taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912029	https://doi.org/10.3897/zookeys.1180.107333.figure1	Figure 1. Evolutionary relationships amongst species in the Pristimantis gladiator complex, under the Maximum Likelihood inference criterion (modified from Franco-Mena et al. (2023)). Red font indicates the new sequence.	Figure 1. Evolutionary relationships amongst species in the Pristimantis gladiator complex, under the Maximum Likelihood inference criterion (modified from Franco-Mena et al. (2023)). Red font indicates the new sequence.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912030	https://doi.org/10.3897/zookeys.1180.107333.figure2	Figure 2. Pristimantis donnelsoni sp. nov. in ethanol. Adult female, holotype, DHMECN 14701, in dorsal, ventral and lateral views. Photographs by JPRP.	Figure 2. Pristimantis donnelsoni sp. nov. in ethanol. Adult female, holotype, DHMECN 14701, in dorsal, ventral and lateral views. Photographs by JPRP.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912031	https://doi.org/10.3897/zookeys.1180.107333.figure3	Figure 3. Ventral view of left hand and foot of Pristimantis donnelsoni sp. nov. Adult female, holotype, DHMECN 14701. Photographs by JPRP.	Figure 3. Ventral view of left hand and foot of Pristimantis donnelsoni sp. nov. Adult female, holotype, DHMECN 14701. Photographs by JPRP.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912032	https://doi.org/10.3897/zookeys.1180.107333.figure4	Figure 4. Lateral views of Pristimantis donnelsoni sp. nov. in life. Adult female, holotype DHMECN 14701. LRC: 15.42 mm. Photographs by Frank Pichardo.	Figure 4. Lateral views of Pristimantis donnelsoni sp. nov. in life. Adult female, holotype DHMECN 14701. LRC: 15.42 mm. Photographs by Frank Pichardo.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912033	https://doi.org/10.3897/zookeys.1180.107333.figure5	Figure 5. Dorsolateral and ventral views in life of Pristimantis gladiator species complex and related species A Pristimantis donnelsoni sp. nov. female, DHMECN 14701, holotype, LRC: 15.42 mm B Pristimantis donnelsoni sp. nov. male, DHMECN 18854, LRC: 13.66 mm from Cerro Candelaria C Pristimantis kayi sp. nov. female, ZSFQ 0775, holotype, LRC: 19.7 mm D Pristimantis kayi sp. nov. male, DHMECN 14450 LRC: 13.58 mm, from Reserva Machay E Pristimantis gladiator sensu stricto female, DHMECN 12463, LRC: 20.38 mm F Pristimantis gladiator sensu stricto, male DHMECN 12474, LRC: 16.87 mm; from Rio Quijos drainage G Pristimantis festae female, DHMECN 16280, LRC: 20.19 mm H Pristimantis festae, male (left), DHMECN 16837, LRC: 16.50 mm; female (right), DHMECN 16836, LRC: 19.80 mm; Photographers: JPRP (A, B), MYM (D, E, F, H); Patricia Bejarano (G); Jose Vieira (C).	Figure 5. Dorsolateral and ventral views in life of Pristimantis gladiator species complex and related species A Pristimantis donnelsoni sp. nov. female, DHMECN 14701, holotype, LRC: 15.42 mm B Pristimantis donnelsoni sp. nov. male, DHMECN 18854, LRC: 13.66 mm from Cerro Candelaria C Pristimantis kayi sp. nov. female, ZSFQ 0775, holotype, LRC: 19.7 mm D Pristimantis kayi sp. nov. male, DHMECN 14450 LRC: 13.58 mm, from Reserva Machay E Pristimantis gladiator sensu stricto female, DHMECN 12463, LRC: 20.38 mm F Pristimantis gladiator sensu stricto, male DHMECN 12474, LRC: 16.87 mm; from Rio Quijos drainage G Pristimantis festae female, DHMECN 16280, LRC: 20.19 mm H Pristimantis festae, male (left), DHMECN 16837, LRC: 16.50 mm; female (right), DHMECN 16836, LRC: 19.80 mm; Photographers: JPRP (A, B), MYM (D, E, F, H); Patricia Bejarano (G); Jose Vieira (C).	