examined	Material examined. (87 ♂, 36 ♀). Type material. Paratype of Smicromyrme erronea cretica Nonveiller, 1972, ♂, Greece, Iraclion, Creta, 25. IX. 1959, J. Stančić [IBSS]. Additional material. 10 ♂ and 4 ♀ from Greece (Lelej et al. 2003); 3 ♂ and 3 ♀ from Turkey (Lelej & Yildirim 2009). 9 ♂ from Azerbaijan [IBSS] (Lelej et al. 2022). Greece: Crete, 1 ♂ [ZIN]; Rhodos, Ixia, 1 – 4. IX. 1962, 2 ♂, PMF Verhoeff // Smicromyrme erronea André, J. Suárez det., 1963 [IBSS]; Hellas, Peloponisos, 5 km S Monemvasia, 26 – 31. VIII. 1983, 1 ♂, Zool. Mus. Copenh. Exped. // Smicromyrme erronea André, B. Petersen det., 1984 [IBSS]; Hellas, Lakonia, 5 km S Monemvasia, 22. VIII. 1983, 1 ♂, G. Chistensen // Smicromyrme erronea André, B. Petersen det., 1984 [IBSS]. Turkey: Prov. Cankiri, Ilgaz Dagi, Ilgaz, ca. 900 m, 13. VII. 1995, 1 ♂, Gelbrecht & E. Schwabe [IBSS]. Serbia: Preševo, Reljani, 18. VII. [19] 56, Bjegović / Vučji Kamen / Smicromyrme errana Nonvll. det. Nonvll., 1976, 1 ♀ [IBSS]; Macedonia [North Macedonia], Dojran, 9. IX. [19] 56, 1 ♀, Nonvll. [Nonveiller] [IBSS]. Greece: Corfu [Kerkira], 1867, 8 ♂, J. Erber [ZIN]; Syra [Syros, Siros], 1867, 4 ♂, J. Erber [ZIN]; 1 ♂ and 4 ♀ from Volos and Poros [NHME, IBSS] (Lelej et al. 2003); Thessaloniki, Sindos, 11. IX. [19] 57, 1 ♀, Jovanović [IBSS]; Hellas, Timfi, Óros, Aristi / Papingo, 900 - 1100 m, 24. VIII. 1983, 1 ♀, Zool. Mus. Copenh. Exped. / Smicromyrme errana Nonv., B. Petersen, 1984 [IBSS]. Bulgaria: 2 ♂ [NHME] (Lelej et al. 2003); Melnik, 16 – 22. VI. 1987, 2 ♂, H. Halada [OLML]; Melnik, 29 – 31. VII. 1982, 1 ♀, H. Halada [OLML]. Turkey: 16 ♂ and 20 ♀ including five pairs taken in copula by E. Yildirim [EMET, IBSS] (Lelej & Yildirim 2009); KasimpaŞa [Istanbul], 1 ♂, Morawitz coll. [ZIN]. Azerbaijan: Kangarli, Chalkhangala, 39 º 25 ’ N, 45 º 13 ’ E, 1445 m, 17. VI. 2020, 1 ♂, M. Maharramov [IBSS]. Tash-Bulag, [North Azerbaijan, Shekinsky Region], 19. VI. 1928, 1 ♂, O. Botsharnikov [ZIN]; Ordubad [Azerbaijan, Nakhchivan], 1892, 1 ♂, E. Reitter [ZIN]; Nakhchivan, Karabaglar, 17. VI. 1960, 1 ♂, G. Dlussky [ZMMU]; same place, 19. VI. 1959, 2 ♂, G. Viktorov [ZMMU]; Arafsa, 29. VI. 1958, 1 ♂, L. Zimina [ZMMU]. Armenia: Garni, 22. VI. 1960, 2 ♂, G. Dlussky [ZMMU]; Khosrov Reserve, 29. VI – 8. VII. 1983, 9 ♂, M. Nesterov [IBSS]; 23. VI. 1980, 2 ♂, V. Ermolenko [IBSS]; Jrvezh, 5. VII. 1976, 2 ♂, V. Gorbatovsky [IBSS]; same place, 1. VII. 1959, 1 ♂, G. Viktorov [ZMMU]; Chimankend, 22. VII. 1960, 1 ♂, E. Antonova [ZMMU].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAAFFA0FF432146FB031233.taxon	distribution	Distribution. Albania, Azerbaijan, Armenia, Bulgaria, Greece, Serbia, North Macedonia, Italy (including Sicily), Turkey, Algeria.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFAAFFA0FF432146FB031233.taxon	discussion	Remarks. In the key to the subgenera, André (1900: 134) wrote: “ Yeux étroitement et plus ou moins profondément échancrés en dedans, très rarement avec une échancrure indistincte [Chez la Mutilla erronea André] ”. According to Articles 11 and 12.1 of the ICZN (1999), Mutilla erronea André, 1900 is an available specific name. A full description of this species is given on pages 360 – 362 [dated 1902]. For a long time I followed the synonymy of Smicromyrme curtiventris (André, 1901) (= S. errana Nonveiller, 1958) proposed by B. Petersen (1988). When I examined the holotype of Mutilla curtiventris André, 1901 in the NHMW in 2011, I understood that this synonymy is wrong and M. curtiventris belongs to the genus Skorikovia Ovtchinnikov, 2002, where it becomes a new junior subjective synonym of S. trinotata (Costa, 1858). Such a form with a shortened metasoma is already known for this species (S. pouzdranensis Hoffer, 1936).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA8FFA1FF4326AFFD60130B.taxon	description	(Figs 11 – 15, 20 – 23)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA8FFA1FF4326AFFD60130B.taxon	diagnosis	Diagnosis. MALE. Ocelli small, OOD 6 – 7 × diameter of lateral ocellus. Head black. Wings dark. Mesosoma black with ferruginous-red dorsum or totally black. Metasoma with pale band on T 3 and pale apical fringe on T 1 – 2. T 7 apically straight. Body length 9.5 – 12.0 mm. FEMALE. T 2 with basal medial spot of pale setae. T 4 – 5 with black setae. Mesosoma black or dark brown, dorsum dark red with dense yellowish-white setae. Pygidial area very wide. Body length 9.2 – 12.0 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA8FFA1FF4326AFFD60130B.taxon	materials_examined	Material examined. (22 ♂, 10 ♀). Greece: 2 ♂, 5 ♀ (Lelej et al. 2003). Turkey: 2 ♂ (Lelej & Yildirim 2009). Azerbaijan: 3 ♂, 4 ♀ (Lelej et al. 2022). Armenia: Arazdayan, 15. VII. 1961, 10 ♂, 1 ♀, D. Panfilov [ZMMU]; Asni, 28. VI. 1971, 3 ♂, V. Rikhter [IBSS]. Jordan: Wadi el Heidan, 8. V. 1995, 1 ♂, K. Deneš [OLML]. Palestine: 5 km W Jericho, Wadi Qelet St. Georg, 30 ° 57 ʹN 35 ° 08 ʹE, 1 ♂, Ch. Schmid-Egger [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA8FFA1FF4326AFFD60130B.taxon	distribution	Distribution. Azerbaijan, Armenia, Greece, Iran, Turkey, Jordan (new record), Palestine.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA8FFA1FF4326AFFD60130B.taxon	discussion	Remarks. The morphological characters of Dentilla ursina (Nagy, 1972), ♀, especially the wide pygidial area and the shape of the head, as well as the coloration of the mesosoma, are similar to those of Dentilla persica, ♀, and I consider the former to be a synonym of the latter.