taxonID	type	description	language	source
245B87DB9D41171233EFCFC768E9FABC.taxon	description	(Figs 1, 2, 3; Tables 1, 2, 3)	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D41171233EFCFC768E9FABC.taxon	materials_examined	TYPE MATERIAL. — Türkiye • [C 7 Urfa] in fissuris rupium calcareaum regionis calidae ad Mar Jacob supra Orfa (Urfa) Mesopotamiae; 15. V. 1865; Haussknecht s. n.; lectotype: JE [JE 00003786] image!; isolectotypes: B [B 100241949] image!, BM [BM 000632423] image!, G-BOIS [G 00330685] image, G [G 00418089] image!, JE [JE 00003785] image!, K [K 000838527] image!, LE [LE 00013283] image!, MW [MW 0592720] image!), P [P 04022552, P 04022553] images!, W [W 0005250, W 1889 - 0223437] images! OTHER MATERIAL EXAMINED. — Türkiye • B 8 Diyarbakır, Eğil to Dicle 4 km, oak openings, rock crevices; 1042 m alt.; 23. V. 2018; Ertekin 2018 - 085; DUF • C 7 Şanlıurfa, North of Kızılkuyu Wildlife Development Area, near beltway, rock crevices; 729 m alt.; 27. V. 2020; Ö. F. Kaya 6471 • Nemrut mountain, surroundings of Mar Jacob monastery ruins, rock crevices; 712 m alt.; 27. V. 2020; Ö. F. Kaya 6479.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D41171233EFCFC768E9FABC.taxon	distribution	DISTRIBUTION AND ECOLOGY. — R. haussknechtii s. str. is an endemic species native to the Southeastern Anatolia Region and an IranoTuranian element. Bare crevices or rockies on open areas of oak forest form the plant habitat. R. haussknechtii s. str. grows in rockies or in rock areas at oak glades at 700 - 1050 meters altitudes in the Mesopotamian steppes and, three populations of R. haussknechtii s. str. were identified in Şanlıurfa and Diyarbakır provinces. The distribution area of the species for Şanlıurfa is approximately 4 km 2 with an average of 350 - 400 individuals, while it is approximately 2 km 2 for Diyarbakır with an average of 150 - 200 individuals. The main factors such as road construction, land use change, grazing and urbanization threaten the plant environment / habitat. Therefore, it has been suggested that the plant should be evaluated in Vulnerable-VU B 1 b (iii) category according to IUCN Standards & Petitions Committee (2024) Red List categories.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D41171233EFCFC768E9FABC.taxon	discussion	REMARKS The holotype was not specified, in the protologue of Rosularia haussknechtii by Boissier (1872). This was ignored by Chamberlain & Muirhead (1972) when a lectotype should have been determined. On the other hand, according to ICN (Turland et al. 2018), Eggli (1988) used the term holotype instead of lectotype in his original choice of lectotype for Rosularia haussknechtii. This is an error (see point 9.10). However, since this erroneous typing was before January 1, 2001 (see Article 9.23), the original selection by Eggli (1988) is valid. Unnumbered samples first collected by Haussknecht in 1865 from a single locality in Urfa (now Şanlıurfa) were named as Umbilicus haussknechtii by Boissier and Reuter (Boissier 1872). There are 13 sheets collected by Haussknecht only from this type locality and only found in B, BM, G, JE, K, LE, MW, P, W herbaria. Although “ typus ” was written on the sheets found in B, G, P and W herbaria, taxonomists, who examined these samples, stated them as “ isotypus ”. The sheet (MW 0592720) only found in MW was labelled as “ isolectotypus ”.