taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
E4F09F24B2135BA9942025793699F768.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441485	https://doi.org/10.3897/zse.101.160106.figure6	Figure 6. The general view of Biwia zezera, the type species of the genus Biwia, KUN-P 040282, male, 53.8 mm SL, collected from Lake Biwa, Moriyama, Shiga, Japan. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Seigo Kawase.	Figure 6. The general view of Biwia zezera, the type species of the genus Biwia, KUN-P 040282, male, 53.8 mm SL, collected from Lake Biwa, Moriyama, Shiga, Japan. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Seigo Kawase.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
E4F09F24B2135BA9942025793699F768.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
E4F09F24B2135BA9942025793699F768.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
E4F09F24B2135BA9942025793699F768.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441481	https://doi.org/10.3897/zse.101.160106.figure2	Figure 2. The phylogenetic relationship of seven genera within the Biwia – Microphysogobio complex. A. Bayesian inference (BI) tree, posterior probabilities shown at nodes; B. maximum likelihood (ML) tree, bootstrap values shown at nodes.	Figure 2. The phylogenetic relationship of seven genera within the Biwia – Microphysogobio complex. A. Bayesian inference (BI) tree, posterior probabilities shown at nodes; B. maximum likelihood (ML) tree, bootstrap values shown at nodes.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
61412E2A90F55ACDBF771F5A23FBE182.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441491	https://doi.org/10.3897/zse.101.160106.figure12	Figure 12. The general view of Crossocheilogobio tafangensis, the type species of the genus Crossocheilogobio gen. nov., ASIZB 60212, male, 91.2 mm SL, collected from the Xin’anjiang River in the Qiantangjiang River Basin, Tunxi District, Anhui Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 12. The general view of Crossocheilogobio tafangensis, the type species of the genus Crossocheilogobio gen. nov., ASIZB 60212, male, 91.2 mm SL, collected from the Xin’anjiang River in the Qiantangjiang River Basin, Tunxi District, Anhui Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
61412E2A90F55ACDBF771F5A23FBE182.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
61412E2A90F55ACDBF771F5A23FBE182.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
38B58A57926D5CCA81184DD24968169E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441487	https://doi.org/10.3897/zse.101.160106.figure8	Figure 8. The general view of Huigobio chenhsienensis, the type species of the genus Huigobio, SHOU 20231209015, 60.4 mm SL, collected from a tributary of the Chengtanjiang River in the Cao’ejiang River system, Shengzhou City, Zhejiang Province, China. A. lateral view; B. dorsal view; C. ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 8. The general view of Huigobio chenhsienensis, the type species of the genus Huigobio, SHOU 20231209015, 60.4 mm SL, collected from a tributary of the Chengtanjiang River in the Cao’ejiang River system, Shengzhou City, Zhejiang Province, China. A. lateral view; B. dorsal view; C. ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
38B58A57926D5CCA81184DD24968169E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
38B58A57926D5CCA81184DD24968169E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
9601FD1185D255AA9A82E73EBA0A88BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441488	https://doi.org/10.3897/zse.101.160106.figure9	Figure 9. The general view of Mesophysogobio kachekensis, the type species of the genus Mesophysogobio gen. nov., ASIZB 69768, male, 93.2 mm SL, collected from a tributary of the Dasijiang River in the Maolingjiang River system, Shangsi County, Guangxi Zhuang Autonomous Region, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 9. The general view of Mesophysogobio kachekensis, the type species of the genus Mesophysogobio gen. nov., ASIZB 69768, male, 93.2 mm SL, collected from a tributary of the Dasijiang River in the Maolingjiang River system, Shangsi County, Guangxi Zhuang Autonomous Region, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
9601FD1185D255AA9A82E73EBA0A88BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
9601FD1185D255AA9A82E73EBA0A88BB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
1E235459835A51C788C516A008214F4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441486	https://doi.org/10.3897/zse.101.160106.figure7	Figure 7. The general view of Microphysogobio hsinglungshanensis, the type species of the genus Microphysogobio, ASIZB 240788, 51.7 mm SL, collected from the Liuhe River in the Luanhe River Basin, Xinglong County, Hebei Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 7. The general view of Microphysogobio hsinglungshanensis, the type species of the genus Microphysogobio, ASIZB 240788, 51.7 mm SL, collected from the Liuhe River in the Luanhe River Basin, Xinglong County, Hebei Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
1E235459835A51C788C516A008214F4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
1E235459835A51C788C516A008214F4E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
C5CE45A8E8EF504B98F6A25DE867FFF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441490	https://doi.org/10.3897/zse.101.160106.figure11	Figure 11. The general view of Oriengobio fukiensis, the type species of the genus Oriengobio gen. nov., ASIZB 220659, 54.9 mm SL, collected from the Mayangxi River in the Minjiang River Basin, Jianyang District, Fujian Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 11. The general view of Oriengobio fukiensis, the type species of the genus Oriengobio gen. nov., ASIZB 220659, 54.9 mm SL, collected from the Mayangxi River in the Minjiang River Basin, Jianyang District, Fujian Province, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
C5CE45A8E8EF504B98F6A25DE867FFF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
C5CE45A8E8EF504B98F6A25DE867FFF2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
F6D812B9E99351F691550E1942DE7766.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441489	https://doi.org/10.3897/zse.101.160106.figure10	Figure 10. The general view of Platysmacheilus exiguus, the type species of the genus Platysmacheilus, ASIZB 220879, 77.6 mm SL, collected from the Lijiang River in the Pearl River Basin, Pingle County, Guangxi Zhuang Autonomous Region, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	Figure 10. The general view of Platysmacheilus exiguus, the type species of the genus Platysmacheilus, ASIZB 220879, 77.6 mm SL, collected from the Lijiang River in the Pearl River Basin, Pingle County, Guangxi Zhuang Autonomous Region, China. A. Lateral view; B. Dorsal view; C. Ventral view. Scale bar: 10 mm. Photographed by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
F6D812B9E99351F691550E1942DE7766.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441482	https://doi.org/10.3897/zse.101.160106.figure3	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	Figure 3. Lip papillae patterns of the type species of seven genera within the Biwia – Microphysogobio complex, with a diagrammatic illustration showing the structures. A. Biwia zezera, KUN-P 040282; B. Microphysogobio hsinglungshanensis, ASIZB 240788; C. Huigobio chenhsienensis, SHOU 20231209015; D. Oriengobio fukiensis, ASIZB 220659; E. Mesophysogobio kachekensis, ASIZB 69768; F. Platysmacheilus exiguus, ASIZB 220879; G. Crossocheilogobio tafangensis, ASIZB 60212; H. diagrammatic illustration based on Microphysogobio chinssuensis. CPAP = central portion of anterior papillae; LPAP = lateral portion of anterior papillae; MP = medial pad; LL = lateral lobe; BB = barbel; UJ = upper jaw; LJ = lower jaw. The photo of Biwia zezera is provided by Seigo Kawase; the other photos and illustration were made by Zhi-Xian Sun.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
F6D812B9E99351F691550E1942DE7766.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1441484	https://doi.org/10.3897/zse.101.160106.figure5	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	Figure 5. The distributions of the Biwia – Microphysogobio complex and seven genera. A. The complex; B. Biwia; C. Microphysogobio; D. Huigobio; E. Platysmacheilus; F. Crossocheilogobio gen. nov.; G. Oriengobio gen. nov.; H. Mesophysogobio gen. nov.	2025-10-16	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui		Zenodo	biologists	Sun, Zhi-Xian;Tang, Wen-Qiao;Zhao, Ya-Hui			