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA9FFAFFF4323E1FD5C1553.taxon	description	(Figs 24 – 28, 33 – 35)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA9FFAFFF4323E1FD5C1553.taxon	diagnosis	Diagnosis. MALE. Ocelli large, OOD 2 × diameter of lateral ocellus. Head reddish-brown. Wings hyaline with darkened apex. Anterior border of clypeus with narrow medial projection whose dorsal length is about equal to its width. Basal half of T 2 with dense separate punctures. POD: OOD ratio 0.8 × and more, ocelli large. Upper carina of mandible high. Longitudinal carina between antennal tubercles and base of clypeus well developed. First subbasal tooth of the mandible with short carina much less than distance from the base of this carina to the base of the mandible. T 7 apically with weak notch. T 1 length less than its maximal width. Lower subbasal lobe of mandible invaginated below, height of subbasal tooth more than smallest distance between notch and upper carina of mandible. Body length 9.5 – 13.0 mm. FEMALE. T 2 with basal medial spot of pale setae. T 4 – 5 with pale setae. Eye located about same distance from articulation of mandible and posterior margin of vertex. Ratio of smallest distance between eyes to longitudinal eye diameter 1.6 – 1.8 ×. Length T 2 0.8 × its maximal width. Body length 9.0 – 10.0 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA9FFAFFF4323E1FD5C1553.taxon	materials_examined	Material examined. (157 ♂ 3 ♀). Type material. Holotype of Ephutomma quinquedentata mesopotamica Bischoff, 1920, ♂. Additional material. Turkmenistan: Dus-olum [south-western Turkmenistan], J. Pomerantsev / к. Моравица / 5 - dentata F. Mor. 1 ♂ " [ZIN]; Peskimar, D. Glasunov [1893], 1 ♂ [ZIN]; Kopet-Dag, Firyuza, 8. VI. 1903, K. Anger, 3 ♂ [ZIN]; 8, 20. VII. 1991, on light, V. Dubatolov, 2 ♂ [IBSS]; East Kopet-Dag, Charlyk, 37 ° 09 ʹ 45 ʺN 59 ° 49 ʹ 22 ʺE, 26 – 27. VI. 1991, V. Krivokhatsky, 1 ♂ [IBSS]; Bairam-Ali, 27. IX. 1896, K. Anger, 1 ♂ [ZIN]. Afghanistan: (Uruzgan, Kabul), 2 ♂ (Lelej & Kabakov 1980). Herat, Zulfagar [Sulfagar], [28. V. 1893] D. Glasunov / к. Моравица / 5 - dentata F. Mor. 3 ♂ " [ZIN]. Iran: (Hormozgan, Fars, Ilam), 100 ♂ (Lelej & Osten 2004; Lelej et al. 2008) Tehran, 14. VIII. 1951, N. Shutova, 1 ♂ [ZIN]. Razavi Khorasan Prov., Nerduali, [1893], D. Glasunov / к. Моравица / 5 - dentata F. Mor. 2 ♂ " [ZIN]; South Khorasan Prov., Nehbandan, 24. VI – 2. VII. 1901, N. Zarudny, 1 ♂ [ZIN]; Yazd Prov., ca 23 km SW Kharanaq, 1600 m, on light, 32 ° 11 ʹN 54 ° 30 ʹE, A. Timokhov, 1 ♂ [IBSS]. Kerman Prov., 75 km S Sirjan, 9. IV. 1970, Ressel, 1 ♀ [ZSM]; Keshi [Keshit], 28. III – 2. IV. 1901, N. Zarudny, 3 ♂ [ZIN]; 25 km NE Jiroft, 5 km E Saghdar, 2000 – 2400 m, on light, 28 ° 50 ʹN 57 ° 56 ʹE, A. Timokhov, 15 ♂ [IBSS]. Fars Prov., 50 km NNE of Shiraz, Bamoo Res., 29 ° 45 ʹN 52 ° 46 ʹE, 18 – 28. V. 2000, Yu. Marusik, 2 ♂ 1 ♀ [IBSS]; Sistan & Baluchestan, Bampur, 14. V. 1901, N. Zarudny, 1 ♂ [ZIN]; Karavandar [Karvandar], 23 – 24. IV. 1901, N. Zarudny, 4 ♂ [ZIN]; Sargad, 29 – 31. VIII. 1898, N. Zarudny, 2 ♂ [ZIN]; Sargad, Chaashen-Sadk, 28 – 30. IV. 1901, N. Zarudny, 1 ♂ [ZIN]; Shurab [Sor Ab], 27. VI. 1898, N. Zarudny, 1 ♂ [ZIN]; Bazman, 28. VI – 3. VII. 1898, N. Zarudny, 3 ♂ [ZIN]; Makran, Rong River, 29. III. 1901, N. Zarudny, 1 ♂ [ZIN]; Khash, 20. V. 1955, D. Steinberg, 1 ♀ [ZIN]. Israel: 50 km SE Beer Sheva, Wadi En Aqrabbin, Negev, 30 ° 57 ʹN 35 ° 08 ʹE, 8. V. 1996, Ch. Schmid-Egger, 1 ♂ [IBSS]. Jordan: Jemini, 19. XI. 1966, coll. G. Pagliano, 1 ♂ [IBSS]. Oman: NE, S of Sur, 15 km W of Al Askharah, 13. III. 2015, M. Snižek, 1 ♂ [IBSS]. Saudi Arabia: Riyadh, 1 ♂ [OLML].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA9FFAFFF4323E1FD5C1553.taxon	distribution	Distribution. Turkmenistan, Afghanistan, Iran, Iraq, Syria (new record), Jordan (new record), Israel, Palestine, Saudi Arabia (new record), Oman (new record), Egypt (Sinai).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA9FFAFFF4323E1FD5C1553.taxon	discussion	Remarks. The holotype of Мutilla quinquedentata Morawitz, 1890 from Repetek is not found in the ZIN where the Morawitz collection is deposited. This holotype is probably lost. In the ZIN collection there are 7 ♂ from Nerduali, Sulfagar, Peskimar collected by D. Glasunov in the Transcaspian region and northern Iran and Dus-olum collected by J. Pomerantsev in south-western Turkmenistan, all are with Morawitz identification label " 5 - dentata F. Mor., ♂ ". They cannot be syntypes (Lelej 1985), because they were collected by D. Glasunov in 1893 (after the description of М. quinquedentata) (Semenov 1896). Børge Petersen (unpublished) labelled the holotype of Ephutomma quinquedentata mesopotamica Bischoff, 1920, ♂ in MNHU as Dentilla dichroa (Sichel et Radoszkowski, 1869). As this synonymy has not yet been published, I hereby support it and propose it as a new synonymy. The female of D. dichroa (Sichel et Radoszkowski, 1869) may eventually be recognized as D. sabulosa (Skorikov, 1935) due to its co-occurrence in Iran: Sistan and Baluchestan Prov. and especially Fars Prov., 50 km NNE of Shiraz, Bamoo Res., 29 ° 45 ʹN 52 ° 46 ʹE, 18 – 28. V. 2000, Yu. Marusik, 1 ♀ and 2 ♂. There is no direct evidence (pair collected in copula) to support this relationship.