taxonID	type	description	language	source
366BA2148359BE0707BBFD77C1ABF97E.taxon	type_taxon	type species: Pintia boholica Semper, 1898, by original designation.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148359BE0707BBFD77C1ABF97E.taxon	diagnosis	Diagnosis. Members of the genus are small-sized moth with black ground color and yellow stripes of different shapes on wings. Superficially, the genus can be easily distinguished from Cryptophysophilus and Heterusinula in the same clade, both only known from females, by the less-developed yellow patterns on forewing (yellow patterns occupy half of the forewing in Cryptophysophilus and Heterusinula) in female. In female genitalia Docleomorpha is closer to Cryptophysophilus by sharing the presence of strong sclerotisation in antrum (antrum only weakly sclerotised in Heterusinula), and it can be distinguished from Cryptophysophilus by the much shorter sclerotised section on antrum. The male genitalia of Docleomorpha are similar to those of the genus Corma Walker, 1865 (type species Corma obscurata Walker, 1865, by subsequent designation by Kirby (1892 )) but can be distinguished by the presence of juxta and the free distal saccular process (fused with valvula in Corma).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148359BE0707BBFD77C1ABF97E.taxon	distribution	Distribution. The Philippines and Malaysia (Borneo) Checklist of the genus Docleomorpha The D. boholica species-group D. boholica (Semper, 1898) D. epsilon Huang & Horie sp. nov. D. zeta Huang & Horie sp. nov. The D. violae species-group D. violae Huang & Horie sp. nov. The D. zhaoi species-group D. zhaoi Huang & Horie sp. nov.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148359BE0407BBF94CC199FDAF.taxon	diagnosis	Diagnosis. Docleomorpha boholica species-group can be distinguished from violae and zhaoi species-groups by the well-developed, band-like medial stripe on hindwing, the presence of lamella centralis (sensu Volynkin (2024 )), see note below) and the elongated sacculus with its distal section covered with spike-like spines in male genitalia. Notes. The presence of a long triangular sclerite in the valvula (Figs 13, 15 & 16) in D. boholica species-group is unique among Chalcosiinae, and currently no term is available for it in the terminology of Chalcosiinae. Therefore we temporarily adopted the term “ lamella centralis ” from the terminology of Lithosiinae (sensu Volynkin (2024 )) to name it, due to the similar articulation relationship between this sclerite and the distal section of sacculus. However, typical lamella centralis (i. e. figs 14 & 16 in Volynkin (2024 )) usually arises from costa, while the sclerite in Docleomorpha arises from the base of valva. Hence the homology of this structure in Docleomorpha requires further studies. Species included	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148359BE0407BBF94CC199FDAF.taxon	description	The D. boholica species-group	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148359BE0407BBF94CC199FDAF.taxon	description	D. zeta Huang & Horie sp. nov.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0407BBFD38C309FAEB.taxon	description	Figs 1 – 3, 13.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0407BBFD38C309FAEB.taxon	materials_examined	Material examined. 2 males, VIII. 2005, Mt. Balocawe, C Leyte, The Philippines, leg. Danny, ex. coll. C. G. Treadaway, gen. prep. No. SNGHSY 02 (SMF); 1 male, IV. 2011, Mt. Kadohaan, C Leyte, The Philippines (CKH); 1 male, III-V. 2014, Mt. Balocawe, C Leyte, The Philippines, leg. Noel Mohagan (CKH); 1 male, IV. 2011, Mt. Balocawe, C Leyte, The Philippines, leg. Noel Mohagan (CKH).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0407BBFD38C309FAEB.taxon	diagnosis	Diagnosis. Length of forewing 15 – 17 mm in male. In male, Docleomorpha boholica can be distinguished from both D. epsilon sp. nov. and D. zeta sp. nov. superficially by the larger size, the shorter and broader yellow band on forewing and the broader medial stripe on hindwing. In male genitalia, D. boholica can be distinguished from the two congeners in the species-group by the longer uncus, the longer posterior bilateral projections, the semioval distal process of sacculus (ε-shaped in D. epsilon sp. nov. and ζ-shaped in D. zeta sp. nov.) and the longer phallus. The female is unknown.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0407BBFD38C309FAEB.taxon	distribution	Distribution. The Philippines (Bohol and Leyte Islands).