taxonID	type	description	language	source
244787ED623EFFD7FCDDFD7D1D54FC21.taxon	discussion	Notes. This is an exclusively North American genus; however, the Chilean genus Pleolobus Philippi and Philippi has at times been included. Anorus and Pleolobus are characterized among Karumiinae at the world level by adult males with complete elytra and weakly or non-serrate antennae.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	description	(Figs. 1 – 3)	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	diagnosis	Diagnosis. Males of this species (Figs. 1, 2) are readily separated from Dascillus by the prosternal process forming a short denticle near the anterior coxal margin, resulting in the procoxal cavities being contiguous externally (prosternal process completely separating procoxae in Dascillus). From Anorus piceus it may be separated by the sparse, long, mostly erect setae present throughout the dorsal surface and general lack of short decumbent undersetae (A. piceus with dense, recumbent setae dorsally). The scutellar shield is totally devoid of setae (setose in A. piceus). The pronotum is narrower overall than that of A. piceus, with the width across the anterior angles being roughly equivalent to the width across the posterior angles, and the greatest width of the pronotum is at about the midpoint (A. piceus pronotum wider at posterior margin than anterior margin). The elytron does not possess a sulcus along the lateral margin as in A. piceus and Dascillus. The mandibles are less protruding in this species than A. piceus, and the incisor edge possesses three retinacula before the apex and is not longitudinally straight. Females (Fig. 3) are easily recognized by the disposition of the elytra, which are scale-like and do not extend much beyond the scutellar shield.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	description	Redescription of Male. Body slender and nearly parallel-sided, 6 – 12 mm (Fig. 1). Body setae long, thin, bristling. Antennomeres 3 – 10 very weakly serrate, at least 2 times longer than wide. Left mandible always with two retinacula at midlength of mandible and weak basal retinaculum variably developed. Right mandible similar to left. Epicranial suture not discernible externally. Lateral sides of pronotum not explanate, lateral carina usually indistinct anteriorly, posterior margin straight to weakly bisinuate; pronotum widest near middle, anterior and posterior widths subequal. Hypomeron entirely lacking postcoxal process. Prosternal process forming short denticle (as in Fig. 3 B), barely surpassing anterior margin of procoxae, lacking thin lamina between procoxae; mesoventrite lacking prosternal and procoxal rests entirely. Elytra attaining apex of abdomen; punctation variable, generally with scattered punctures; striae not or extremely weakly impressed. Hind wing attaining apex of abdomen; radial cell widely open proximally; wedge cell small, separated from posterior edge by distance greater than length of cell; apical field about one-third length of wing. All tibiae bearing stout spinules along outer margin. All legs with tarsomeres 1 – 4 bearing small fleshy pads beneath, 1 – 2 with smaller pads, not clearly bilobed, 3 – 4 with larger pads, distinctly bilobed. Terminalia with tergite IX (Figs. 2 C, F) with broad V-shaped emargination along posterior margin; tergite X (Figs. 2 C, F) evenly rounded to subtruncate apically; sternite IX (Figs. 2 B, E) rounded apically; phallobase of aedeagus (Figs. 2 A, D, G – J) distinctly emarginate basally, slightly broader than long; parameres longer than phallobase, very gradually narrowing from base to apex, lateral margin slightly convex, not at all recurved apically, bearing short setae ventromedially in apical half; dorsal lobe of penis measured from anterior margin of basal strut to posterior apex approximately as long as paramere, gradually tapering, apex evenly rounded, never appearing cleft; ventral lobe of penis approximately two-thirds length of dorsal lobe, clawlike, evenly curved ventrally to sharply pointed apex. Description of Female. Body stout (Figs. 3 A, B). Head globose, eye small, subequal to size of second antennomere. Frons smooth and glabrous, temples and occiput rugosely punctate and sculptured, densely covered with long setae. Mandibles stout, well sclerotized, with acute retinaculum preapically on dorsal cutting surface, left mandible with additional small tooth more apically, ventral cutting surface of both with basally positioned, obtuse lobe. Labrum weakly bilobed. Pronotum highly transverse, 2.5 times wider than long, anterior margin strongly and posterior margin weakly bisinuate, pronotum appearing evenly arcuate laterally but lateral carina absent. Scutellar shield transverse, devoid of setae. Elytron weakly sclerotized, shorter than pronotum, transverse, trapezium-shaped, elytra not in contact medially and separated by scutellar shield. Hind wing absent. Abdominal sternite IX distinctly sclerotized, contrasting with previous segments. Terminalia (Figs. 3 C, D) with tergite IX not observed, seemingly weakly sclerotized; sternite VIII (spiculum ventrale) appearing narrowly emarginate, without anterior strut or lobe; proctiger not observed; paraprocts very short, with ventral baculi converging apically; gonocoxites distinctly longer than paraprocts, each coxite with longitudinal baculum with basal thickening; gonostyli present, palpiform.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	discussion	Note on Synonymy and Type Material. Although male specimen series from particular localities seem to be consistent in morphological detail, from the specimens we have examined there is no apparent clinal variation or line of demarcation in width vs. length, pronotal shape (the primary character in the original diagnosis of A. arizonicus) or coloration in males (Fig. 1). Blaisdell (1934) made the comment that “ Arizonicus is a very distinct species, ” but he only examined the type series of five males and apparently only observed two males of what he considered to be A. parvicollis, those being the type and an additional specimen in CASC from Phoenix, Arizona. Additionally, no significant differences were observed among male genitalia (Fig. 2), which are shown here to be distinct from other known species in the family. Clearly a twospecies system representing this complex of populations is a misleading arrangement. Horn (1894) did not mention the number of specimens he used for his original description of A. parvicollis. Examination of the Horn collection (MCZ) revealed 10 male specimens, one labeled “ lectotype ” and the remainder each labeled “ paratype ”. All should be considered syntypes. Two additional male specimens in the LeConte collection (MCZ) are probably syntypes. The holotype of A. arizonicus is a male deposited in CASC (type no. 3735). Four male paratypes are also present in CASC, bearing the same data.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	distribution	Distribution. Throughout the low desert areas of Arizona, south into Sonora (and possibly Sinaloa), Mexico (Fig. 4). The records from California and Nevada represent new state records for this species. Previous records of this species from Baja California, Mexico, are quite possibly in error, since we have seen no specimen vouchers. However, if A. parvicollis is present in Baja California, they may be expected to occur in the lower Colorado Desert in the northeastern portion of the state. The specimens putatively from Sinaloa were intercepted at the United States border within produce shipments that originated from Sinaloa and subsequently transported through this species’ range in Sonora.