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA7FFADFF432590FEAB12E7.taxon	description	(Fig. 63)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA7FFADFF432590FEAB12E7.taxon	diagnosis	Diagnosis. MALE. Ocelli large, OOD 2 × diameter of lateral ocellus. Head reddish-brown. Wings hyaline with darkened apex. Anterior border of clypeus with narrow medial projection whose dorsal length about equal to its width. Basal half of T 2 with dense separate punctures. POD: OOD ratio 0.8 × and more, ocelli large. Upper carina of mandible high. Longitudinal carina between antennal tubercles and base of clypeus well developed. First subbasal tooth of mandible with short carina much less than distance from base of this carina to base of mandible. T 7 apex rounded. T 1 length more than its maximal width. Body length 7.5 – 11.0 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA7FFADFF432590FEAB12E7.taxon	materials_examined	Material examined. (66 ♂). Type material. Holotype and 18 paratypes of D. irana [ZIN]. Holotype of D. kompantsevi [ZMMU]. Additional material. Iran: Hormozgan, 7 ♂ (Lelej & Osten 2004). Fars, Estahban, 15 - 18. VI. 2010, R. Yakovlev, 9 ♂ [IBSS]; India: Rajasthan, 15 km W Jaisalmer, 24. VIII. 2004, on light, T. Osten, 4 ♂; 7 km W Jaisalmer, 22. VIII. 2004, on light, T. Osten, 2 ♂; 15 km SE Jaisalmer, 20. VIII. 2004, on light, T. Osten, 1 ♂ [IBSS, SMNS]. Pakistan: Sindh Prov., Tharparker distr., Thar desert, 1. VII. 2016, G. Lakho, 2 ♂ [IBSS]; Hyderabad distr., Miani Forest, 1. II. 2016, G. Lakho, 1 ♂ [IBSS]; Pashwar - Islamabad Road, bridge across Indus River, 33 ° 53 ʹN, 72 ° 18 ʹE, 1. IX. 2005, S. Ovtchinnikov, 1 ♂ [ZIN]; Baluchistan, 10 km W of Kach vill., 30 ° 25 ʹN, 67 ° 16 ʹE, 28. VII. 2005, S. Ovtchinnikov, 18 ♂ [ZIN]; Wam vill. Pil Forest, 2213 m, 30 ° 26 ʹN, 67 ° 26 ʹE, 29. VII- 5. VIII. 2005, S. Ovtchinnikov, 1 ♂ [ZIN]; Zijarat - Multan Road, 50 km E of Zijarat, 1486 m, 30 ° 18 ʹN, 68 ° 30 ʹE, 7. VIII. 2005, S. Ovtchinnikov, 1 ♂ [ZIN].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA7FFADFF432590FEAB12E7.taxon	distribution	Distribution. Iran (Sistan and Baluchestan, Hormozgan), India (Rajasthan) (new record), Pakistan (Sind, Baluchistan) (new record).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA7FFADFF432590FEAB12E7.taxon	discussion	Remarks. Examination of D. irana specimens from India and Pakistan shows that ratio of first abscissa of radial sector (RS 1 ʹ) to second abscissa (RS 1 ʺ) of forewing varies (0.6 – 1.5 ×) and wing venation characters of holotype of D. kompantsevi are within this variation. Therefore, I consider D. kompantsevi to be a junior subjective synonym of D. irana.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFADFF43234EFCE014E7.taxon	diagnosis	Diagnosis. MALE. POD: OOD ratio 0.5 – 0.6 ×, ocelli small. Upper carina of mandible weak, mandible almost flat. Longitudinal carina between antennal tubercles and base of clypeus scarcely visible. Body length 7.5 – 8.5 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFADFF43234EFCE014E7.taxon	materials_examined	Material examined. (5 ♂). Type material. Holotype and 1 paratype of D. beludzhistana [ZIN]. Additional material. Iran: Fars, Estahban, 15 - 18. VI. 2010, R. Yakovlev, 2 ♂ [IBSS]; Loristan, Machmudvan, 4 - 5. VI. 2020, 1 ♂, R. Yakovlev [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFADFF43234EFCE014E7.taxon	distribution	Distribution. Iran (Fars, Loristan, Sistan & Baluchestan).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	description	(Figs 29 – 32, 42 – 44)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	diagnosis	Diagnosis. MALE. Ocelli large, OOD 2 × diameter of lateral ocellus. Head reddish-brown. Wings hyaline with darkened apex or preapical area. Anterior border of clypeus with narrow medial projection whose dorsal length about equal to its width. Basal half of T 2 at least with dense separate punctures. POD: OOD ratio 0.8 × and more, ocelli large. Upper carina of mandible high. Longitudinal carina between antennal tubercles and base of clypeus well developed. First subbasal tooth of mandible with short carina approximately equal to or smaller than this tooth and much less than distance from base of this carina to base of mandible. T 7 apex with weak notch. T 1 length less than its maximal width. Lower subbasal lobe of mandible scarcely invaginated below, height of subbasal tooth less than smallest distance between notch and upper carina of mandible. Body length 7.2 – 9.9 mm. FEMALE. T 2 with basal medial spot of pale setae. T 4 – 5 with pale setae. Eyes located at about same distance from articulation of mandible and posterior margin of vertex. Ratio of smallest distance between eyes to longitudinal eye diameter 1.6 – 1.8 ×. Length T 2 0.9 – 0.95 × its maximal width. T 2 laterally strongly rounded. Pygidial area granulose at least in apical half. Mesosoma dorsally with reticulate punctures; scutellar scale small, without distinct tubercles around scutellar scale. Body length 6.4 – 7.2 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	description	Description. MALE. Body length 7.2 – 9.9 mm. Head width equal mesosoma width including tegulae. Clypeus deeply concave with narrow median process anteriorly and short acute basal median carina. Scape distinctly bicarinate beneath, upper ridge visible basally. Ocelli large, POD: OOD 0.8 ×; POD equal to maximal diameter of anterior ocellus; distance between posterior ocellus and posterior head margin 1.9 × POD. Frons with longitudinal median furrow. F 1 1.4 × its width, 2.7 × pedicel, and 0.55 × F 2, F 2 1.1 × F 3. Antennal sockets with arcuate carina. Mandible quadridentate with strong curved upper carina, with acuminated subbasal tooth beneath, first subbasal tooth of mandible with short carina approximately equal to or smaller