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912034	https://doi.org/10.3897/zookeys.1180.107333.figure6	Figure 6. Pristimantis donnelsoni sp. nov. dorsal and ventral views of the preserved types series A DHMECN 4779, female B DHMECN 4782, female C female, ZSFQ 1097 D holotype, 14701, female E DHMECN 4781, male F DHMECN 16613, male G DHMECN 16610, male H ZSFQ 1098, male I DHMECN 4778, male. Photographs by JPRP and David Brito Zapata.	Figure 6. Pristimantis donnelsoni sp. nov. dorsal and ventral views of the preserved types series A DHMECN 4779, female B DHMECN 4782, female C female, ZSFQ 1097 D holotype, 14701, female E DHMECN 4781, male F DHMECN 16613, male G DHMECN 16610, male H ZSFQ 1098, male I DHMECN 4778, male. Photographs by JPRP and David Brito Zapata.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912035	https://doi.org/10.3897/zookeys.1180.107333.figure7	Figure 7. Life dorsal and ventral views of Pristimantis donnelsoni sp. nov., in situ A, B holotype male DHMECN 14701, LRC: 15.42 mm, from Chamana Reserve C, D paratype female DHMECN 13321, LRC: 15.47 mm, from Cerro Candelaria E, F paratype female DHMECN 16175, LRC: 13.71 mm, from Cerro Candelaria G, H paratype female DHMECN 18159, LRC: 18.72 mm, from Hacienda Guamag I, J adult male DHMECN 18185, LRC: 14.26 mm, from Finca Palmonte. Photographs by JPRP.	Figure 7. Life dorsal and ventral views of Pristimantis donnelsoni sp. nov., in situ A, B holotype male DHMECN 14701, LRC: 15.42 mm, from Chamana Reserve C, D paratype female DHMECN 13321, LRC: 15.47 mm, from Cerro Candelaria E, F paratype female DHMECN 16175, LRC: 13.71 mm, from Cerro Candelaria G, H paratype female DHMECN 18159, LRC: 18.72 mm, from Hacienda Guamag I, J adult male DHMECN 18185, LRC: 14.26 mm, from Finca Palmonte. Photographs by JPRP.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912036	https://doi.org/10.3897/zookeys.1180.107333.figure8	Figure 8. Skull morphology diagram in dorsal (left column) and ventral (right column) views in four species of the Pristimantis myersi species group: A, B P. donnelsoni sp. nov., DHMECN 4808 C, D P. kayi sp. nov., DHMECN 14447 E, F P. gladiator DHMECN 12465 G, H P. festae DHMECN 9294. Detail of the skull bones abbreviations: pmax = premaxilla; max = maxilla; spheth = sphenethmoides; fpar = frontoparietal; pter = pterigoides; pro = prootic; q = quadratojugal; sp = squamosal; exo = exoccipital; prph = parasphenoides. Frontoparietal fontanelle not coloured.	Figure 8. Skull morphology diagram in dorsal (left column) and ventral (right column) views in four species of the Pristimantis myersi species group: A, B P. donnelsoni sp. nov., DHMECN 4808 C, D P. kayi sp. nov., DHMECN 14447 E, F P. gladiator DHMECN 12465 G, H P. festae DHMECN 9294. Detail of the skull bones abbreviations: pmax = premaxilla; max = maxilla; spheth = sphenethmoides; fpar = frontoparietal; pter = pterigoides; pro = prootic; q = quadratojugal; sp = squamosal; exo = exoccipital; prph = parasphenoides. Frontoparietal fontanelle not coloured.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912037	https://doi.org/10.3897/zookeys.1180.107333.figure9	Figure 9. Boxplot of body size (SVL) and tympanic diameter (TD) in four closely-related species of the Pristimantis myersi species group. * Asterisks represent species with significant differences.	Figure 9. Boxplot of body size (SVL) and tympanic diameter (TD) in four closely-related species of the Pristimantis myersi species group. * Asterisks represent species with significant differences.