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	description	Figs 4 – 7, 14, 15 & 20.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	materials_examined	Material examined. Holotype. male, “ Cyclosia | Boholica | Bohol. Philippines Semp. ” / “ Joicey | Bequest. | Brit. Mus. | 1934 - 120 ” / “ Mindanao | Philippines | I. I. Mounsay ” / QR-code label with unique number “ NHMUK 015106663 ”, gen. slide No. NHMUK 014331807 (NHMUK). Paratypes. 1 male, 3 females, 2011, Mt. Balocawe, C Leyte, The Philippines, genitalia preparation No. KH 54 (female) and KH 55 (male) (CKH); 1 male, IV. 2012, Mt. Halcon, Mindoro, The Philippines (CKH); 1 female, IV-VI. 2021, Mt. Halcon, Mindoro, The Philippines (CKH).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	diagnosis	Diagnosis. In male, Docleomorpha epsilon sp. nov. can be distinguished from D. boholica superficially by the smaller size, the slender and more obliquely placed yellow band on forewing and the narrower medial stripe on hindwing. In male genitalia, D. epsilon sp. nov. can be distinguished from D. boholica by the shorter and narrower uncus, the shorter posterior bilateral projections, the ε-shaped distal process of sacculus (semilunar-shaped in D. boholica) and the shorter phallus. The comparison with D. zeta sp. nov. is provided below in the diagnosis of that species.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	description	Description. External morphology of adults. Male. Length of forewing 12.5 – 14 mm. Antennae black with long rami. Head, thorax and abdomen black dorsally and yellow ventrally. Abdomen ringed with yellow between segments. Forewing ground color black with a rodlike yellow stripe extending from costa to vein CuA 2 in male and to vein CuP in female. Hindwing ground color black, in male the patterns consisted of a pale yellow stripe stretching along costal area and a yellow club-like stripe extending from wing base to submarginal zone medially. Male genitalia. Uncus nearly rectangular with its distal margin slightly concave, bearing setae along lateral and distal margins. Scaphium moderately sclerotised, band-like. Tegumen nearly trapezoid with straight margin, about twice the width of the uncus. Posterior bilateral projection blade-like with the bases of the projections connected medially by a band-like sclerite. Vinculum moderately broad, nearly the same width of the tegumenal lobe. Juxta M-shaped. Saccus distally rounded and broad U-shaped. Valva broad (ca. 3.5 × uncus length) and short (ca. 2 × uncus length), costa stripe-like, simple; valvula membranous and fan-shaped, covered sparsely with setae; sacculus band-like, elongated (ca. 2 × uncus length), with the outer margin covered with numerous spike-like spines from the medial to distal section and the distal section formed an ε-shaped process. Lamella centralis long triangular, extended from the base of valva to the distal section of sacculus. Phallus elongated, simple and cylindrical with coecum well-developed. Female. Length of forewing 15 – 15.5 mm. Antennae similar to male but with much shorter rami. Head black with metallic greenish blue scales dorsally, thorax and abdomen similar in male. Forewing apex less pointed than in male with similar yellow pattern, and bearing an additional metallic greenish blue patch at submarginal area. On hindwing the two stripes expanded, fused together and formed a large yellow patch, with the black pattern between reduced to a short stripe stretching from wing base to the medial section of discal cell. Female genitalia. Papillae anales long (ca. 0.5 × apophyses posteriores length). Apophyses posteriores slightly longer than apophyses anteriores, both heavily sclerotised. Ostium rounded. Antrum funnel-shaped, with the posterior section sclerotized and the anterior section somewhat membranous. Ductus bursae membranous. Corpus bursae membranous and globular.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	distribution	Distribution. The Philippines (Mindanao, Mindoro and Leyte Islands).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835ABE0507BBFAF4C362FC63.taxon	etymology	Etymology. The specific epithet epsilon is derived from the lower-case Greek letter “ ε ”, referring to the shape of the distal section of sacculus. It is a noun in apposition.