than this tooth and much less than distance from base of this carina to base of mandible; first preapical tooth less than second preapical tooth, latter with tuft of setae beneath. Vertex sparsely punctate, genae densely punctate. Pronotal dorsum almost straight anteriorly, pronotal width 1.2 × propodeal width (on spiracle line). Metasternal process with medial carina, apically without denticles. Posterior coxae carinate inside. Tegulae slightly projecting over mesoscuto-scutellar suture, shining, glabrous, with a few punctures inside. Propodeum abrupt, reticulate with median dorsal longitudinal cell distinctly delimited posteriorly by carina interrupted medially. S 1 carinate beneath, T 1 0.85 × its maximal width; T 2 with long lateral felt lines, S 2 with very short one and approximately 3 × less than distance between felt line and posterior border of S 2; T 2 with dense large punctures, sparser on disc. T 7 apically straight or with weak notch, densely punctured. Genitalia (Fig. 32). Head, palps, mesosoma, tegulae, legs, T 1 basally, and S 1 yellowish-red. Antenna yellowish-red. Mandible yellowish-red with dark denticles. Metasoma brownish. Meso- and metatibial spurs whitish. Wings hyaline with pale brown veins, forewing (distal of cells) slightly infuscated, darker toward anterior margin. Body and legs clothed with recumbent short and scattered long erect whitish pubescence; T 2 – 6 and S 2 – 6 with apical fringes of whitish setae which denser on T 2; felt lines on T 2 and S 2 whitish. FEMALE. Body length 6.4 – 7.2 mm. Head pale red, roundish, width equal to pronotum. Surface of vertex shiny, with wide dense punctuation, sparser on occipital area, gena densely punctate Eyes oval, large, clearly protruding from head profile and strongly convex; interocular distance 1.7 × maximum orbital diameter. Clypeus with prominent upper carina, clearly visible from above, ending in shiny basal tubercle. Mandible tridentate, ferruginous, weakly curved, darkish in apical half. Antennae entirely pale reddish, with curved scape. Mesosoma pale red, subrectangular, 1.2 × than broad; pronotum just slightly arched, with sharp angles, less rounded and evanescent than posterior ones; lateral margins wavy. Mesosoma dorsally shiny, with punctuation larger and denser than head; in posterior half, interpuctual spaces very small but protruding in jagged denticles, aligned in arcuate rows. Scutellar scale large and rounded. Pleurae with slightly punctate on mesopleural area. Propodeum feebly arched, without a distinct angle between propodeal and dorsal surface of mesosoma. Legs including calcaria pale red. Metasoma brown with reddish segment 1. Width of T 1 0.45 × than T 2. T 2 sparsely and finely punctate, dorsally with variolate punctures that laterally are larger, denser, and sometime confluent. Pygidial area (Fig. 44). Pubescence on head yellowish, with recumbent yellowish short setae mostly on vertex; erect setae uniformly occurring, longer ones close to eyes and on occipital side; shorter erect setae occur also on clypeal margin and around upper carina, as well as on scapes, pedicels and F 1, Mesosoma dorsally with recumbent yellowish setae, with long erect yellowish setae on lateral and posterior margins, shorter setae occur on dorsum and on pronotal margin. T 2 with medial subbasal spot of whitish setae, with long erect scattered yellowish setae and brownish recumbent setae; posterior margin with yellowish-whitish fringe of short and recumbent setae, just slightly forward extended in middle. Pubescence of T 3 – 5 mixed between long erect and short recumbent yellowish-whitish setae. Felt lines whitish, 1.2 × longer than their distance from posterior margin of T 2.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	materials_examined	Material examined. (6 ♂ 21 ♀). Type material. Holotype, ♂, Iran, Isfahan, Kashan, 34 ° 01 ' 37.98 " ʹN 51 ° 21 ʹ 19.86 " E, 975 m, 16. IX. 2013, R. Dehghani [IBSS]. Paratypes, 5 ♂ and 21 ♀ with the same label [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	distribution	Distribution. Iran (Isfahan).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	etymology	Etymology. This species is dedicated to Rouhullah Dehghani, who studied the importance of this species for human health and natural history in central Iran and collected the type series.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA5FFAAFF43254FFD0317D0.taxon	biology_ecology	Natural history. (Dehghani et al. 2018, Kassiri et al. 2021). During 2013 – 2015, 49 individuals stung by velvet ants in the residential areas of Kashan were studied. The identification of the stinging agent was made according to the morphological speciation of the insect samples collected in the houses of the injured. Stinging complications in individuals were studied according to the clinical manifestation and the time of stinging, which was from June to September. The stinging agent was identified as velvet ants, Dentilla sp. [Dentilla dehghanii sp. nov. AL]. The first sign of the sting was a severe and sharp pain. The highest percentage of redness in individuals was 47 % on the first day, and the lowest was 2 % four or five days after the sting. Intense itching was one of the main symptoms of velvet ant stings. In the final stages of pain and itching, signs of hemolysis and bruising were observed as brown spots. Due to the high number of stings in the carpet weaving workshops, living rooms and bedrooms, it was concluded that this wasp is a domestic pest.