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D41171233EFCFC768E9FABC.taxon	description	DESCRIPTION Caudex well developed, 4 - 8 mm diam. Rosettes persistent after flowering, semiglobular, 2 - 5 cm diam.; offsets many; scapes lateral, 5 - 10 cm long, green or pink. Rosette leaves obovate-spathulate to narrowly spathulate, 18 - 30 × 3 - 6 mm, uninerved, attenuate at base, densely pectinate-ciliate with scattered glandular hairs on both surfaces, apex subacute. Lower cauline leaves linear, oblong-linear, 8 - 12 × 1.5 - 2 mm; median leaves linear or oblong-linear, 16 - 24 × 2 - 3 mm, dull green or deep pink, hirsute, densely ciliate, apex obtuse; floral leaves smaller and densely glandular pilose 7 - 12 × 2 mm. Inflorescence a lax thyrse, densely hairy, 8 - 17 flowered, always lateral, 1 - 3 per rosette, axis glandular-hairy in upper 2 / 3 imbricate leafy. Flowers 5 - merous, narrowly campanulate. Pedicel 1 - 4 mm. Bracts 2 - 4 mm, bracteoles lanceolate, hairy. Calyx 4 - 5 mm, densely hairy, tube 1 - 1.5 mm, teeth lanceolate, 3 - 4 mm. Corolla 10 - 12 mm, pale yellow, back of petals densely pilose, petals connate 2 / 3 to 3 / 4, corolla lobes spreading, 3 - 4 mm, ovate, yellow, with purple or pinkish veined, margins denticulate, mucronate, c. 0.5 mm, corolla tubes greenish, 7 - 8 mm, densely pilose, longer than the calyx. Stamens 10, biseriate, longer stamen 7 - 8 mm, shorter stamens 5 - 6 mm long. Follicles 5 - 6 mm, finely hairy, styles 1 mm, stigmas capitate. Flowering in May. Rocky places, open areas of oak forest.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D44171230C4C9906EBFFA0A.taxon	description	(Fig. 3)	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D44171230C4C9906EBFFA0A.taxon	materials_examined	TYPE MATERIAL. — Türkiye • Tunceli, Munzur dağı, above Ovacık, limestone rocks; 2800 m alt.; 17. VII. 1957; P. H. Davis & I. C. Hedge (D. 31242); E! OTHER MATERIAL EXAMINED. — Türkiye • East Anatolia, in Armeniae petrosis; s. d.; Aucher-Éloy 2652; G! • Same data; Aucher-Éloy 2653; G! • Erzincan, N. W. side of Tayran dağı; 1500 - 1800 m alt.; Watson et al. 32981.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D44171230C4C9906EBFFA0A.taxon	distribution	DISTRIBUTION AND ECOLOGY. — R. pseudohaussknechtii is an endemic species native to Eastern Anatolia, identified from the Munzur Mountains and the nearby Tayran Mountains. The distribution area of the species is limited and it grows on calcareous rocks at elevations between 1500 and 2800 meters. The distribution area and population status of R. pseudohaussknechtii could not be evaluated due to insufficient data. While R. haussknechtii s. str. is distributed in Southeastern Anatolia, R. pseudohaussknechtii is distributed in Eastern Anatolia. Some morphological characteristics of both species are shown in detail in Table 2.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
245B87DB9D44171230C4C9906EBFFA0A.taxon	description	DESCRIPTION Caudex slender, 2 - 3 mm diam. Rosettes depressed 1 - 1.5 cm diam; scapes lateral, 6 - 12 cm, densely hairy. Rosette leaves oblong to oblong-spathulate, 8 - 12 × 2 - 3 mm, ciliate. Cauline leaves sparsely; lower narrowly obovate-spathulate, 7 - 12 × 2 - 3 mm, obtuse; median and upper leaves shorter and narrower, oblong to linear-spathulate, 6 - 9 × 1 - 1.5 mm, densely hairy. Inflorescence a lax thyrse, densely hairy, 7 - 13 flowered, lateral, 1 - 3 per rosette. Flowers 5 - merous, narrowly campanulate. Pedicel 3 - 6 mm. Bracts minute, c. 2 mm, hairy. Calyx 4 - 5 mm, densely hairy, teeth lanceolate, connate at base. Corolla 9 - 11 mm, pale pink, back of petals pilose; petals connate 2 / 5 to 1 / 2, corolla lobes 5 - 6 mm, ovate, apiculate, as long as or slightly longer than corolla tube. Flowering in July. Limestone rocks 1700 - 2800 m.	en	Kaya, Ömer Faruk, Ertekin, Alaattin Selçuk (2025): An emended description for Rosularia haussknechtii (Boiss. & Reuter) A. Berger (Crassulaceae) from Southeastern Anatolia. Adansonia (3) 47 (18): 319-326, DOI: 10.5252/adansonia2025v47a18, URL: https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/adansonia2025v47a18.pdf