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	description	Figs 8 – 10, 16 & 17	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	materials_examined	Material examined. Holotype. male, “ KH 56 ” / “ Philippines | Leyte Is. Mt. Balocaue | 2011 ”, genitalia preparation No. KH 56 (CKH, will be deposited in a public collection later). Paratypes. 1 male, 5. VI. 2002, Mt. Halcon, Mindoro, The Philippines, leg. native collector, genitalia preparation No. KH 57 (CKH); 1 male, VI. 2000, Mt. Halcon, N Mindoro, The Philippines, leg. N. Mohagan, genitalia preparation No. MfNHSY 023 (MfN).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	diagnosis	Diagnosis. Docleomorpha zeta sp. nov. can be distinguished from D. epsilon sp. nov. superficially by the shorter, somewhat broader and less obliquely placed yellow band on forewing. In male genitalia, it can be distinguished by the shorter uncus with medially concave lateral margins (lateral margins straight in D. epsilon sp. nov.) and the ζ-shaped distal process of sacculus (ε-shaped in D. epsilon sp. nov.). The comparison with D. boholica is provided above in the diagnosis of that species.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	description	Description. External morphology of adults. Length of forewing 12 – 13.5 mm in male. Antennae black, bipectinate. Head, thorax and abdomen black dorsally and yellow ventrally. Abdomen ringed with yellow between segments. Forewing ground color black with a rodlike yellow stripe extending from costa to vein CuP. Hindwing ground color black, the yellow patterns consisted of a pale yellow stripe stretching along costal area and a yellow club-like stripe extending from wing base to submarginal zone medially. Male genitalia. Uncus nearly rectangular, with its distal and lateral margins slightly concave, bearing setae along lateral and distal edges. Scaphium moderately sclerotised, band-like. Tegumen nearly trapezoid, about 1.5 × the width of the uncus. Posterior bilateral projection blade-like with the bases of the projections connected medially by a band-like sclerite. Vinculum moderately broad, nearly the same width of the tegumenal lobe. Juxta M-shaped. Saccus distally rounded and broad U-shaped. Valva broad (ca. 3.5 × uncus length) and short (ca. 2 × uncus length), costa stripe-like, simple; valvula membranous and fan-shaped, covered sparsely with setae; sacculus band-like, elongated (ca. 2 × uncus length), with the distal one third of the outer margin covered with numerous spike-like spines and the distal section formed a ζ-shaped process. Lamella centralis long triangular, extended from the base of valva to the distal section of sacculus. Phallus elongated, simple and cylindrical with coecum well-developed. Female unknown.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	distribution	Distribution. The Philippines (Mindoro and Leyte Islands).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835BBE0507BBFC67C340F811.taxon	etymology	Etymology. The specific epithet zeta is derived from the lower-case Greek letter “ ζ ”, referring to the shape of the distal section of sacculus. It is a noun in apposition.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFF35C1ACFE12.taxon	diagnosis	Diagnosis. Docleomorpha violae species-group can be distinguished from boholica and zhaoi species-groups by the broad, long dew-like medial stripe on hindwing and the beak-like distal saccular process in male genitalia. Species content The D. violae species-group	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFF35C1ACFE12.taxon	description	D. violae Huang & Horie sp. nov.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	description	Figs 11 & 18	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	materials_examined	Material examined. Holotype. male, “ Borneo Mt. Kinabalu | Kalanggaan, 1400 m | 9. - 13. VIII. 2005, LF | leg. W. Mey & K. Ebert. ” / “ MfNHSY 011 ” / “ ZFMK | LepDNA 0000479 ” (MfN).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	diagnosis	Diagnosis. Docleomorpha violae sp. nov. can be distinguished from its congeners superficially by the narrower forewing, the gradually narrowed yellow band on forewing and the well-developed medial stripe on hindwing. In male genitalia, D. violae sp. nov. can be distinguished from all the congeners by the smaller valvula, the ill-developed juxta, the absence of lamella centralis and the smooth, beak-like distal process of sacculus.