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA3FFA8FF43277DFDE1114F.taxon	description	(Figs 39 – 41)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA3FFA8FF43277DFDE1114F.taxon	diagnosis	Diagnosis. FEMALE. T 2 with basal medial spot of pale setae. T 4 – 5 with pale setae. Eye located at about same distance from articulation of mandible and posterior margin of vertex. Ratio of smallest distance between eyes to longitudinal eye diameter 1.6 – 1.8. Length T 2 0.9 × its maximal width. T 2 laterally strongly rounded (Fig. 39). Body length 7.0 – 7.5 mm. MALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA3FFA8FF43277DFDE1114F.taxon	materials_examined	Material examined (2 ♀). Type material. Holotype and paratype of Dentilla speciosa [ZIN, IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA3FFA8FF43277DFDE1114F.taxon	distribution	Distribution. Eastern Afghanistan.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA1FFB6FF4326C3FE85114E.taxon	description	(Figs 58 – 62)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA1FFB6FF4326C3FE85114E.taxon	diagnosis	Diagnosis. MALE. This species easily differs from species of dichroa species-group by having basal half of T 2 with coarse elongate foveae mixed with longitudinal striae (at least dense separate punctures in other species), darkened apical half of fore and hind wings (at least darkened apical spot of fore wing in other species). Body length 7.2 – 12.0 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA1FFB6FF4326C3FE85114E.taxon	materials_examined	Material examined. (172 ♂). Type material. Holotype and 170 paratypes of D. rasnitsyni from United Arab Emirates and Oman [RMNH, IBSS, SMNH, NHME] (Lelej & van Harten 2011). Additional material. Oman: NE, NW of Ibri, S of Dank, 9. III. 2015, M. Snižek, 1 ♂ [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA1FFB6FF4326C3FE85114E.taxon	distribution	Distribution. United Arab Emirates, Oman.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFA1FFB6FF4326C3FE85114E.taxon	biology_ecology	Natural history. This nocturnal species inhabits mountainous areas, i. e. stony deserts, and is rarely found in sandy deserts.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF43234EFB9F1557.taxon	description	(Fig. 64)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF43234EFB9F1557.taxon	diagnosis	Diagnosis. MALE. The male can be distinguished from Palaearctic D. dichroa (Sichel et Radoszkowski 1869) in having the apex of T 7 rounded (emarginated in dichroa), in having golden felt lines on T 2 and S 2 (whitish in dichroa), in having more infuscated forewing (infuscated spot beyond marginal cell in dichroa), in having differing shape and setae of volsella (long setae of basivolsella project over apex of gonostylus in dichroa). Body length 8.0 – 12.8 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF43234EFB9F1557.taxon	materials_examined	Material examined. (27 ♂). Type material. Holotype and 16 parat ypes of D. ehrenbergi from Yemen and Saudi Arabia [ZMAN, IBSS, RMNH, ZMMU] (Lelej & van Harten 2006). Additional material. Eritrea: Massaua [Mitsiwa], 1892, 1 ♂, Ragazzi [MNHU]. Israel: Arava Valley, 40 km N Elat, Yotvata, 29 53 N 35 02 E, 3. VI. 1991, 9 ♂, Müller [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF43234EFB9F1557.taxon	distribution	Distribution. Israel (new record), Yemen, Saudi Arabia, Eritrea (new record).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF4320CDFB1212E8.taxon	description	(Fig. 54)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF4320CDFB1212E8.taxon	diagnosis	Diagnosis. FEMALE. Head as wide as mesosoma, head and mesosoma closely punctured, metasoma almost round (dorsal view) with fine and somewhat sparse punctures; mesosoma subquadrate. Head, antennae, mesosoma, legs and apical two or three metasomal segments light red; basal 1 – 3 (4) metasomal segments very dark brown, almost black. Pubescence greyish white, sparse but long, especially on metasoma. Body length 4.0 – 5.9 mm. MALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF4320CDFB1212E8.taxon	materials_examined	Material examined. No material examined.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF4320CDFB1212E8.taxon	distribution	Distribution. India (Gujarat, Maharashtra).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB7FF4320CDFB1212E8.taxon	discussion	Remarks. Børge Petersen saw the holotype of Mutilla malinka Nurse, 1903 (Petersen in litt., Dec. 12, 1980) and identified the female specimen from Bombay (India: Maharashtra, Mumbai) in the collection of NHMD as Dentilla malinka. Recently (Boutin & Vilhelsem 2024) the latter specimen has been figured.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB5FF432595FC6A1137.taxon	diagnosis	Diagnosis. MALE. POD: OOD ratio 0.8 × and more, ocelli medial. Upper carina of mandible high. Longitudinal carina between antennal tubercles and base of clypeus well developed First subbasal tooth of mandible inside with long carina approximately equal to distance from base of carina to mandibular base. Second subbasal tooth of mandible located closer to first subbasal tooth than apical tooth. Distance between lateral ocelli and occipital carina 1.73 × POD. Body length 6.9 – 12.3 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB5FF432595FC6A1137.taxon	materials_examined	Material examined. No specimens examined.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB5FF432595FC6A1137.taxon	distribution	Distribution. Yemen (Socotra, Samha).