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED623EFFD1FCF4FBB91EB5FD4E.taxon	materials_examined	Specimens Examined (215). UNITED STATES: ARIZONA: La Paz Co.: 1 mile northwest of Bouse, 1 June 1946, J. R. Slevin (2 ♂♂, CASC); 6 miles southeast of Parker, 7 May 1966, Gorodenski, Davidson and Cazier (1 ♂, ASUHIC); Maricopa Co.: 5 miles east of Mesa, 30 May 1964, S. A. Gorodenski (2 ♂♂, ASUHIC); 5 miles west of Wickenburg, 2,4700 elevation, 5 June 1978, C. Bellamy (1 ♂, LACM); Bush highway junction with Usery Pass road, 23 April 1981, W. B. Warner (1 ♂, ASUHIC); Sunflower, J. McCormick (1 ♂, ASUHIC); Salt River at Blue Point Forest Campground, 27 April 1974, Rockwood, Draper and Kolner (3 ♂♂, ASUHIC); Thunderbird Regional Mountain Park, 23 May 1975, R. S. Wielgus and F. F. Hasbrouck (12 ♂♂, ASUHIC); Wickenburg, 16 May 1937, E. Van Dyke (2 ♂♂, CASC); Wickenburg, 11 May 1938, L. M. Martin (5 ♂♂, LACM); Wickenburg, 1 June 1959, L. A. Stange (4 ♂♂, LACM); Mohave Co.: Lower Boner Canyon, 34.7431 ° 113.5903 °, 15 April – 24 June 2017, Barrier Pitfall Trap, M. A. Johnston (3 ♂♂, MAJC); Pima Co.: 8 km E of Robles Junction, Avra Valley, 32 ° 040 N 111 ° 150 W, 14 – 24 April 1996, UV light, D. Yanega (3 ♂♂, INHS); Ajo, 25 April 1935, F. H. Parker (3 ♂♂, UCRC); Baboquivari Canyon, 9 April 1963, Timberlake (2 ♂♂, UCRC); Santa Catalina Mountains, 8 May 1933, Bryant (1 ♂, CASC); Tucson, 16 May 1937, Bryant (4 ♂♂, CASC); Tucson, 14 May 1940, L. M. Martin (10 ♂♂, LACM); Tucson, 31 May 1956, J. L. Van Deren (1 ♀, UAIC); same except found in a new house, laid eggs (1 ♀, UAIC); Tucson, 5 May 1998, R. L. Otto (4 ♂♂, UAIC); Vail, Mountain Creek Ranch, 32 ° 04.990 N, 110 ° 39.560 W, 1,100 m, 2 – 9 May 2006, Malaise trap in dry wash, M. E. Irwin (1 ♂, SBMNH); Sonoran Desert Museum, 21 April 1961 (1 ♂, SBMNH); Pinal Co.: 10 miles south of Florence, 32.902 ° 111.281 °, 25 April 2015, M. A. Johnston (40 ♂♂, MAJC); same except M. Gimmel and L. Indruchová (27 ♂♂, ASUHIC); same except Gimmel, Johnston and Anzaldo (22 ♂♂, MAJC); Yuma Co.: Aztec, 16 April 1954 (4 ♂♂, LACM); Dateland, 32 ° 4704200 N 113 ° 3202600 W, 15 February – 5 May 2011, W. B. Warner (1 ♂, MAJC; 1 ♀ imaged by William B. Warner, deposited in the Albert Allen Collection of Coleoptera in Star, Idaho); Mohawk, 14 April 1963, Timberlake (1 ♂, UCRC); Yuma, 27 March 1971, J. Kirkpatrick (4 ♂♂, ASUHIC). CALIFORNIA: Riverside Co.: 18 miles west of Blythe, 29 April 1952, Timberlake (2 ♂♂, UCRC); Chuckwalla Road 2 miles east of junction with highway 10, 12 April 1986, J. Rifkind and P. Gum (1 ♂, LACM); Cottonwood Mountains 25 miles east of Indio, 1,8000 elevation, 18 April 1986, J. P. and K. E. S. Donahue (4 ♂♂, LACM); Pinto Wells at Joshua Tree National Monument, 28 April 1962, E. L. Sleeper (1 ♂, CASC); San Bernardino Co.: 44 km S of Needles, 34.4388 ° N, 114.6517 ° W, 28 April – 4 May 2007, Malaise trap in dry wash with flowering Cercidium, M. E. Irwin (4 ♂♂, SBMNH). NEVADA: Clark Co.: Nellis Dunes northeast of Las Vegas, 36 ° 1703000 N 114 ° 5704900 W, 11 June 2010, W. E. Steiner, J. M. Swearingen (2 ♂♂, USNM). MEXICO: SINALOA: intercepted at Nogales, Arizona, on tomato fruit from Culiacán, 24 April 1939 (1 ♂, USNM); intercepted at Nogales, Arizona, on tomato wrapper from San Blas, 12 May 1932, E. C. Harrison (1 ♂, USNM) [these two interception records not included in map]; SONORA: 16 miles south of Hermosillo, 16 February 1963, P. H. Arnaud (7 ♂♂, CASC); 4 miles north of Navojoa, 30 March 1961, K. D. Payton (6 ♂♂, UCRC); Bahia Cholla, 12 April 1954, Menke and Stange (1 ♂, LACM); Keno Bay [= Bahia Kino], 4 April 1953, M. C. Cushner (6 ♂♂, CASC); Puerto Pe ~ nasco, 16 May 1972, Dr Lenczy (7 ♂♂, USNM); San Pedro Bay, 3 April 1953, P. H. Arnaud (2 ♂♂, CASC). Doubtful locality, not mapped: CALIFORNIA: Orange Co.: Seal Beach, 16 May 1938, D. Poole (2 ♂♂, LACM).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	description	(Figs. 5 – 7)	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	diagnosis	Diagnosis. As in A. parvicollis, males of this species (Fig. 5) are readily separated from Dascillus by the prosternal process forming a short denticle near the anterior coxal margin, resulting in the procoxal cavities being contiguous externally (prosternal process completely separating procoxae in Dascillus). It may be separated from A. parvicollis by the dense, recumbent setae present throughout the dorsal surface, including the scutellar shield, and lack of long erect setae, though many specimens have a pelage of suberect setae on the elytra (A. parvicollis bearing sparse, long, mostly erect setae). The pronotum is distinctly narrowest across the anterior angles, then the pronotum gradually widens posteriorly for two-thirds of the length of pronotum, then narrows slightly to the posterior angles (Fig. 5 A). The elytron is more similar to that of Dascillus than to that of A. parvicollis, both in disposition of setae and the presence of a sulcus along the lateral margin extending from behind the humerus nearly to the apical angle. The mandibles are more elongate in this species than A. parvicollis, the incisor edge is longitudinally straight anterior to the labrum, and there are two retinacula (proximal one longer and more acute) before the apex. Females are easily recognized by the disposition of the elytra, which extend about a third to a half of the distance to the tip of the abdomen (Fig. 6; see also Blaisdell 1934: fig. 3).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	description	Redescription of Male. Body parallel-sided, moderately slender, 8 – 14 mm (Fig. 5 A). Body setae of two types, most fairly short and recumbent; elytra, especially laterally, and appendages also bearing scattered moderately long, bristling setae. Antennomeres 3 – 10 weakly serrate, at least 2 times longer than wide. Left mandible bearing two strong retinacula, lacking basal angulation. Right mandible with two retinacula, basal retinaculum very strong and finger-like, more apical retinaculum blunt, appearing as a lobe on the incisor edge near the base of the apical tooth and often nearly absent. Epicranial suture faintly traceable. Lateral sides of pronotum weakly explanate, lateral carina always distinct to anterior margin, posterior margin weakly but distinctly bisinuate; pronotum widest in posterior third, anterior margin always narrower than posterior margin. Hypomeron with distinct but short postcoxal process. Prosternal process acute apically, lateral margins converging near middle of procoxae, continued as thin vertical lamina often concealed by procoxae (as in Fig. 6 B); mesoventrite with abrupt mesal elevation projecting anteriorly, bearing small procoxal rests. Elytra attaining the apex of abdomen, with weakly punctate striate. Hind wing exceeding apex of abdomen; radial cell completely closed proximally; wedge cell medium-sized, separated from posterior edge by distance about half length of cell; apical field about one-third length of wing. All tibiae lacking stout spinules along outer margin. All legs with tarsomeres 1 – 4 bearing fleshy bilobed pads beneath, tarsomere 1 with relatively small lobes. Terminalia with tergite IX (Fig. 5 G) with very broad and shallow V-shaped emargination along posterior margin; tergite X (Fig. 5 G) evenly broadly rounded apically; sternite IX (Fig. 5 F) rounded apically; phallobase of aedeagus (Figs. 5 B – D) weakly emarginate to rounded basally, slightly broader than long; parameres narrow, much longer than phallobase, gradually narrowing from base to apex, lateral margin slightly concave, recurved apically, bearing very few scattered, fine setae in apical half; dorsal lobe of penis comparatively narrow, measured from anterior margin of basal strut to posterior apex approximately as long as paramere, variably narrowed apically, apex evenly to narrowly rounded and never appearing cleft; ventral lobe of penis nearly as long as dorsal lobe, curved dorsally around middle, then curved ventrally near apex, apex narrowly rounded. Redescription of Female. Resembling male in most characters except: Body stouter (Fig. 6), wider in apical half. Elytra shorter than abdomen, about 1.5 – 2 × as long as pronotum, lacking striae. Hind wing less than half length of abdomen. Terminalia (Fig. 7) with tergite IX not observed; sternite VIII (spiculum ventrale) extremely weakly lobed medially; proctiger narrowly triangular; paraprocts short, with ventral baculi converging apically; gonocoxites slightly shorter than paraprocts, each coxite with proximal plate more heavily sclerotized; gonostyli apparently absent.