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912038	https://doi.org/10.3897/zookeys.1180.107333.figure10	Figure 10. Oscillogram and spectrogram of the advertisement call of Pristimantis donnelsoni sp. nov., adult male, paratype DHMECN 4774.	Figure 10. Oscillogram and spectrogram of the advertisement call of Pristimantis donnelsoni sp. nov., adult male, paratype DHMECN 4774.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912039	https://doi.org/10.3897/zookeys.1180.107333.figure11	Figure 11. Distribution of four closely-related species of the Pristimantis myersi species group in the Andes of Ecuador.	Figure 11. Distribution of four closely-related species of the Pristimantis myersi species group in the Andes of Ecuador.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
85F826E58E0F5EDABF3AA1FF49A9BDDE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912040	https://doi.org/10.3897/zookeys.1180.107333.figure12	Figure 12. Habitat vegetation at type localities and etymology of the two new frogs described herein. Photographs by JPRP and Zane Libke A panoramic view of the upper Pastaza Valley and its adjacent mountains, main geographic barriers promoting speciation at a regional scale, at back Tungurahua Volcano and mountains south of the Pastaza River habitat of Pristimantis donnelsoni sp. nov., image from the slopes of Cerro Mayordomo type locality of Pristimantis kayi sp. nov., in the northern mountains to the Pastaza River B Tungurahua Volcano eruption, from Chamana Reserve C Montane forest in Candelaria Reserve D Don Nelson Palacios (+) during an expedition; we tribute his work with the naming of Pristimantis donnelsoni sp. nov. E Sumaco Volcano seen from Wawa Sumaku, habitat of P. kayi sp. nov. F Montane forest and subparamo in Leito, Patate G Andreas Kay (+); we honour his devotion to nature photography and research in the upper Pastaza Valley and through Ecuador by naming Pristimantis kayi sp. nov.	Figure 12. Habitat vegetation at type localities and etymology of the two new frogs described herein. Photographs by JPRP and Zane Libke A panoramic view of the upper Pastaza Valley and its adjacent mountains, main geographic barriers promoting speciation at a regional scale, at back Tungurahua Volcano and mountains south of the Pastaza River habitat of Pristimantis donnelsoni sp. nov., image from the slopes of Cerro Mayordomo type locality of Pristimantis kayi sp. nov., in the northern mountains to the Pastaza River B Tungurahua Volcano eruption, from Chamana Reserve C Montane forest in Candelaria Reserve D Don Nelson Palacios (+) during an expedition; we tribute his work with the naming of Pristimantis donnelsoni sp. nov. E Sumaco Volcano seen from Wawa Sumaku, habitat of P. kayi sp. nov. F Montane forest and subparamo in Leito, Patate G Andreas Kay (+); we honour his devotion to nature photography and research in the upper Pastaza Valley and through Ecuador by naming Pristimantis kayi sp. nov.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912029	https://doi.org/10.3897/zookeys.1180.107333.figure1	Figure 1. Evolutionary relationships amongst species in the Pristimantis gladiator complex, under the Maximum Likelihood inference criterion (modified from Franco-Mena et al. (2023)). Red font indicates the new sequence.	Figure 1. Evolutionary relationships amongst species in the Pristimantis gladiator complex, under the Maximum Likelihood inference criterion (modified from Franco-Mena et al. (2023)). Red font indicates the new sequence.