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB5FF432595FC6A1137.taxon	biology_ecology	Natural history. The species occurs from the coastal belt up to 1000 m and has been found in several different habitats, including sandy or rocky coastal plains, succulent and open deciduous shrublands of inland sheltered valleys, and Dracaena woodlands (Lo Cascio et al. 2012).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBFFFB5FF432595FC6A1137.taxon	discussion	Remarks. In the legend of the figures (Lo Cascio et al. 2012), fig. 11 belongs to D. purcharti and not to D. ehrenbergi (M. Romano per. comm.). The female of D. socotrana (holotype) and four males of D. purcharti were collected by J. Batelka (7 – 8. XI. 2010, Wadi Ayhaft, 200 m), and I am sure that they belong to the same species. There is no direct evidence (pair collected in copula) to support this relationship.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43232AFC8215D7.taxon	description	(Figs 51 – 53)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43232AFC8215D7.taxon	diagnosis	Diagnosis. FEMALE. Head, mesosoma and legs ferruginous red, metasoma black, T 2 with round medial basal spot and narrow marginal band slightly protruding in the middle, T 3 – 4 with broad band, all with pale golden setae. Scutellar scale not very distinct, hardly protruding, pygidial area with parallel striae diverging apically. Body length 7.0 mm. MALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43232AFC8215D7.taxon	materials_examined	Material examined (2 ♀). Type material. Paratype of Smicromyrme arabica Hammer, 1962: 1 ♀, Fiyush Dist [rict] near Aden [Yemen], 1915 // Ind [ian] Mus [eum], [R. B. S.] Sewell, Desert fauna // Paratype // Smicromyrme arabica, m [ihi], Type, ♀, det. Hammer " [NHMW]. Other material. Yemen: Wadi Bana, 26. X. 1992, A. van Harten, 1 ♀ [ZMAN].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43232AFC8215D7.taxon	distribution	Distribution. Yemen.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43232AFC8215D7.taxon	discussion	Remarks. Dentilla arabica may be the opposite sex of D. testacea or D. ehrenbergi, since all three species were collected from nearby localities (Lelej & van Harten 2006).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43213EFAF81334.taxon	diagnosis	Diagnosis. FEMALE. A female of Dentilla belonging to the group that includes species without spots of setae on T 2, head and mesosoma red. Body length 5.5 – 6.5 mm. MALE. Unknown (see Remarks to Dentilla purcharti).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43213EFAF81334.taxon	materials_examined	Material examined. No specimens examined.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43213EFAF81334.taxon	distribution	Distribution. Yemen (Socotra).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB5FF43213EFAF81334.taxon	biology_ecology	Natural history. Data from labels indicate a broad phenology for this species, extended at least from November to April, and a wide distribution on the island. One of the specimens was collected at Homhil (about 1000 m) in a stony area subject to intensive grazing. Other traits of its biology are still unknown (Lo Cascio et al. 2012).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB2FF4326CCFE881217.taxon	description	(Figs 55 – 57)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB2FF4326CCFE881217.taxon	diagnosis	Diagnosis. MALE. Anterior border of clypeus with broad medial projection whose dorsal length about half its width. Mandible quadridentate, below with subbasal tooth. Length of first metasomal segment less than its maximum width. OOD nearly 2.0 × diameter of lateral ocellus. Legs pale, slightly brownish. Head and mesosoma brownish red. Metasternum 4 - toothed, lateral teeth clearly higher than median teeth. Apical part of forewings darkened. First mandibular inner subbasal tooth small, sharp. Body length 8.0 – 12.5 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB2FF4326CCFE881217.taxon	materials_examined	Material examined. (4 ♂). Morocco: Assa-Zag Province, Aouinet - Torkoz, Draa, 20 – 24. X. [19] 74, J. Mateu // Eremomyrme saharica (Gin.), ♂, J. Suárez det., 1976, 2 ♂ [IBSS]; Maader Aster, Oued, Tigzirt, Draa, 26. X. 1971, J. Mateu // Eremomyrme saharica (Gin.), ♂, J. Suárez det., 1976, 1 ♂ [IBSS]. Algeria: Beni Abbés, Sahara Algérien, 4 – 11. IV. 1965, J. Mateu // Eremomyrme saharica (Gin.), ♂, J. Suárez det., 1976, 1 ♂ [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB2FF4326CCFE881217.taxon	distribution	Distribution. Western Sahara, Morocco, Algeria.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBDFFB2FF4326CCFE881217.taxon	discussion	Remarks. My record of this species from Sudan (Lelej 2002) was wrong and belongs to the Dentilla bischoffi Lelej, sp. nov.