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	distribution	Distribution. Primarily along the drier slopes and valleys of the Coast Ranges in southern California but extending at least to Owens Valley (Inyo Co.) to the northwest and south into Baja California (Fig. 8).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	discussion	Note on Variation and Type Material. LeConte (1859 a) did not mention how many specimens his original description was based on. Four male specimens are deposited in the LeConte collection (MCZ), three of which bear a gold circle (= “ California ”); the latter should probably be considered syntypes. The examined specimens of A. piceus demonstrate less individual and inter-populational variation than is seen in A. parvicollis. Specimens of the former from California and Baja California tend to be similar in terms of coloration and sclerotization. Specimens from the northeastern extent of its range in Nevada tend to be slightly smaller and lighter in color, and the female elytra are proportionately shorter than those of the single female known from California.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6238FFDBFF06FD051D93FCCB.taxon	materials_examined	Specimens Examined (671). UNITED STATES: CALIFORNIA: Inyo Co.: 7 miles west of Westgard Pass, 25 June 1953, H. Washburn (1 ♂, LACM); Alabama Hills, 36.60083 °, 118.10615 °, 1,350 m, 1 June 2016, UV / MV light, M. and L. Gimmel, A. Mayor, M. A. Johnston, C. Wirth, ML 83 (1 ♂, SBMNH); Antelope Spring, 8 mi SW of Deep Springs, 29 June 1961, J. A. Litsinger (1 ♂, MAIC); same except 9 July 1962 (1 ♂, MAIC); Argus Mountains, 8 June 1938, J. A. Comstock (6 ♂♂, LACM); Baker Creek Campground, Big Pine, 37 ° 10.180 N, 118 ° 18.550 W, 8 June 2011, M. Kippenhan (1 ♂, MAIC); Big Pine Canyon, 8,5000, 8 August 1973, G. R. Ballmer (2 ♂♂, UCRC); Independence, 6 June 1918, L. L. Muchmore (2 ♂♂, LACM); Independence, 1 June 1972 (4 ♂♂, UCRC); Junction Ranch, 28 June 1969, L. Lester (2 ♂♂, LACM); Wyman Canyon, White Mountains, 9 July 1967, S. and S. Fromer (3 ♂♂, UCRC); 5 mi W of Big Pine, Baker Creek, 37.15424 ° N, 118.34295 ° W, 4,939 ft, 17 May 2004, UV light and trap, Thomas E. Dimock, 1121 (4 ♂♂, SBMNH); Kern Co.: Democrat Hot Springs, 2,2000 elevation, 16 June 1969, D. Levin (1 ♂, LACM); Keene, 3,000 ft, 7 July 1975, A. Fuller (1 ♂, SBMNH); Rancheria Creek in Piute Mountains, 4,3500 elevation, T 29 S R 33 E S 23, 1 June 1973, J. P. and K. E. Donahue (2 ♂♂, LACM); Los Angeles Co.: 1 mile south of Valyermo, 19 May 1964, M. J. Wargo (3 ♂♂, UCRC); 2.5 miles SSW of Valyermo, 20 June 1959, N. McFarland (5 ♂♂, LACM); 5 miles north of Beverly Hills in Santa Monica Mountains, 1,1000 elevation, 13 July 1955, N. McFarland (3 ♂♂, LACM); Albright (2 ♂♂, LACM); Altadena, 10 May 1959, J. R. Northern (1 ♂, LACM); Angeles National Forest, San Dimas Experimental Forest, Bell Canyon, 34.1826 ° N, 117.7938 ° W, 13 June 2007, M. Caterino and S. Chatzimanolis (5 ♂♂, SBMNH); Angeles National Forest, Ruby Canyon, 34.6060 ° N, 118.5523 ° W, 22 June 2007, at light, M. Caterino and S. Chatzimanolis (4 ♂♂, SBMNH); Angeles National Forest, San Dimas Experimental Forest, Big Dalton Canyon, 34.1793 ° N, 117.7985 ° W, 27 July 2007, M. Caterino (1 ♂, SBMNH); Angeles National Forest, Shake Canyon, 34.7009 ° N, 118.5250 ° W, 2 July 2007, at light, M. Caterino (1 ♂, SBMNH); Big Pine Ranger Station, 5 July 1975, A. V. Evans (16 ♂♂, LACM); Big Pines, 13 July 1971 (6 ♂♂, LACM); Big Pines Recreation Area, 7 July 1983, S. Ziff (3 ♂♂, LACM); Big Pines Recreation Area, 20 July 1983, S. Ziff (2 ♂♂, LACM); Bouquet Canyon, 23 June 1937, N. Westerlund (3 ♂♂, LACM); Bouquet Canyon, 26 June 1937, J. A. Comstock (9 ♂♂, LACM); Bouquet Canyon, 7 July 1937, N. Westerlund and L. M. Martin (16 ♂♂, LACM); Burbank, 21 May 1930, C. H. Hicks (1 ♂, CASC); Burbank, 23 May 1930, C. H. Hicks and F. C. Hadden (3 ♂♂, LACM); Burbank, 30 June 1958 (2 ♂♂, LACM); Chatsworth, 26 June 1920, L. L. Muchmore (2 ♂♂, LACM); Claremont, 19 May 1962, T. Reimo (2 ♂♂, UCRC); Devil’ s Punch Bowl, 21 June 1964, D. G. Marqua and P. H. Sullivan (1 ♂, LACM); Devil’ s Punchbowl, 11 June 1975 (15 ♂♂, LACM); Eaton Canyon Park, 5 June 1964, R. H. Crandall (2 ♂♂, LACM); Glendale, 3 July 1965, E. I. Schlinger (1 ♂, UCRC); Gorman, 22 July 1938, J. von Bloeker (6 ♂♂, LACM); Juniper Hills, 8 June 1975, A. V. Evans (23 ♂♂, LACM); Juniper Hills, 4,3000 elevation, 14 July 1979, C. L. Bellamy (2 ♂♂, LACM); La Canada, 8 July 1950, D. C. Blodget (3 ♂♂, LACM); La Canada, 21 June 1969 (1 ♂, UCRC); La Crescenta, 19 June 1955, J. Roberds (6 ♂♂, LACM); La Verne, 24 June 1961, R. S. Erdman (4 ♂♂, LACM); Lake Hughes, 13 July 1938 (1 ♂, LACM); Littlerock, [day / month blank] 1945 (1 ♂, LACM); Littlerock, [day blank] May 1947, G. P. Mackenzie (1 ♂, LACM); Mint Canyon, 21 June 1949, H. E. Cott (1 ♂, SBMNH); Mount Wilson (2 ♂♂, LACM); Pomona, 6 April 1931, W. D. Clancy (1 ♂, UCRC); Santa Monica, 22 July 1943, G. P. Mackenzie (1 ♂, UCRC); Sierra Madre, 11 July 1944 (1 ♂, LACM); Sunland, 12 May 1940 (1 ♂, LACM); Tanbark Flats, 4 July 1950, D. C. Blodget (1 ♂, LACM); Tujunga, 19 June, 1955, J. Roberts (1 ♂, LACM); Tujunga, 14 May 1969, J. C. von Bloeker (30 ♂♂, LACM); Tujunga Canyon, 15 May 1969, A. Strong (1 ♂, UCRC); Littlerock, 8 June 1941, G. P. Mackenzie (1 ♂, UCRC); Topanga Canyon, 6 June 1978 (1 ♂, LACM); Pasadena, 25 May 1894 (1 ♂, LACM); Pasadena, [day blank] May [year blank] (1 ♂, LACM); San Marino, 5 July 1942, G. P. Mackenzie (1 ♂, UCRC); Valyermo, 1 July 1967, S. Ziff (2 ♂♂, LACM); West Los Angeles, 17 March 1967, B. S. Cheary (1 ♂, UCRC); [county record only], [day blank] June 1938, J. Schwartz (1 ♂, LACM); Orange Co.: 4 miles east of Olive, 1 July 1975, J. Wilcox (1 ♂, UCRC); Orange Park, 11 May 1978, B. P. Harris (1 ♂, LACM); Riverside Co.: 4 miles south of Beaumont, at Potrero Creek, 1 July 1990, G. R. Ballmer and D. C. Hawks (1 ♂, UCRC); 6 kilometers north of Desert Center, 18 March 1986, W. E. Steiner and J. E. Lowry (2 ♂♂, USNM); Beaumont, 10 June 1942, E. Van Dyke (1 ♂, CASC); Bee Canyon near Strawberry Canyon, 25 May 1969 (♂, LACM); Box Spring Mountains, 30 June 1964, G. E. Wallace (1 ♂, UCRC); same except C. D. Howell (4 ♂♂, UCRC); Big Morongo, 22 April 1985, M. Clement (3 ♂♂, UCRC); Big Morongo Canyon, 2,0000 elevation, 2 May 1969, C. D. Howell (3 ♂♂, UCRC); Boggert Trail, T 5 S R 4 E S 14, 25 April 1998, J. D. Barry (5 ♂♂, UCRC); Bradshaw Trail Road 4 miles west of Graham Pass road, 33 ° 25.60 N 115 ° 14.40 W, 11 April 1994, R. R. Snelling (5 ♂♂, LACM); Citrus Experimental Station, 13 May 1964, P. A. Rauch (1 ♂, UCRC); Corona, 5 June 1955, R. H. Alvarado (1 ♂, UCRC); Deep Canyon, Sec. 6 T 7 s, R 6 E, 16 May 1975, L. E. Clark (9 ♂♂, UCRC); Dripping Springs, 33 ° 270 N 116 ° 580 W, 13 May 1999 (3 ♂♂, UCRC); Gavilan Hills, 15 May 1965 (11 ♂♂, UCRC); Indian Wells, 24 April 1953, G. Yamamoto (1 ♂, LACM); Lake Elsinore, 28 May 1981, G. Pratt (2 ♂♂, UCRC); Lost Palms Canyon in Joshua Tree National Monument, 26 May 1963, E. Sleeper (1 ♂, CASC); Morongo Valley 1 mile north of Indian Avenue, 3 May 1969, B. Hunt (2 ♂♂, UCRC); P. L. Boyd Desert Research Center, 3.5 miles south of Palm Desert, 3 May 1974, D. L. Dickson (1 ♂, UCRC); Pinyon Flats, 6 June 1970, Cazier, Francke and Welch (12 ♂♂, ASUHIC); Pinyon Flats in San Jacinto Mountains, 27 May 1939, J. G. Shanafelt (2 ♂♂, LACM); Pinyon Flats in San Jacinto Mountains, 27 June 1952, B. Tinglof (2 ♂♂, LACM); Pinyon Flats, 4,0000 elevation, 26 May 1980, C. L. Bellamy (1 ♂, LACM); Riverside, 2 May 1961, L. D. Anderson (1 ♂, UCRC); Riverside, 5 May 1961, O. L. Brawner (2 ♂♂, UCRC); Riverside, 2 May 1961, C. S. Papp (5 ♂♂, LACM); Riverside, 1 May 1962, B. Ewing (4 ♂♂, UCRC); Riverside, 7 May 1962, E. I. Schlinger (3 ♂♂, UCRC); Riverside, 26 June 1962 (1 ♂, UCRC); Riverside, 14 May 1964, M. E. Irwin (1 ♂, UCRC); Riverside, 29 May 1964, M. E. Irwin (1 ♂, UCRC); Riverside, 13 May 1972, A. Tabet (8 ♂♂, UCRC); Riverside, 16 May 1972, A. Tabet (3 ♂♂, UCRC); Riverside, 23 May 1972, A. Tabet (2 ♂♂, UCRC); Riverside, 2 June 1972, A. Tabet (2 ♂♂, UCRC); San Bernardino National Forest, San Jacinto River, 33.7244 ° N, 116.8061 ° W, 24 May 2005, at light, M. Caterino (5 ♂♂, SBMNH); Split Rock, Joshua Tree National Monument, 24 May 1966, J. Wilcox (3 ♂♂, UCRC); Tahquitz Canyon, 