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912033	https://doi.org/10.3897/zookeys.1180.107333.figure5	Figure 5. Dorsolateral and ventral views in life of Pristimantis gladiator species complex and related species A Pristimantis donnelsoni sp. nov. female, DHMECN 14701, holotype, LRC: 15.42 mm B Pristimantis donnelsoni sp. nov. male, DHMECN 18854, LRC: 13.66 mm from Cerro Candelaria C Pristimantis kayi sp. nov. female, ZSFQ 0775, holotype, LRC: 19.7 mm D Pristimantis kayi sp. nov. male, DHMECN 14450 LRC: 13.58 mm, from Reserva Machay E Pristimantis gladiator sensu stricto female, DHMECN 12463, LRC: 20.38 mm F Pristimantis gladiator sensu stricto, male DHMECN 12474, LRC: 16.87 mm; from Rio Quijos drainage G Pristimantis festae female, DHMECN 16280, LRC: 20.19 mm H Pristimantis festae, male (left), DHMECN 16837, LRC: 16.50 mm; female (right), DHMECN 16836, LRC: 19.80 mm; Photographers: JPRP (A, B), MYM (D, E, F, H); Patricia Bejarano (G); Jose Vieira (C).	Figure 5. Dorsolateral and ventral views in life of Pristimantis gladiator species complex and related species A Pristimantis donnelsoni sp. nov. female, DHMECN 14701, holotype, LRC: 15.42 mm B Pristimantis donnelsoni sp. nov. male, DHMECN 18854, LRC: 13.66 mm from Cerro Candelaria C Pristimantis kayi sp. nov. female, ZSFQ 0775, holotype, LRC: 19.7 mm D Pristimantis kayi sp. nov. male, DHMECN 14450 LRC: 13.58 mm, from Reserva Machay E Pristimantis gladiator sensu stricto female, DHMECN 12463, LRC: 20.38 mm F Pristimantis gladiator sensu stricto, male DHMECN 12474, LRC: 16.87 mm; from Rio Quijos drainage G Pristimantis festae female, DHMECN 16280, LRC: 20.19 mm H Pristimantis festae, male (left), DHMECN 16837, LRC: 16.50 mm; female (right), DHMECN 16836, LRC: 19.80 mm; Photographers: JPRP (A, B), MYM (D, E, F, H); Patricia Bejarano (G); Jose Vieira (C).	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912036	https://doi.org/10.3897/zookeys.1180.107333.figure8	Figure 8. Skull morphology diagram in dorsal (left column) and ventral (right column) views in four species of the Pristimantis myersi species group: A, B P. donnelsoni sp. nov., DHMECN 4808 C, D P. kayi sp. nov., DHMECN 14447 E, F P. gladiator DHMECN 12465 G, H P. festae DHMECN 9294. Detail of the skull bones abbreviations: pmax = premaxilla; max = maxilla; spheth = sphenethmoides; fpar = frontoparietal; pter = pterigoides; pro = prootic; q = quadratojugal; sp = squamosal; exo = exoccipital; prph = parasphenoides. Frontoparietal fontanelle not coloured.	Figure 8. Skull morphology diagram in dorsal (left column) and ventral (right column) views in four species of the Pristimantis myersi species group: A, B P. donnelsoni sp. nov., DHMECN 4808 C, D P. kayi sp. nov., DHMECN 14447 E, F P. gladiator DHMECN 12465 G, H P. festae DHMECN 9294. Detail of the skull bones abbreviations: pmax = premaxilla; max = maxilla; spheth = sphenethmoides; fpar = frontoparietal; pter = pterigoides; pro = prootic; q = quadratojugal; sp = squamosal; exo = exoccipital; prph = parasphenoides. Frontoparietal fontanelle not coloured.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912040	https://doi.org/10.3897/zookeys.1180.107333.figure12	Figure 12. Habitat vegetation at type localities and etymology of the two new frogs described herein. Photographs by JPRP and Zane Libke A panoramic view of the upper Pastaza Valley and its adjacent mountains, main geographic barriers promoting speciation at a regional scale, at back Tungurahua Volcano and mountains south of the Pastaza River habitat of Pristimantis donnelsoni sp. nov., image from the slopes of Cerro Mayordomo type locality of Pristimantis kayi sp. nov., in the northern mountains to the Pastaza River B Tungurahua Volcano eruption, from Chamana Reserve C Montane forest in Candelaria Reserve D Don Nelson Palacios (+) during an expedition; we tribute his work with the naming of Pristimantis donnelsoni sp. nov. E Sumaco Volcano seen from Wawa Sumaku, habitat of P. kayi sp. nov. F Montane forest and subparamo in Leito, Patate G Andreas Kay (+); we honour his devotion to nature photography and research in the upper Pastaza Valley and through Ecuador by naming Pristimantis kayi sp. nov.	