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	description	(Figs 36 – 38)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	diagnosis	Diagnosis. MALE. Anterior border of clypeus with broad medial projection whose dorsal length is about half its width. Mandible below with weak apical widening. OOD slightly larger than diameter of lateral ocellus. Head darker than mesosoma. Metasternum 2 - toothed with medial notch. Body length 9.0 – 12.0 mm. FEMALE. T 2 with broad basal band of long appressed silvery setae. T 4 – 5 with pale setae. Eyes closer to posterior margin of vertex than to articulation of mandible. Ratio of smallest distance between eyes to longitudinal diameter of eye is 1.3 ×. Body length 7.0 – 9.0 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	materials_examined	Material examined. (557 ♂, 6 ♀). Type material. Holotypes of Smicromyrme sabulicola Skorikov, 1935, ♀ and Dentilla zarudnyi Lelej, 1985, ♂ [ZIN] and 62 paratypes of D. zarudnyi Lelej, 1985, ♂ from Turkmenistan, Iran, Afghanistan [ZMMU, ZIN, IBSS, NHMD, MZLU]. Additional material. Iran: 58 ♂ (Lelej & Osten 2004). United Arab Emirates: 424 ♂ and 3 ♀. Oman: 5 ♂ (Lelej & van Harten 2011). Iran: Yazd Prov., ca 23 km SW Kharanaq, 1600 m, on light, 32 ° 11 ʹN 54 ° 30 ʹE, A. Timokhov, 1 ♂ [IBSS]; Horrnozgan, Bila'i, 13 – 14. V. 1977 (loc. no. 329), 1 ♀, Exped. Nat. Mus. Praha [Praha]. Pakistan: Baluchistan, Taftan-Quetta road, 160 km W of Quetta, 22. VI. 2003, S. Ovtchinnikov, 3 ♂; 490 km W of Quetta, 26. VII. 2005, S. Ovtchinnikov, 1 ♂ [ZIN]. India: Rajasthan, 15 km W Jaisalmer, 24. VIII. 2004, on light, T. Osten, 1 ♂ [IBSS]. United Arab Emirates: Jebel Hafeet, 293 m, Green Mub dam, 24 ° 03 ʹ 18.5 ʺN, 55 ° 26 ʹ 36.9 ʺE, 4. XI. 2010, 1 ♀, H. Roberts [IBSS]. Oman: South of Sur, 15 km W of Al Askharah, 13. III. 2015, M. Snižek, 1 ♂ [IBSS].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	distribution	Distribution. Afghanistan, Iran, Pakistan (new record), India (new record), Turkmenistan (Lelej 2002); Oman, United Arab Emirates (Lelej & van Harten 2011).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	biology_ecology	Natural history. This nocturnal species inhabits mountainous areas, i. e. stony deserts, and is rarely found in sandy deserts.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBAFFB2FF4322D4FCE617FB.taxon	discussion	Remarks. The record of D. zarudnyi from Yemen (Lelej & van Harten 2011) was incorrect; this species was not recorded by Lelej & van Harten (2006) from Yemen, nor by Lo Cascio et al. (2012) from Socotra. The female of D. zarudnyi Lelej, 1985 may eventually be recognized as D. sabulicola (Skorikov, 1935) due to its co-occurrence in Iran (Sistan & Baluchestan: Bampur and its vicinities, 14 ♂, 1 ♀, 12 – 18. IV. 1901, N. Zarudny), and United Arab Emirates (Wadi Bih dam, 19 ♂, 1 ♀, 30. IV – 4. VI. 2008, LT, leg. A. van Harten). There is no direct evidence (pair collected in copula) to support this relationship.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF43238DFD8A15BD.taxon	description	(Figs 65 – 67)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF43238DFD8A15BD.taxon	diagnosis	Diagnosis. MALE. The male of this species differs from Palaearctic nocturnal species by large apical lobe beneath of mandible. By wide median clypeal process this species similar with D. sabulicola (Skorikov) and D. saharica (Giner) but differs, except mandible shape, by elongate body, and by elongate metasomal segment 1 (length less its maximum width in D. sabulicola and D. saharica). FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF43238DFD8A15BD.taxon	materials_examined	Material examined. (10 ♂). Type material. Holotype [SMNS] and nine paratypes [SMNS, IBSS] of D. osteni from Iran (Lelej & Osten 2004).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF43238DFD8A15BD.taxon	distribution	Distribution. Iran (Hormozgan).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4325B2FD2817C0.taxon	description	(Figs 45, 46)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4325B2FD2817C0.taxon	diagnosis	Diagnosis. MALE. Mandible below without large apical lobe. Length of first metasomal segment less than its maximum width. OOD nearly 2.0 × diameter of lateral ocellus. T 1 dark or slightly red. Mandible quadridentate with lower subbasal tooth. Legs black. Medial cell of propodeum not bordered posterad by carina. Head and antennae ferruginous-red. The lower parts of the pleura and the sterna of mesosoma blackened. The legs are black, the tibial spurs pale. The wing edge is darker brown. Body length 12.0 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4325B2FD2817C0.taxon	materials_examined	Material examined. (3 ♂). Type material. Holotype of Ephutomma semirubra Bischoff, 1920. Additional material. Egypt: Mersa-Matruh, 2 ♂ [NHMW].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4325B2FD2817C0.taxon	distribution	Distribution. Libya, Egypt (new record).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4320CDFECF13AF.taxon	description	(Figs 47 – 50)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4320CDFECF13AF.taxon	diagnosis	Diagnosis. MALE. Ocelli large, OOD 2 × diameter of lateral ocellus. The head is reddish-brown. Wings are hyaline with darkened apex. Anterior border of clypeus with broad medial projection whose dorsal length is about half its width. Propodeum abrupt, reticulate with median dorsal longitudinal cell distinctly delimited posteriorly by two denticles. Whole body light yellow-brown. Body length 10.4 mm. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4320CDFECF13AF.taxon	materials_examined	Material examined. (8 ♂). Type material. Lectotype and three paralectotype of Mutilla testacea in MNHU from “ Arabia deserta ” [Saudi Arabia]. Additional material. 3 ♂ from Yemen and 1 ♂ from Saudi Arabia (Lelej & van Harten 2006).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4320CDFECF13AF.taxon	distribution	Distribution. Yemen, Saudi Arabia.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFBBFFB3FF4320CDFECF13AF.taxon	discussion	Remarks. Francisco-Javier Suárez unpublished labelled the specimen in MNHU with Klug's “ testacea Kl. ” label as the lectotype. Since no validly published lectotype designation has yet been made, I hereby designate the specimen so labelled by Suárez as the lectotype. My designation of the holotype of this species was wrong (Lelej & van Harten 2006).