8 June 1957, Menke and Stange (2 ♂♂, LACM); Temecula Canyon, Santa Margarita River, 7500 elevation, 18 May 1968, L. O. Tejada (1 ♂, UCRC); Tenaja Campground, 14 May 1972, A. Tabet (3 ♂♂, UCRC); Tenajas Ranger Station, 29 July 1967, J. A. Honey (3 ♂♂, LACM); Hurkey Creek Campground, 2 May 1972 (1 ♂, USNM); Whitewater Canyon, 16 May 1969, B. Duff (1 ♂, USNM); Whitewater Canyon, 33 ° 6202200 N 116 ° 4104200 W, J. D. Barry (1 ♂, UCRC); Whitewater Canyon, T 2 S R 3 E S 26, 650 m elevation, 6 May 1997, H. W. Park (3 ♂♂, UCRC); [county record only] 13 May 1966, M. W. Stone (3 ♂♂, UCRC); 24 May 1966, M. W. Stone (3 ♂♂, UCRC); Whitewater Canyon, 33.9692 ° N, 116.6521 ° W, 15 May 2006, M. Caterino and S. Chatzimanolis (6 ♂♂, SBMNH); San Bernardino Co.: 44 km S of Needles, 34.4388 ° N, 114.6517 ° W, 28 April – 4 May 2007, Malaise trap in dry wash with flowering Cercidium, M. E. Irwin (3 ♂♂, SBMNH); Alta Loma, 11 June 1955, R. H. Alvarado (2 ♂♂, UCRC); Apple Valley, 8 July 1959, N. McFarland (1 ♂, LACM); Baldy Mesa, 13 June 1937, J. A. Comstock (2 ♂♂, LACM); Cajon Pass, 24 June 1937 (1 ♂, LACM); Cajon Pass, 24 June 1949, H. E. Cott (1 ♂, SBMNH); Clark Mountain, T 17 N R 13 E SE quarter S 22, 25 May 1985, J. P. and K. E. Donahue (3 ♂♂, LACM); Cucamonga Canyon, 22 May 1972, T. Stemwedel (1 ♂, LACM); Deep Creek Campground, 15 June 1937, Menke and Stange (4 ♂♂, LACM); Fontana, 19 May 1959, R. J. Jeffrey (2 ♂♂, UCRC); same except 20 May 1959 (2 ♂♂, UCRC); Horse Spring Campground in San Bernardino Mountains, 5700 ’ elevation, 13 June 1986, J. P. and K. E. S. Donahue (6 ♂♂, LACM); Joshua Tree National Park, 49 Palms Oasis Trail, 34.1109 ° N, 116.1072 ° W, 27 – 28 April 2012, UV light, M. S. Caterino, A. K. Tishechkin and M. P. Richmond (4 ♂♂, SBMNH); Joshua Tree National Park, 49 Palms Oasis Trailhead, 34.1193 ° N, 116.1118 ° W, 27 – 28 April 2012, D. Yanega (1 ♂, SBMNH); Joshua Tree National Park, 49 Palms Oasis, 34.1058 ° N, 116.1051 ° W, 27 – 28 April 2012 (1 ♂, SBMNH); Joshua Tree National Park, Smithwater Canyon, 34.0213 ° N, 116.2961 ° W, 25 – 27 April 2013, night collecting, M. S. Caterino and E. Shafer (1 ♂, SBMNH); Lower Covington Flat in Joshua Tree National Monument, 28 May 1960, E. Sleeper (1 ♂, CASC); Lytle Creek, 8 June 1928, E. Van Dyke (1 ♂, CASC); Mill Creek, 957 m elevation, 31 May 1994, M. Cooperband (1 ♂, UCRC); Oak Glen, 2 June 1970, M. J. Wargo (1 ♂, UCRC); Ontario, 1 June 1938 (1 ♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 2 August 1994, J. Freilich (1 ♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 28 May 1995, J. Freilich (1 ♂, UCRC); Pioneertown, UC Burns Reserve, 1350 m elevation, 1 July 1995, J. Freilich (2 ♂♂, UCRC); Providence Mountains, 7 May 1939, J. A. Comstock (2 ♂♂, LACM); San Bernardino National Forest, Gobbler’ s Knob, 34.3116 ° N, 117.5835 ° W, 3 July 2005, M. Caterino (3 ♂♂, SBMNH); Seven Oaks, 8 July 1950 (1 ♂, LACM); Twentynine Palms, 26 April 1949 (2 ♂♂, LACM); UC Burns Reserve, 34.1403 ° N, 116.4542 ° W, 22 May 2005, at light, M. Caterino (1 ♂, SBMNH); Victorville, 20 May 1935, C. Dammers (2 ♂♂, LACM); Wildwood Canyon, 5 miles east of Calimesa, 28 July 1965, E. I. Schlinger (1 ♂, UCRC); Wrightwood, 19 June 1966, R. H. Crandall (6 ♂♂, LACM); Wrightwood, 20 July 1983, S. Ziff (38 ♂♂, LACM); Wrightwood, 2 June 1984, S. Ziff (1 ♂, LACM); San Diego Co.: Alpine, 20 April 1960, S. C. Williams (2 ♂♂, ASUHIC); Borrego, 5 April 1940 (3 ♂♂, LACM); Borrego, 27 April 1955, Timberlake (1 ♂, UCRC); Boulder Oaks, 30 June 1957, D. Peterson (2 ♂♂, UCRC); Boulevard near I- 8 exit, 3 June 1995 (1 ♂, MAJC); Culp Canyon, 2 April 1962, J. C. Hall (1 ♂, UCRC); Encinitas, 26 May 1932, H. L. McKenzie (3 ♂♂, UCRC); Indian Flats Campground, 3,8000 elevation, 28 May 1968, J. A. Honey (9 ♂♂, LACM); Jacumba, [day / month blank] August 1955, D. Cox (1 ♂, ASUHIC); La Puerta Valley, 23 April 1938 (4 ♂♂, LACM); La Puerta Valley, 20 May 1939, L. M. Martin (3 ♂♂, LACM); Lawson Valley, 12 June 1939, J. von Bloeker (4 ♂♂, LACM); Oak Grove, 28 June 1966, J. R. Dixon (2 ♂♂, LACM); Palm Canyon, Borrego Valley, 28 May 1936, Timberlake (15 ♂♂, UCRC); Palm Canyon, Borrego Desert, 2 April 1946, A. L. Melander (1 ♂, UAIC); Palm Canyon, Borrego Valley, 26 April 1953 (2 ♂♂, LACM); Palomar Mountain, 22 June 1940, G. P. Mackenzie (1 ♂, UCRC); Point Loma, 17 June 1952, R. V. D. Rosch (1 ♂, UCRC); Scissors Crossing, Valle de San Felipe, 2,240 – 2,8000 elevation, 4 May 1969, E. I. Schlinger (2 ♂♂, UCRC); Sentenac Canyon, 26 April 1953, G. Yamamoto (1 ♂, LACM); Sentanac Canyon, 1,800 – 1,9000 elevation, 4 May 1968, J. DeVol (1 ♂, UCRC); Torrey Pines, 6 July 2009, N. Bloomfield (1 ♂, USNM); [county record only], F. E. Blaisdell (1 ♀, CASC); [county record only] 10 June 1966, M. W. Stone (1 ♂, UCRC); 27 June 1967, M. W. Stone (2 ♂♂, UCRC); San Luis Obispo Co.: 7 mi E of Pozo, 4 July 1967, blacklight, C. W. Baker and D. H. Wilson (1 ♂, SBMNH); Carrizo Plain National Monument, 35.0440 ° N, 119.6009 ° W, 24 May 2003, mercury vapor light, M. Caterino and P. Jump (13 ♂♂, SBMNH); Carrizo Plain National Monument, Selby Campground, 35.1273 ° N, 119.8430 ° W, 5 June 2004, at light, M. Caterino and P. Jump (3 ♂♂, SBMNH); Los Padres National Forest, 27 June 1987, L. Hunsaker (1 ♂, SBMNH); Mountains west of La Panza, 18 May 1928, J. O. Martin (4 ♂♂, CASC); Paso Robles, 24 May 1928, L. S. Slevin (1 ♂, CASC); Paso Robles, 12 June 1971, C. E. Langstrom (1 ♂, LACM); Santa Margarita, 12 June 1971, C. E. Lengstrom (4 ♂♂, LACM); Santa Barbara Co.: 0.75 mi S of Ballinger Wash, 34.876 ° N, 119.482 ° W, 2,840 ft, 13 June 2007, UV light trap, P. M. Jump (3 ♂♂, SBMNH); same except 20 June 2007 (5 ♂♂, SBMNH); same except 7 June 2007 (2 ♂♂, SBMNH); Ballinger Canyon, check dam, 34.87665 °, 119.48225 °, 860 m, 6 May 2017, UV light trap, Peter Jump (3 ♂♂, SBMNH); state record only: May 1908, R. Woglum (1 ♂, UCRC). NEVADA: Clark Co.: Charleston Peak, 4 June 1946 (3 ♂♂, CASC); Nye Co.: Mercury Nevada Test Site, 18 May 1960 (1 ♂, MAIC; 6 ♂♂, USNM); same except 19 May 1960 (3 ♂♂, USNM); same except 19 May 1961 (3 ♂♂, USNM); same except 25 May 1961 (1 ♂, MAIC; 3 ♂♂, USNM); same except 7 June 1961 (1 ♂, USNM); same except 16 June 1964 (1 ♀, USNM); same except 20 June 1964 (1 ♂, 2 ♀♀, MAIC; 1 ♂, 3 ♀♀, USNM); same except 12 June 1965 (1 ♂, USNM). MEXICO: BAJA CALIFORNIA [NORTE]: 4 miles south of La Rumorosa, 4,4000 elevation, 7 July 1967, M. E. Irwin (5 ♂♂, UCRC); 12 miles east of El Rosario, 10 June 1979 (4 ♂♂, UCRC); Arroyo del Rosario 3 miles above Rosario, 26 April 1963, P. H. Arnaud (5 ♂♂, CASC); La Zapopita, Valle de Trinidad, 7 April 1961, F. S. Truxal (7 ♂♂, LACM); La Zapopita, Valle de Trinidad, 16 April 1961, F. S. Truxal (9 ♂♂, LACM); Parque Nacional Constitución de 1857, 12 km S of Laguna Hanson, Sierra Juárez, 31.96 ° N, 115.99 ° W 1,700 m, 15 June 2019, A. Shamaev (3 ♂♂, MAIC); San Telmo Canyon in San Pedro Martir, 30 May 1981, Werner, Olson, Hetz, Thomas, Burn, Frank and MacLachlan (4 ♂♂, UAIC); Santo Tomas, 8 August 1973, S. C. Williams and K. B. Blair (2 ♂♂, CASC); Tijuana, 18 June 1964 (1 ♂, UCRC). Doubtful localities, not mapped: CALIFORNIA: Yolo Co.: Davis, 10 May 1960, J. L. Bath (2 ♂♂, UCRC). BAJA CALIFORNIA SUR: Santa Inez, 26 April 1979, D. B. Weissman (1 ♂, CASC) [probably in Baja Norte or mislabeled]. WYOMING: Converse Co.: Douglas, 27 July 1964, H. B. Leech (1 ♂, CASC) [almost certainly mislabeled].	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6232FFD8FCD6FCE31F70FEC2.taxon	discussion	Notes. This genus contains approximately 30 additional species distributed in the Old World (Fang et al. 2020; Jin et al. 2013 b, 2016, 2017; Terzani et al. 2017; Wang et al. 2019 a, b), where the type species, Dascillus cervinus (Linnaeus, 1758), occurs. The genus can only be diagnosed in its present constitution using a combination of characters (Jin et al. 2013 b). Consequently, monophyly of the genus is an open question, particularly with regard to the North American species, representatives of which were not considered in detail in the most comprehensive modern treatment of Jin et al. (2013 b). Larval evidence is consistent with the view that the North American species is an isolated lineage (Grebennikov and Scholtz 2003). It is worth noting that Jin et al. (2013 b: 559) credited the fixation of Chrysomela cervina as the type species of Dascillus to the listing found in Thomson (1859: 107); however, Latreille (1802: 98) already established this as the type species according to subsequent monotypy (see ICZN 1999: Art. 69.3). Therefore, the synonymy of Atopa proposed by Latreille (1802: 99) is solidified by objective synonymy of the two genera, since they share the same type species.