Figure 12. Habitat vegetation at type localities and etymology of the two new frogs described herein. Photographs by JPRP and Zane Libke A panoramic view of the upper Pastaza Valley and its adjacent mountains, main geographic barriers promoting speciation at a regional scale, at back Tungurahua Volcano and mountains south of the Pastaza River habitat of Pristimantis donnelsoni sp. nov., image from the slopes of Cerro Mayordomo type locality of Pristimantis kayi sp. nov., in the northern mountains to the Pastaza River B Tungurahua Volcano eruption, from Chamana Reserve C Montane forest in Candelaria Reserve D Don Nelson Palacios (+) during an expedition; we tribute his work with the naming of Pristimantis donnelsoni sp. nov. E Sumaco Volcano seen from Wawa Sumaku, habitat of P. kayi sp. nov. F Montane forest and subparamo in Leito, Patate G Andreas Kay (+); we honour his devotion to nature photography and research in the upper Pastaza Valley and through Ecuador by naming Pristimantis kayi sp. nov.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912041	https://doi.org/10.3897/zookeys.1180.107333.figure13	Figure 13. Holotype of Pristimantis kayi sp. nov. in life, adult female, ZSFQ 0775. LRC: 19.7 mm. Photographs by Jose Vieira.	Figure 13. Holotype of Pristimantis kayi sp. nov. in life, adult female, ZSFQ 0775. LRC: 19.7 mm. Photographs by Jose Vieira.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912042	https://doi.org/10.3897/zookeys.1180.107333.figure14	Figure 14. Colour variation of Pristimantis kayi sp. nov. A MZUTI 2209, adult female, LRC: 20.15 mm, from Cordillera de Guacamayos B MZUTI 2210, adult female, LRC: 19.90 mm, from Cordillera de Guacamayos C MZUTI 2211, adult male, LRC: 14.30 mm, from Cordillera de Guacamayos D MZUTI 2213, adult male, LRC: 15.10 mm, from Cordillera de Guacamayos E MZUTI 2214, adult female, LRC: 18.73 mm, from Cordillera de Guacamayos F ZSFQ 774, adult female, LRC: 18.96 mm from Sumaco Volcano G ZSFQ 782, adult female, LRC: 30.03 mm, from Sumaco Volcano. Photographs by Lucas Bustamante (A, B, C, D, E) and Jose Vieira (F, G).	Figure 14. Colour variation of Pristimantis kayi sp. nov. A MZUTI 2209, adult female, LRC: 20.15 mm, from Cordillera de Guacamayos B MZUTI 2210, adult female, LRC: 19.90 mm, from Cordillera de Guacamayos C MZUTI 2211, adult male, LRC: 14.30 mm, from Cordillera de Guacamayos D MZUTI 2213, adult male, LRC: 15.10 mm, from Cordillera de Guacamayos E MZUTI 2214, adult female, LRC: 18.73 mm, from Cordillera de Guacamayos F ZSFQ 774, adult female, LRC: 18.96 mm from Sumaco Volcano G ZSFQ 782, adult female, LRC: 30.03 mm, from Sumaco Volcano. Photographs by Lucas Bustamante (A, B, C, D, E) and Jose Vieira (F, G).	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912043	https://doi.org/10.3897/zookeys.1180.107333.figure15	Figure 15. Preserved holotype of Pristimantis kayi sp. nov. ZSFQ 0775, on its dorsal, ventral and lateral views. Photographs by David Brito-Zapata.	Figure 15. Preserved holotype of Pristimantis kayi sp. nov. ZSFQ 0775, on its dorsal, ventral and lateral views. Photographs by David Brito-Zapata.