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4322E6FBB215F3.taxon	diagnosis	Diagnosis. MALE. " The males I have placed here are also extremely similar to those of the previous species [Ephutomma continua aurea, ♂ non ♀ = Dentilla ehrenbergi according to Lelej & van Harten 2006], but differ morphologically in that the middle basal field of the middle segment [propodeum] is not bordered by a lamella. In addition, the second tergite of the male also has a stronger shine. Length: 9 – 11 mm] " (Bischoff 1920: 153). Mandible broad, tridentate with subbasal tooth below. Clypeus with a broad medial projection whose dorsal length is about half its width, and belongs to species group Dentilla saharica. FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4322E6FBB215F3.taxon	materials_examined	Material examined. (10 ♂). Holotype, ♂, Algeria: “ SO Algerien [SE Algeria] / Ravis, 15 – 17.4.14 / Bar. v. Geyr S. G. ” [MNHU]. Paratypes (8 ♂ from Algeria and 1 ♂ from Sudan), 9 ♂, “ SO Algerien [SE Algeria] / Ravis, 15 – 17.4.14 / Bar. v. Geyr S. G. ” [MNHU]; Tig'amaiin-en-tisita [Algeria, Tamanraset region], 25 – 30.4.14, Bar. v. Geyr S. G. [MNHU]; Gara Djenoun [Djanet, Algeria] 10 – 14.3.14, Bar. v. Geyr S. G. [MNHU]; Sudan: Chartum, Hartmann [MNHU].	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4322E6FBB215F3.taxon	distribution	Distribution. Algeria, Sudan.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4322E6FBB215F3.taxon	discussion	Remarks. The type series of Ephutomma continua geyri Bischoff consists of 1 ♀ (holotype) and 10 ♂ (paratypes) as designated by Bischoff (1920). Actually, the female belongs to the genus Pseudophotopsis André, 1896 and the male to the genus Dentilla. I have studied in the MNHU the male of Pseudophotopsis fumata Bischoff, 1920 with the same label as the holotype of Ephutomma continua geyri Bischoff, ♀ " SO Algerien [SE Algeria] / Ravis, 15 – 17.4.14 / Bar. v. Geyr S. G. " and I predict that they are the opposite sexes of the same species. The paratypes (male) of Ephutomma continua geyri are described here as Dentilla bischoffi sp. nov.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4320CDFAEA125F.taxon	diagnosis	Diagnosis. MALE. “ A second specimen from Tunisia, Gabés (Weiss 1906), [M. P.] is distinguished [from semirubra - AL] by the somewhat longer, horizontal surface of the middle segment, which is less steeply truncated at the back. I call this form, which should probably be regarded as a separate subspecies, gabesiana n. subsp. The basal field of the middle segment is only vaguely present in both forms. Length: 12 mm. ” (Bischoff 1920: 154). FEMALE. Unknown.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4320CDFAEA125F.taxon	materials_examined	Material examined. No specimen examined.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4320CDFAEA125F.taxon	distribution	Distribution. Tunisia.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB0FF4320CDFAEA125F.taxon	discussion	Remarks. The shape of propodeal posterior face in other species (examined in long series of males) can vary from abrupt to gentle. I think that gabesiana Bischoff, 1920 is simple synonym of semirubra Bischoff, 1920.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB1FF432632FDD210D2.taxon	description	(Figs 68 – 71)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB1FF432632FDD210D2.taxon	diagnosis	Diagnosis. MALE. Unknown. FEMALE. Eyes well protruding from head contour (face view). Head has no developed posterior angles (face view). Longitudinal diameter of eye 2.3 × distance from posterior edge of eye to posterior edge of vertex. T 2 with one median subbasal spot of pale setae. T 4 – 5 with black setae. Body length 7.5 mm.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB1FF432632FDD210D2.taxon	materials_examined	Material examined. Holotype only.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB1FF432632FDD210D2.taxon	distribution	Distribution. Azerbaijan.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB8FFB1FF432632FDD210D2.taxon	discussion	Remarks. This species should be placed in the genus Skorikovia Ovtchinnikov, 2002, based on the shape of the eyes and the very broad scutellar scale, where it may be the opposite sex of Skorikovia transcaucasica (Lelej, 1985), which is known only by the male.	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB9FFBEFF4326F5FF421377.taxon	description	(Figs 72 – 76)	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB9FFBEFF4326F5FF421377.taxon	distribution	Distribution. Russia (Crimea, Dagestan, South and East of European part, South Ural), Bulgaria, Croatia, Czech Republic, Greece, Hungary, Italy (Toscana, Campania, Puglia), Montenegro, Slovakia, Austria, Turkey (new record) (Istanbul, Manisa, Kastamonu, Erzurum), Syria (new record).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
277E1331FFB9FFBEFF4326F5FF421377.taxon	discussion	Remark. After further detailed examination of the holotype of Mutilla curtiventris André, 1901 I conclude that it belongs to the genus Skorikovia Ovtchinnikov, 2002 and is a new junior subjective synonym of Skorikovia trinotata (Costa, 1858). Such a form with a shortened metasoma is already known for this species (pouzdranensis Hoffer).	en	Lelej, Arkady S. (2025): Revision of the genus Dentilla Lelej in Lelej et Kabakov, 1980 (Hymenoptera: Mutillidae). Zootaxa 5633 (2): 288-315, DOI: 10.11646/zootaxa.5633.2.4, URL: https://doi.org/10.11646/zootaxa.5633.2.4