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	description	(Figs. 9 – 11)	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	diagnosis	Diagnosis. By far the largest and most robust dascillid species in North America, D. davidsoni (Figs. 9 – 11) can easily be recognized by its size (10 mm or greater in total length), robust overall body shape, and the wide prosternal process separating the procoxal cavities externally and reaching their posterior margin (coxae contiguous in Anorus species).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	description	Redescription of Male. Body parallel-sided, 10 – 18 mm (Fig. 9). Body setae variable, recumbent, never bristling. Antennomeres 3 – 10 serrate, subtriangular. Left mandible with one distinct, blunt retinaculum and a second weak and very blunt retinaculum apicad. Right mandible with single retinaculum. Epicranial suture indistinct but traceable. Lateral sides of pronotum explanate, dorsal and ventral surfaces well demarcated by complete lateral carina, posterior margin strongly bisinuate; pronotum evenly narrowing from base to apex. Prosternal process relatively large, distinctly surpassing posterior margin of procoxae; mesoventrite with anteriorly projected prosternal rest (boatshaped cavity to accept prosternal process), bearing distinct procoxal rests. Hypomeron with distinct angulate postcoxal process. Elytra weakly punctatestriate, striae becoming slightly impressed posteriorly. Hind wing about 2.5 times longer than greatest width, attaining apex of abdomen; radial cell completely closed proximally; wedge cell large, separated from posterior edge by distance less than half length of cell; apical field about one-fourth length of wing. All tibiae lacking stout spinules along outer margin. All legs with tarsomeres 1 – 4 bearing fleshy bilobed pads beneath. Terminalia with tergite IX (Fig. 10 C) with broad V-shaped emargination along posterior margin; tergite X (Fig. 10 C) evenly rounded apically; sternite IX (Fig. 10 B) rounded apically; phallobase of aedeagus (Figs. 10 A, D – G) weakly emarginate basally, slightly broader than long; parameres longer than phallobase, narrowing from base to apex, lateral margin straight until recurved laterally near apex, bearing scattered long setae laterally in apical half; dorsal lobe of penis measured from anterior margin of basal strut to posterior apex approximately as long as paramere, elongate triangular, apex weakly to broadly rounded and weakly sclerotized medially, sometimes appearing cleft; ventral lobe of penis distinctly shorter than dorsal lobe, triangular, evenly tapering to apex, curved ventrally near apex, apex truncate to slightly emarginate. Redescription of Female. Similar to male except: Larger, body stouter (Fig. 11 A), widest in apical half of elytra. Epicranial suture very distinct and sulcate. Hind wing slightly narrower than in male, about 2.6 times longer than greatest width but attaining apex of abdomen. Terminalia (Figs. 11 B, C) with tergite IX distinctly broadly emarginate along anterior margin; sternite VIII (spiculum ventrale) narrowly triangular; proctiger narrowly triangular; paraprocts relatively short, with ventral baculi converging apically; gonocoxites subequal in length to paraprocts, each coxite with proximal lobe heavily sclerotized ventrally, triangularly produced medially; gonostyli absent. Note on Synonymy and Variation. After examination of 630 specimens of North American Dascillus, we concluded that the two traditionally recognized names refer to expressions of conspecific color morphs. The traditional concept of D. davidsoni, here referred to as the “ davidsoni-morph ” (Fig. 9 A), refers to specimens with a lighter reddish-brown cuticle with pale setae forming a dark double zig-zag across the elytra. The traditional concept of D. plumbeus, here referred to as the “ plumbeus- morph ” (Fig. 9 B), includes specimens without a zig-zag pattern, only rows of pale to dark setae on the elytral interstriae, often giving a pinstriped appearance. Additionally, the cuticle of the elytra in the latter tends to be darker, piceous to nearly black, although dark or light reddish-brown cuticle can occur along with either setal pattern (e. g., Figs. 9 C, D). Consequently, abraded individuals are nearly impossible to definitively assign to morph. On the other hand, particular morphs tend not to be mixed at a given locality, and there are very few individuals that are intermediate, i. e., that express a faint zig-zag pattern. Regions that include predominantly plumbeus- morph include the Sierra Nevada foothills and the interior Coast Range north of San Francisco Bay, while the davidsoni- morph dominates the southern and western sides of the San Francisco Bay and the Pacific slopes of the Coast Ranges. Localities from which we have seen both morphs include Martinez (Contra Costa Co.), Mill Valley (Marin Co.), Hopland (Mendocino Co.), Mount Saint Helena (Napa Co.), Mount Hamilton (Santa Clara Co.), and Sylvania (county unknown), while faint-patterned intermediate individuals were prevalent along a transition zone in the central portion of the northern Coast Ranges, including Fort Seward (Humboldt Co.) and Clearlake Oaks (Lake Co.). In addition to variation in color pattern, the last abdominal ventrite (ventrite 5) in the male is variably emarginate at the apex, but this is not observably correlated with other variation. Horn (1880), in the original description of D. plumbeus, used the relative lengths of antennomeres 3 and 4 to separate the two Dascillus species, but we were unable to make use of this character across our larger sampling of specimens and localities. Such synonymy and individual variation is not without precedent in the genus Dascillus. Terzani et al. (2017) published on the widespread, variable species D. cervinus of the Western Palearctic region. They observed variation in the coloration of both the integument and setation, the shape of abdominal sternite IX, and even among structures of the aedeagus, such as the ventral lobe of the penis, which ranged from parallel-sided to narrowing, and the dorsal lobe of the penis, which ranged from indistinctly to distinctly projecting medially (see Terzani et al. 2017: fig. 2). Additionally, apparent variation in the shape of the parameres should be interpreted with caution; these structures are flexible and highly sensitive to distortion resulting from mounting medium, level of clearing, and from the amount of extrusion of the penis. The very limited genitalic variation (Figs. 10 A, D – G) seen in D. davidsoni as circumscribed herein falls well within the magnitude of intraspecific variability observed within certain Old World Dascillus species concepts.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	distribution	Distribution. Central California north to Humboldt County, south to Santa Barbara County, and west to the western foothills of the Sierra Nevada (Fig. 12). Specimen records from Arizona, Oregon, and Mexico (see below) seem to be erroneous, although its occurrence in southwestern Oregon would not be unexpected.	