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912044	https://doi.org/10.3897/zookeys.1180.107333.figure16	Figure 16. Detailed ventral view of hand and feet of the holotype ZSFQ 0775 of Pristimantis kayi sp. nov. Photographs David Brito-Zapata.	Figure 16. Detailed ventral view of hand and feet of the holotype ZSFQ 0775 of Pristimantis kayi sp. nov. Photographs David Brito-Zapata.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912045	https://doi.org/10.3897/zookeys.1180.107333.figure17	Figure 17. Dorsal and ventral views of preserved type series of Pristimantis kayi sp. nov. A holotype ZSFQ 0775, female; Paratypes B ZSFQ 0779, female C DHMECN 16212, female D MZUTI 5162, female E MZUTI 4657, male F ZSFQ 0773, male G DHMECN 15233, male H DHMECN 18440, male; Photographs by JPRPR and David Brito-Zapata.	Figure 17. Dorsal and ventral views of preserved type series of Pristimantis kayi sp. nov. A holotype ZSFQ 0775, female; Paratypes B ZSFQ 0779, female C DHMECN 16212, female D MZUTI 5162, female E MZUTI 4657, male F ZSFQ 0773, male G DHMECN 15233, male H DHMECN 18440, male; Photographs by JPRPR and David Brito-Zapata.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912034	https://doi.org/10.3897/zookeys.1180.107333.figure6	Figure 6. Pristimantis donnelsoni sp. nov. dorsal and ventral views of the preserved types series A DHMECN 4779, female B DHMECN 4782, female C female, ZSFQ 1097 D holotype, 14701, female E DHMECN 4781, male F DHMECN 16613, male G DHMECN 16610, male H ZSFQ 1098, male I DHMECN 4778, male. Photographs by JPRP and David Brito Zapata.	Figure 6. Pristimantis donnelsoni sp. nov. dorsal and ventral views of the preserved types series A DHMECN 4779, female B DHMECN 4782, female C female, ZSFQ 1097 D holotype, 14701, female E DHMECN 4781, male F DHMECN 16613, male G DHMECN 16610, male H ZSFQ 1098, male I DHMECN 4778, male. Photographs by JPRP and David Brito Zapata.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912037	https://doi.org/10.3897/zookeys.1180.107333.figure9	Figure 9. Boxplot of body size (SVL) and tympanic diameter (TD) in four closely-related species of the Pristimantis myersi species group. * Asterisks represent species with significant differences.	Figure 9. Boxplot of body size (SVL) and tympanic diameter (TD) in four closely-related species of the Pristimantis myersi species group. * Asterisks represent species with significant differences.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912046	https://doi.org/10.3897/zookeys.1180.107333.figure18	Figure 18. Call of Pristimantis kayi sp. nov. (adult male, MZUTI 852). The species has different types of calls, represented as follows: A single note call B pulsed note calls C complex calls (non-pulsed and pulsed notes).	Figure 18. Call of Pristimantis kayi sp. nov. (adult male, MZUTI 852). The species has different types of calls, represented as follows: A single note call B pulsed note calls C complex calls (non-pulsed and pulsed notes).	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
10218311988457C8AE068B321C63C959.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/912039	https://doi.org/10.3897/zookeys.1180.107333.figure11	Figure 11. Distribution of four closely-related species of the Pristimantis myersi species group in the Andes of Ecuador.	Figure 11. Distribution of four closely-related species of the Pristimantis myersi species group in the Andes of Ecuador.	2023-09-26	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina		Zenodo	biologists	Reyes-Puig, Juan Pablo;Urgiles-Merchan, Miguel;Franco-Mena, Daniela;Guayasamin, Juan M.;Batallas, Diego;Reyes-Puig, Carolina			