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	discussion	Note on Type Material and Spelling. LeConte (1859 b) did not mention how many specimens he used for the original description of D. davidsoni, but there must have been at least two since he compared male and female characters. However, Horn (1880: 84) stated “ The female of Davidsoni is unknown to us ”, implying that LeConte mistook one or more of his male specimens for a female. Our examination of photographs of specimens in the LeConte collection (MCZ) revealed 11 specimens, all male; these 11 should probably be considered syntypes. Horn (1880) did not mention how many specimens he used for the original description of D. plumbeus, but there must have been at least two since he compared male and female characters. Examination of photographs of specimens labeled as this species in the LeConte and Horn collections (MCZ) revealed eight specimens, including six males and two (probable) females; these eight should probably be considered syntypes. One specimen is labeled “ lectotype ”, but this designation has never been published. For this species, Horn apparently observed genuine females, describing them as much larger (20 mm) and more robust, having the last ventrite slightly shorter, and the flight wings less developed than those of the male and “ probably unsuitable for flight. ” Two large specimens of plumbeus- morph from the Horn collection in the MCZ appear to be females. The subsequent spelling Dascillus davidsoni LeConte is in prevailing usage (Horn 1880; Jin et al. 2013 b; Leng 1920; O’ Keefe 2002) and should be maintained (ICZN 1999: Art. 33.3.1).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6231FFC0FF02FE911A62FD4E.taxon	materials_examined	Specimens Examined (630). NOTE: The various morphs of this species are indicated by letters in square brackets after the numbers of specimens (d = davidsoni- morph; p = plumbeus- morph; i = intermediate). UNITED STATES: CALIFORNIA: Alameda Co.: Alameda, 22 May 1932, E. S. Ross (1 ♂ [d], CASC); Albany Hill, 16 April 1998, Robert L. Langston (1 ♂ [d], CASC); Berkeley, 13 May 1917, R. Hopping collection (1 ♂ [d], CASC); Berkeley, 25 April 1921, J. O. Martin (1 ♂ [d], CASC); Berkeley, 26 April 1924, F. C. Hadden (1 ♂ [d], LACM); Berkeley, April 1925, R. M. and G. E. Bohart (1 ♂ [d], CASC); Berkeley, February 1932 (1 ♀ [d], LACM); Berkeley, 8 May 1938, J. G. Shanafelt (1 ♂ [d], LACM); Berkeley, 24 April 1948, D. W. Buddy (1 ♂ [d], CASC); Berkeley, 16 May 1949, R. S. Beal (1 ♂ [d], ASUHIC); CSCH Campus, Hayward, 17 May 1967, Kenneth France (1 ♂ [d], CASC); “ Dimond G. ”, 14 May 1900, F. E. Blaisdell (1 ♂ [d], CASC); Hayward, May 1911 (2 ♂♂ [d], LACM); hills back of Oakland, 19 April 1919, Van Dyke collection (5 ♂♂ [d], CASC); Livermore, 11 May 1930, Van Dyke collection (1 ♂ [d], CASC); Mills College, 19 April 1926, Thos. Craig (1 ♂ [d], CASC); Niles, 14 April 1914, E. J. Branagan (1 ♂ [d], UCRC); Niles, 19 April 1914, Blaisdell collection (1 ♂ [d], CASC); Niles Canyon, 23 May 1917, W. M. Gifford (1 ♂ [d], CASC); Oakland, Van Dyke collection (2 ♂♂ [d], CASC); Oakland, April 1901, Van Dyke (8 ♂♂ [d], CASC); Oakland, 10 April 1925, F. C. Hadden (1 ♂ [d], LACM); Oakland, 18 April 1931, E. S. Ross (1 ♂ [d], CASC); same except 21 May 1932 (1 ♂ [d], CASC); Oakland, May 1982, F. R. Barnes (1 ♂ [d], CASC); Oakland, E. J. Blum (2 ♂♂ [d], CASC); Oakland Hills, 18 April 1926, Van Dyke collection (4 ♂♂ [d], CASC); Oakland Hills, 12 April 1940, J. J. Marquis (5 ♂♂ [d], CASC); Pleasanton, 23 April 1939, G. S. Mansfield (1 ♂ [d], CASC); San Leandro, Elsie Avenue, 5 May 1983, Thomas W. Davies (1 ♂ [d], CASC); Strawberry Canyon, 29 April 1956, C. L. Hogue (2 ♂♂ [d], LACM); Sunol, 14 May 1922, F. E. Blaisdell (1 ♂ [d], CASC); Sunol Valley Regional Park, 9 May 1964, W. K. Fox (1 ♂ [d], ASUHIC); Sunol Valley Regional Park, 10 May 1967, V. Lee (1 ♂ [d], CASC); Tunnel Road, 13 May 1917, W. M. Giffard (1 ♂ [d], CASC); [county record only], Van Dyke collection (3 ♂♂ [d], CASC); [county record only], R. Hopping collection (4 ♂♂ [d], CASC); [county record only], April, Koebele collection (1 ♂ [d], CASC); [county record only], Leng, R. Hopping collection (1 ♀ [d], CASC); [county record only], Blaisdell collection (1 ♀ [d], CASC); [county record only] (1 ♀ [d], CASC); Butte Co.: Oroville, 9 April 1928, H. H. Keifer (2 ♂♂ [p], CASC); Oroville, 30 April 1971 (5 ♂♂ [p], UCRC); Oroville, 2 May 1971 (4 ♂♂ [p], UCRC); Calaveras Co.: Mokelumne Hill, Blaisdell collection (15 ♂♂ [p], CASC); same except 3 May 1893 (1 ♂ [p], CASC); same except 11 May 1893 (4 ♂♂ [p], CASC); Mokelumne Hill, 20 May 1942, F. E. Blaisdell (1 ♂ [p], CASC); Colusa Co.: [county record only], 20 May 1951, M. Marquis (1 ♂ [d], CASC); Contra Costa Co.: Concord, 26 April 1942, H. P. Chandler (3 ♂♂ [d], CASC); Danville, 8 April 1962, R. Brown (1 ♂ [d], CASC); same except 14 May 1962 (1 ♂ [d], CASC); Lafayette, 10 May 1947, T. W. Fisher (1 ♂ [d], UCRC); Lafayette, 22 May 1951 (2 ♂♂ [d], CASC); Martinez, 21 April 1909, Van Dyke collection (1 ♂ [d], CASC); same except April 1910 (1 ♀ [d], CASC); same except June 1910, Van Dyke collection (1 ♂ [p], CASC); Moraga Ranch, 7 May 2009, S. E. Haskins (1 ♂ [d], MAJC); Mount Diablo, 21 June 1931 (4 ♂♂ [d], LACM); Mount Diablo, 24 April 1927, B. E. White (1 ♂ [d], CASC); Mount Diablo, 24 April 1932, O. Bryant (4 ♂♂ [d], CASC); Mount Diablo, base, 28 April 1932, O. Bryant (1 ♂ [d], CASC); Mount Diablo, 30 April 1933, E. S. Ross (1 ♂ [d], CASC); Mount Diablo, 21 May 1937 (1 ♂ [d], CASC); Mount Diablo, 18 May 1951 (2 ♂♂ [d], CASC); Mount Diablo, northeast slope of North Peak, 30 April 1967, R. M. Brown (1 ♂ [d], CASC); Russelmann Park, east slope of Mount Diablo, 19 April 1931, E. P. Van Duzee (1 ♂ [d], CASC); Russelmann Park, Mount Diablo, 17 May 1960, D. C. Rentz (1 ♂ [d], CASC); Russelmann Park, Mount Diablo, 24 April 1966, R. M. Brown (11 ♂♂ [d], CASC); Russelmann Park, 2 miles SE Clayton, 1,200 ft, 20 May 1967, Paul H. Arnaud, Jr. (1 ♂ [d], CASC); Russelmann Park, 20 May 1967, J. W. Tilden (1 ♂ [d], CASC); Walnut Creek, Castle Rock Park, 10 May 1975, Harvey I. Scudder (1 ♂ [d], CASC); [county record only], 20 July 1911, F. W. Nunenmacher (1 ♂ [d], CASC); El Dorado Co.: Camino, 12 May 1979, D. Plubell (1 ♂ [p], CASC); Coloma, 260 m, hillside, under oaks, 24 May 1981, Paul H. Arnaud, Jr. (1 ♂ [p], CASC); Diamond Springs, 15 May 1954, Truxal and Martin (1 ♂ [p], LACM); Pine Hill, west of Rescue, 1,5000, 2 May 1970, D. P. Levin (2 ♂♂ [p], LACM); Humboldt Co.: Blocksburg, 21 May 1933, Brunson P. Bliven collection (1 ♂ [d], CASC); same except 8 April 1934 (2 ♂♂ [d], CASC); Blocksburg, 25 May 1935, H. J. Rayner (1 ♂ [d], CASC); Blocksburg, 19 April 1936, B. P. Bliven (2 ♂♂ [d], CASC); same except 3 May 1936 (1 ♂ [d], CASC); Benbow, 23 April 1941, B. P. Bliven (1 ♂ [d], CASC); Eureka, 3 June 1935, E. O. Essig (1 ♂ [p], CASC); Fort Seward, 21 May 1935, F. R. Platt (1 ♂ [d], LACM; 1 ♂ [d], UCRC); Fort Seward, 18 May 1935, E. O. Essig (1 ♂ [i], CASC); same except 5 June 1935 (1 ♀ [i], CASC); same except 20 May 1935, H. J. Rayner (2 ♂♂ [i], CASC); same except 24 May 1935 (2 ♂♂ [i], CASC); same except 28 May 1935 (1 ♂ [i], CASC); Fort Seward, 23 May 1935, E. W. Baker (1 ♂ [i], CASC); Fort Seward, 25 May 1935, E. O. Essig (1 ♂ [i], CASC); same except 25 May 1935 (1 ♂ [i], CASC); Garberville, 28 April 1940, B. P. Bliven (2 ♂♂ [d], CASC); Korbel, 5 June 1916, F. E. Blaisdell (1 ♂ [d], CASC); Miranda, 25 April 1937, B. P. Bliven (1 ♂ [p], CASC); Lake Co.: 8 miles north of Kelseyville, 11 May 1952, sweeping, F. B. Turner (1 ♂ [p], LACM); Anderson Springs, 20 May 1951, W. R. Bauer (1 ♂ [p], CASC); same except 17 May 1952 (1 ♂ [p], CASC); Blue Lakes, 16 May 1922, E. P. Van Duzee (1 ♂ [p], CASC); Clearlake Oaks, 19 May 1951, Don Burdick (1 ♂ [i], CASC); Clearlake Oaks, 20 May 1951, J. W. Tilden (1 ♂ [i], CASC); Lower Lake, 13 May 1922, E. P. Van Duzee (2 ♂♂ [p], CASC); Snow Mountain, 22 May 1971, J. Kraemer (1 ♂ [p], CASC); Soda Bay Springs, Kelseyville, 15 May 1922, E. P. Van Duzee (1 ♂ [p], CASC); Upper Lake, 17 May 1922, E. P. Van Duzee (1 ♂ [d], CASC); [county record only], May 1895, Van Dyke collection (11 ♂♂ [p], CASC); [county record only], Van Dyke collection (5 ♂♂ [p], CASC); Marin Co.: Cypress Ridge, 27 April 1920, J. O. Martin (1 ♂ [d], CASC); Fairfax, 11 May 1919, E. P. Van Duzee (4 ♂♂ [d], CASC); same except 9 May 1920 (1 ♂ [d], CASC); same except 4 May 1928 (1 ♂ [d], CASC); Fairfax, 12 April 1925, C. L. Fox (1 ♂ [d], CASC); Fairfax, 1 May 1949, W. A. Russell (1 ♂ [d], CASC); Lagunitas, 11 May 1924, E. P. Van Duzee (1 ♂ [d], CASC); Lake Lagunitas, 29 May 1959, D. C. Rentz (1 ♂ [d], CASC); Mill Valley, 28 April 1913, R. Hopping collection (2 ♂♂ [d], CASC); same except 10 May 1914 (1 ♂ [d], CASC); same except 22 April 1915 (1 ♂ [d], CASC); Mill Valley, 15 May 1948, Frances O. Leech (1 ♀ [d], CASC); same except 24 May 1948 (1 ♂ [d], CASC); Mill Valley, 25 May 1948, H. B. Leech (1 ♂ [d], CASC); same except 1 June 1948 (1 ♂ [d], CASC); same except 7 June 1948 (1 ♂ [d], CASC); same except 16 May 1949 (1 ♂ [d], CASC); same except 22 May 1949 (2 ♂♂ [d], CASC); Mill Valley, June 1948, E. S. Ross (1 ♀ [d], CASC); Mill Valley, 9 May 1950, W. H. Leech (1 ♂ [d], CASC); Mill Valley, 16 May 1959, J. L. Bath (1 ♂ [p], UCRC); Mill Valley, 26 April 1965, P. H. Arnaud, Jr. (1 ♂ [d], CASC); same except 2 – 4 May 1965 (1 ♂ [d], CASC); same except 6 – 10 May 1965 (2 ♂♂ [d], CASC); Mill Valley, 5 May 1965, Loyal Weitz (1 ♂ [d], CASC); Mill Valley, chaparral zone, May 1980, E. S. Ross (1 ♀ [d], CASC); Mount Tamalpais, 28 April 1907, Blaisdell collection (2 ♂♂ [d], CASC); Mount Tamalpais, Old Stage Road, at flower of blue flowering Lupinus, 22 May 1999, Paul H. Arnaud, Jr. (1 ♂ [d], CASC); Muir Woods, 21 April 1921, J. O. Martin (1 ♂ [d], CASC); Novato, 23 April 1961, D. C. Rentz (1 ♂ [d], CASC); Paradise Cove, 6 May 1951, J. W. Green (1 ♂ [d], CASC); Phoenix Lake vicinity, 3 May 1962, D. C. Rentz (1 ♂ [d], CASC); Ross, 28 April 1918, E. P. Van Duzee (1 ♂ [d], CASC); Ross, 9 April 1939, B. E. White (1 ♂ [d], CASC); Samuel P. Taylor State Park, along Lagunitas Creek, 18 May 1977, D. D. Wilder (2 ♂♂ [d], CASC); [county record only], April, Blaisdell collection (2 ♂♂ [d], CASC); between Alp. [illegible], 18 May 1958, D. C. Rentz (1 ♂ [d], CASC); [county record only], Bolter collection (2 ♂♂ [d], INHS); Mariposa Co.: Jerseydale, 8 miles northeast of Mariposa, 3,5000, 14 May 1974, Oakley Shields (1 ♂ [p], LACM); Mendocino Co.: Hopland, 9 May 1926, E. P. Van Duzee (2 ♂♂ [d], CASC); Hopland Experimental Station, ¼ mile southwest of Poor Ranch, 28 April 1972, E. Rogers (2 ♂♂ [p], CASC); Hopland Field Station, Kelsey Cab. Orchard area, 2,500 – 2,800 ft, 14 May 1977, R. Greenberg (1 ♂ [p], CASC); same except M. P. Pavel (1 ♂ [p], CASC); Hopland Field Station, Headquarters Lake, 900 – 1600 ft, 15 May 1977, M. P. Pavel (1 ♂ [p], CASC); same except N. J. Vandenberg (1 ♂ [p], CASC); Orrs Springs, 7 May 1936, Van Dyke collection (1 ♂ [d], CASC); Philo, 13 May 1933 (1 ♂ [d], CASC); Philo, 14 May 1933, J. W. Tilden (1 ♂ [d], CASC); same except 20 May 1938 (2 ♂♂ [d], CASC); Ukiah Grade, 10 May 1926, E. P. Van Duzee (1 ♂ [p], CASC); Yorkville, 17 May 1929, E. P. Van Duzee (3 ♂♂ [d], CASC); same except 8 May 1935 (1 ♂ [d], CASC); Monterey Co.: Arroyo Seco, 10 May 1969, J. W. Tilden (1 ♂ [d], CASC); Arroyo Seco, 14 May 1994, J. G. Edwards (1 ♂ [d], CASC); Bradley, 28 April 1955, R. P. Allen (4 ♂♂ [d], CASC); Lockwood, 3 April 1957, R. P. Allen (20 ♂♂ [d], CASC); same except 25 April 1958 (6 ♂♂ [d], CASC); Paraiso Springs, 3 May 1922, L. S. Slevin collection (1 ♂ [d], CASC); same except 4 May 1922 (3 ♂♂ [d], CASC); same except 5 May 1922 (1 ♂ [d], CASC); Napa Co.: 2 miles north-northeast of Angwin, north side of Howell mountain, 1,300 ft, 25 April 1976, H. B. Leech (1 ♂ [p], CASC); same except 2 May 1976 (1 ♂ [p], CASC); same except 4 May 1976 (1 ♂ [p], CASC); same except 26 April 1979 (1 ♂ [p], CASC); same except 3 May 1979 (1 ♂ [p], CASC); same except 11 May 1979 (1 ♂ [p], CASC); same except 24 April 1980 (1 ♂ [p], CASC); same except 2 May 1980 (1 ♂ [p], CASC); same except 6 May 1980 (1 ♂ [p], CASC); same except 7 May 1980 (1 ♂ [p], CASC); same except 7 May 1981 (2 ♂♂ [p], CASC); same except 29 April 1982 (1 ♀ [p], CASC); same except 4 May 1984 (1 ♂ [p], CASC); same except 18 May 1976, F. O. Leech (1 ♂ [p], UCRC); same except “ adult died ” 15 March 1982, “ reared from 45 mm long larva ex garden soil, 7. III. 1981; pupated Sept. 1981, adult emerged 28. X. 1981 ” (1 ♀ [p], CASC); same except “ 18. III. 1981 as larva in soil ” and “ found in garden soil, 12 inches below surface; reared; as adult in soil, late October 1981, died 3. I. 1982 ” (1 ♂ [p], CASC); Angwin, 16 May 1957, Laurell Rouse (1 ♂ [p], CASC); same except 25 May 1957 (1 ♂ [p], CASC); same except 23 May 1959 (1 ♂ [p], CASC); Angwin, 16 May 1958, Elizabeth McCart (2 ♂♂ [p], CASC); Angwin, 9 May 1960, Marilyn Brandt (1 ♂ [p], CASC); Angwin, 20 May 1961, [collector illegible] (1 ♂ [p], CASC); Angwin, 2 May 1962, B. Easterbrook (1 ♂ [p], CASC); Angwin, 18 May 1971, T. Griswold (1 ♂ [p], CASC); Angwin, 25 May 1972, G. Miller (1 ♂ [p], CASC); Angwin, 10 May 1972 (1 ♂ [p], CASC); Angwin, 30 May 1972 (1 ♂ [p], CASC); Angwin, 6 May 1977 (1 ♂ [p], CASC); Angwin, 10 May 1960, L. Eighme (1 ♂ [p], CASC); same except 3 June 1975 (2 ♂♂ [p], CASC); same except 5 June 1975 (1 ♂ [p], CASC); same except 10 May 1978 (1 ♂ [p], CASC); same except 6 June 1978 (1 ♂ [p], CASC); same except 30 May 1984 (1 ♂ [p], CASC); Angwin, 8 May 1980, Peter Cowley (1 ♂ [p], CASC); Angwin, no date, H. B. Leech (5 ♂♂ [p], UCRC); Calistoga, 1 June 1948, D. Giuliani (1 ♂ [p], CASC); Conn Dam, 18 May 1968, E. S. Ross (1 ♂ [p], CASC); Huichica Creek, 17 June 1934, J. W. Tilden (1 ♂ [i], CASC); Mount Saint Helena, 16 June 1915, Van Dyke collection (1 ♀ [p], CASC); Mount Saint Helena, 9 June 1918, E. P. Van Duzee (2 ♂♂ [p], CASC); Mount Saint Helena, 26 May 1951 (1 ♂ [p], CASC); Mountain Mill House, Highway 29, 12 km north of Calistoga, 475 – 550 m, 11 June 1988, Paul H. Arnaud, Jr. (1 ♂ [p], CASC); Napa, 10 June 1931, A. T. McClay (1 ♂ [d], CASC); Saint Helena, in vineyard, 26 April 1986, Ed Salvestrin (1 ♂ [d], CASC); Sanitarium, 10 May 1960, R. Ratzlaff (1 ♂ [p], CASC); Nevada Co.: 6 miles south of Grass Valley, 18 May 1930, E. P. Van Duzee (1 ♂ [p], CASC); Grass Valley, 17 May 1930, E. P. Van Duzee (2 ♂♂ [p], CASC); Nevada City, 16 May 1930, E. P. Van Duzee (1 ♂ [p], CASC); Placer Co.: 10 miles northeast of Auburn, 5 May 1958, A. E. Menke (1 ♂ [p], LACM); Auburn, 20 April 1969, B. Swisley (1 ♂ [p], CASC); Auburn, 7 May 1969, B. Paul (1 ♂ [p], CASC); [county record only], Van Dyke collection (4 ♂♂, 1 ♀ [p], CASC); Foresthill, 9 February 1977, E. L. Klee (1 ♂ [d], CASC); San Benito Co.: Pinnacles National Monument, 26 June 1967, P. S. Bartholomew (1 ♂ [d], CASC); San Francisco Co.: San Francisco, 31 May 1905, Van Dyke collection (1 ♂ [d], CASC); San Francisco, 1 May 1931, J. A. Kusche (3 ♂♂ [d], CASC); San Luis Obispo Co.: Cerro Alto, 12 May 1979, Selfridge (1 ♂ [d], SBMNH); Reservoir Canyon, 29 April 1979, L. Verhovek (1 ♂ [d], SBMNH); San Luis Obispo, 24 April 1918, E. P. Van Duzee (2 ♂♂ [p], CASC); Tassajara Canyon, 30 April 1978, R. Hervey (1 ♂ [d], SBMNH); [county record only], 18 June 1989, L. Penzo (1 ♂ [d], SBMNH); San Mateo Co.: Crystal Springs Road, 16 April 1957, Cushner (4 ♂♂ [d], CASC); Crystal Springs Road, 27 April 1957 (11 ♂♂ [d], CASC); same except D. Rentz (4 ♂♂ [d], CASC); East Palo Alto, 6 May 1955, P. S. Bartholomew (1 ♂ [d], CASC); Jasper Ridge, Stanford University, 20 April 1952, P. S. Bartholomew (3 ♂♂ [d], CASC); same except 22 April 1962 (1 ♂ [d], CASC); Menlo Park, 10 May 1955, DHB (5 ♂♂ [d], CASC); Redwood City, 11 April 1943 (1 ♂ [p], UCRC); San Mateo, 25 April 1949, R. S. Beal (1 ♂ [d], ASUHIC); San Mateo, Van Dyke collection (1 ♂ [d], CASC); Westridge, 31 June 1959, P. S. Bartholomew (1 ♂ [d], CASC); Santa Barbara Co.: Refugio Canyon, 15 April 1962, P. van Duinwyk (1 ♂ [d], SBMNH); Santa Clara Co.: Alma, 28 May 1916, C. L. Fox (1 ♂ [d], CASC); Alma, Van Dyke collection (2 ♂♂ [d], CASC); Almaden, 28 April 1946, L. L. Lewallen (2 ♂♂ [d], UCRC); Alum Rock Park, San Jose, 26 June 1921, L. S. Slevin (2 ♂♂ [d], CASC); Alum Rock Park, 23 April 1948, P. H. Arnaud (1 ♂ [d], CASC); Alum Rock Park, San Jose, 22 April 1951, J. W. Green (1 ♂ [d], CASC); Alum Rock Park, 22 April 1958, William Walls (1 ♂ [d], CASC); Alum Rock Park, 5 May 1965, on wild blackberry, D. H. Kavanaugh (1 ♂ [d], CASC); Alum Rock Park, 1 May 1975, Harvey I. Scudder (2 ♂♂ [d], CASC); same except 25 April 1979 (1 ♂ [d], CASC); near Calaveras Reservoir, 13 May 1961 (1 ♂ [d], CASC); same except G. I. Hage (1 ♂ [d], CASC); Cupertino, 4 May 1937, K. S. Hagen (1 ♂ [d], CASC); Cupertino, 12 April 1950, R. S. Beal (2 ♂ [d], ASUHIC); Los Altos, 7 May 1921, W. M. Giffard (3 ♂♂ [d], CASC); Los Gatos, 23 April 1909, L. S. Slevin collection (1 ♂ [d], CASC); same except 20 May 1910 (1 ♂ [d], CASC); Los Trancos Creek, Stanford University, 10 May 1952, P. S. Bartholomew (1 ♂ [d], CASC); Mount Hamilton, 20 May 1933, grasses and low herbage, E. S. Ross collection (1 ♂ [p], CASC); Mount Hamilton, 14 April 1934, in poppy flowers (1 ♂ [d], CASC); Mountain View, 1909, R. Hopping collection (2 ♂♂ [d], 1 ♀ [d], CASC); Palo Alto, 22 March 1892 (1 ♂ [d], LACM); Palo Alto, 5 April 1892 (2 ♂♂ [d], LACM); Palo Alto, 6 April 1892 (6 ♂♂ [d], LACM); Palo Alto, 12 April 1892 (1 ♂ [d], LACM); Palo Alto, 20 April 1892, L. J. Muchmore (1 ♂ [d], LACM); Palo Alto, 25 April 1892 (1 ♂ [d], LACM); Palo Alto, 4 May 1892 (1 ♂ [d], LACM); Palo Alto, 21 April 1893 (1 ♂ [d], LACM); Palo Alto, 21 April 1895 (1 ♂ [d], LACM); Palo Alto, 22 April 1895 (1 ♂ [d], LACM); Palo Alto, April 1896 (1 ♂ [d], LACM); Palo Alto, 8 April 1896 (2 ♂♂ [d], LACM); Palo Alto, 16 April 1896 (1 ♀ [d], LACM); 22 April 1897 (1 ♂ [d], LACM); Palo Alto, April 1908, Van Dyke collection (1 ♂ [d], CASC); Palo Alto, 9 April 1981, S. F. Muzzio (1 ♂ [d], CASC); San Jose, April 1906 (1 ♂ [d], LACM); San Jose, 6 May 1956, Dave Ribble (1 ♂ [d], INHS); San Jose, front porch lights, 13 November 1960, student collector (1 ♀ [d], CASC); San Jose, 5 May 1965, Balcom (1 ♀ [d], CASC); San Jose, ex grass blade, 3 May 1976, Suzanne Muzzio (1 ♂ [d], CASC); same except ex Pinus, 16 April 1978 (1 ♂ [d], CASC); same except ex grass, 28 April 1979 (1 ♂ [d], CASC); Sand Hill Road, Stanford University, 30 March 1952, P. S. Bartholomew (1 ♂ [d], CASC); Silver Creek Hills, 11 April 1961, R. L. Macdonald (1 ♂ [d], UCRC); Silver Creek Hills, 28 April, 1966, A. R. Stephen (1 ♂ [d], CASC); Silver Creek Hills, 9 May 1968, A. and A. Gillogly (2 ♂♂ [d], LACM); Stanford University, May 1898, Kuwana (1 ♂ [d], LACM); Stanford University, 18 April 1902 (1 ♂ [d], LACM); Stanford University, Mar 19 [blank] (1 ♂ [d], LACM); Stanford University, 11 April 1905 (1 ♂ [d], LACM); Stanford University, 13 April 1905 (1 ♂ [d], LACM); Stanford University, 20 March 190 [blank] (1 ♂ [d], LACM); Stanford University, 4 May 190 [blank] (1 ♂ [d], LACM; 1 ♂ [d], UCRC); Stanford University, 20 March 190 [blank] (1 ♂ [d], LACM); Stanford University, April 1923 (1 ♂ [d], LACM); Stanford University, 13 April 1923 (2 ♂♂ [d], LACM); Stanford University, 20 April 1923 (1 ♂ [d], LACM); Stanford University, April 1925 (1 ♂ [d], LACM); Stanford University, May 1928, Hungate (1 ♂ [d], LACM); Stanford University, April 1927, Shedd (1 ♂ [d], LACM); same except May 1927 (1 ♂ [d], LACM); Stanford University, April 1929 (1 ♂ [d], LACM); Stanford University, 30 April 1933, D. Scott, Jr. (1 ♂ [d], CASC); Stanford University, 28 April 1934 (1 ♂ [d], CASC); Stanford University, 13 May 1950, P. S. Bartholomew (2 ♂♂ [d], CASC); same except 21 May 1951 (1 ♂ [d], CASC); same except 10 May 1952 (1 ♂ [d], CASC); same except 5 May 1953 (1 ♂ [d], CASC); same except 17 May 1953 (4 ♂♂ [d], CASC); same except 27 April 1954 (1 ♂ [d], CASC); same except 29 April 1954 (1 ♂ [d], CASC); same except 17 April 1960 (1 ♂ [d], CASC); Stevens Creek, 5 May 1940, G. S. Mansfield collection (1 ♂ [d], CASC); Stevens Creek, 19 May 1974, E. L. Klee (3 ♂♂ [d], CASC); Stevens Creek area, 20 April 1950, Alice and J. G. Edwards (1 ♂ [d], CASC); Stevens Creek area, 28 May 1952 (1 ♂ [d], CASC); Sunnyvale, 29 March 1949, T. H. Lauret (1 ♂ [d], CASC); Sunnyvale, May 1971, D. H. Bartholomew (1 ♀ [d], CASC); Uvas Creek, 13 May 1979, J. G. Edwards (1 ♂ [d], CASC); William Street Park, San Jose, 14 May 1965, in tall grass, D. H. Kavanaugh (1 ♂ [d], CASC); [county record only], 25 April 1950, Alice and J. G. Edwards (3 ♂♂ [d], CASC); [county record only], 6 – 27 April 1974, P. S. Bartholomew (1 ♀ [d], CASC); Shasta Co.: South Fork Cow Creek, Millville, 7,000 ft, 13 May 1949, H. P. Chandler (1 ♂ [p], CASC); [county record only], F. W. Nunenmacher, 6 May 1913 (1 ♂ [p], CASC); Solano Co.: Putah Canyon, 16 April 1958, A. Menke (2 ♂♂ [p], LACM); Solano Lake, 24 April 1970, R. Melton (1 ♂ [p], LACM); Vacaville, 9 July 1967, Verda Fletcher (3 ♂♂ [p], CASC); Sonoma Co.: 3 miles northwest of Healdsburg, host weeds, 28 March 1986, John Guardeno (1 ♂ [d], CASC); 5.5 miles east of Guerneville, 23 April 1966, Peter Rubtzoff (1 ♂ [d], CASC); Bennett Mountain Lake, west of Kenwood, 1,180 ft, H. B. Leech (1 ♂ [d], CASC); Cazadero, 31 March 1934, G. E. Bohart (1 ♂ [d], CASC); Eldridge, 1 May 1912, Van Dyke collection (2 ♂♂ [i], CASC); same except June 1912 (5 ♂♂ [p], CASC); same except 9 June 1912 (1 ♂ [p], CASC); Geyserville, 27 April 1971 (1 ♂ [p], UCRC); Geyserville, 10 May 1979, E. L. Klee (1 ♂ [p], CASC); Healdsburg, 16 May 1974 (1 ♂ [d], CASC); Laguna de Santa Rosa, 31 April, 1960, J. Sedlacek (4 ♂♂ [d], CASC); Mark West Springs, 10 May 1930, E. P. Van Duzee (1 ♂ [d], CASC); same except 14 May 1930 (1 ♂ [d], CASC); Mount Saint Helena, 6 May 1930, Van Dyke collection (1 ♂ [d], CASC); Santa Rosa, 16 April 1890 (4 ♂♂ [d], LACM); Santa Rosa, Ricksecker (1 ♂ [d], CASC); Santa Rosa, 15 April 1934 (1 ♂ [d], LACM); Santa Rosa, 12 May 1940, J. W. Green collection (1 ♂ [d], CASC); Sobre Vista, 15 April 1910, Van Dyke collection (2 ♂♂ [i], CASC); same except 24 April 1910 (1 ♂ [d], CASC); same except 31 April 1910 (3 ♂♂ [d], CASC); [county record only] (6 ♂♂ [d], LACM); [county record only], 19 May 1952, J. Quast (1 ♂ [d], LACM); [county record only], Koebele collection (1 ♂ [d], 1 ♂ [p], CASC); Sutter Co.: Yuba City, 29 May 1971, Marc P. Hayes (1 ♂ [p], SBMNH); Yolo Co.: Capay, 29 April 1934, B. E. White collection (1 ♂ [p], CASC); Davis, 22 May 1948, A. T. McClay (4 ♂♂ [p], CASC); same except 5 May 1947 (1 ♂ [p], UCRC); same except 14 April 1950 (1 ♂ [p], CASC); same except 13 May 1950 (3 ♂♂ [p], CASC); same except 12 May 1951 (7 ♂♂ [p], CASC); Davis, 27 April 1958, F. D. Parker (1 ♂ [p], UCRC); Davis, 20 April 1960, L. Traynham (1 ♂ [p], SBMNH); Davis, 1 May 1964, R. L. Macdonald (1 ♂ [p], UCRC); Putah Canyon, 25 May 1938, J. W. Tilden (1 ♂ [p], CASC); Putah Canyon, 24 April 1950, E. J. Taylor (1 ♂ [p], CASC); Putah Creek, 15 April 2000, S. E. Haskins (1 ♂ [p], MAJC); same except 9 – 29 May 2000 (2 ♂♂ [p], MAJC); Rumsey, 29 April 1934, B. E. White collection (1 ♂ [p], CASC); same except 3 May 1936 (1 ♂ [p], CASC); same except 3 May 1938 (1 ♂ [p], CASC); Rumsey, 17 May 1958, A. E. Menke (2 ♂♂ [p], LACM); Yuba Co.: Marysville, 7 August 1937, in peach orchard (1 ♂ [d], LACM); county unknown: Sylvania, April, Ricksecker (4 ♂♂ [i], CASC); Sylvania, 5 May 1895, R. Hopping collection (4 ♂♂ [d], CASC); [state record only], 26 April 1908, R. Hopping collection (1 ♂ [d], CASC); [state record only], Leng, R. Hopping collection (1 ♀ [d], CASC); [state record only], Fall, Fenyes collection (1 ♂ [p], CASC); Rust, 30 April 1913, L. S. Slevin collection (1 ♂ [d], CASC); [no locality], ex Carl Fuchs collection via E. R. Leech collection (1 ♀ [d], CASC); [no locality], J. W. Green collection (1 ♀ [d], CASC); [no locality] E. S. Ross collection (1 ♂ [p], CASC). Doubtful localities, not mapped: ARIZONA: Santa Cruz Co.: Patagonia, April 1976, R. T. Ross (1 ♂ [d], CASC); OREGON: Marion Co.: Salem, 20 March 1961 (1 ♂ [d], ASUHIC); MEXICO: country record only, Ricksecker (1 ♂ [d], CASC).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
244787ED6228FFC1FEEFFDAE1F02FC23.taxon	discussion	LeConte (1853: 229) stated that he had “ not seen this species, but learn from my father that it occurs in Georgia. ” Gemminger and Harold (1869: 1615) listed it from “ Georgia amer. ” Horn (1880: 84) stated that it “ should be stricken from our [United States] lists, it is a native of Santo Domingo. ” This taxon, to our knowledge, has not been treated in the literature or placed in checklists since 1880. It is currently under study by M. A. Ivie and K. Guerrero (M. A. Ivie, in litt.).	en	Johnston, M. Andrew, Gimmel, Matthew L. (2020): Review of North American Dascillidae (Coleoptera: Dascilloidea), with Descriptions of Dramatic Female Wing Reduction. The Coleopterists Bulletin 74 (4): 731-757, DOI: 10.1649/0010-065X-74.4.731, URL: http://dx.doi.org/10.1649/0010-065x-74.4.731