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	description	Description. External morphology of adult. Length of forewing 13 mm in male. Antennae black, bipectinate. Head, thorax and abdomen black dorsally and yellow ventrally. Abdomen with yellow rings between segments. Forewing ground color black with a rodlike yellow stripe extending from costa to vein CuP, gradually narrowed from costa to the distal end. Hindwing ground color black, the yellow patterns consisted of a reduced pale yellow stripe stretching along the costal area, and a yellow, long dew-shaped stripe extending and broadening from wing base to submarginal zone medially. Male genitalia. Uncus rectangular with its distal margin concave medially, bearing setae along lateral and distal edges. Scaphium membranous. Tegumen trapezoid, about twice the width of the uncus. Posterior bilateral projection rodlike, with the tip of each lobe bent upwards and hooked and the bases of the projections connected medially by a short band-like sclerite. Vinculum moderately broad, nearly the same width of the tegumenal lobe. Juxta horseshoe-like and ill-developed, mostly fused with the inner margin of vinculum. Saccus broad U-shaped. Valva broad (ca. 2 × uncus length) and short (ca. 1.3 × uncus length), costa stripe-like, simple; valvula membranous and fan-shaped, with setae sparsely scattered; sacculus moderately broad (about 0.65 × uncus width) and short (ca. 1.4 × uncus length), with its distal one third formed a beak-like process. Phallus cylindrical, elongated and narrowed medially, with coecum well-developed. Female unknown.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	distribution	Distribution. Malaysia (Borneo).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	etymology	Etymology. The specific epithet violae is named after Ms Viola Richter (MfN, Berlin, Germany), who helped the first author greatly during his visits to the collections of MfN. It is a noun in the genitive case.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0007BBFDE3C1D7F992.taxon	discussion	Remarks. The juxta and sacculus of D. violae sp. nov. are very different from typical Docleomorpha, e. g. D. boholica species-group, as already described above, and its final placement requires the confirmation from molecular approaches.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0E07BBF960C1ABFEFA.taxon	diagnosis	Diagnosis. Docleomorpha zhaoi species-group can be distinguished from boholica and violae species-groups by the small sized adults, the reduced yellow patterns on both wings and the conical uncus, the well-developed, rounded juxta with blade-like dorsal process, and the presence of an oval sclerotized plate on valvula and a L-like basal saccular process in male genitalia. Species content The D. zhaoi species-group	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA214835EBE0E07BBF960C1ABFEFA.taxon	description	D. zhaoi Huang & Horie sp. nov.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	description	Figs 12 & 19	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	materials_examined	Material examined. Holotype. male, “ ZSY 256 | Kina Balu ” / “ Borneo: Sabah | near Keningau, Mt. | Trus Madi, 1180 m, | 18 – 23. IV. 2013 ” (CHSY, will be deposited in a public collection later).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	diagnosis	Diagnosis. Length of forewing 10 mm in male. Docleomorpha zhaoi sp. nov. can be distinguished from its congeners superficially by the small size and the reduced yellow patterns on both wings. In male genitalia, D. zhaoi sp. nov. can be easily distinguished from all the congeners by the conical uncus (rectangular in other congeners), the well-developed, rounded juxta with two dorsal blade-like processes, the presence of a weakly sclerotized plate above the ventral margin of valva and the sacculus with a L-like basal saccular process.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	description	Description. External morphology of adult. Length of forewing 10 mm in male. Antennae black, bipectinate. Head, thorax and abdomen black dorsally and yellow ventrally. Abdomen with yellow rings between segments. Forewing ground color black with a prominent yellow spot situated at costa and an obscure yellow stripe situated right after the yellow spot, strongly suffused with black. Hindwing ground color black, the yellow patterns consisted of a reduced pale yellow stripe stretching along the costal area and a yellow, trapezoid spot situated in postdiscal area with its basal section obscure and suffused with black. Male genitalia. Uncus conical, bearing setae along lateral margin. Scaphium membranous. Tegumen trapezoid, slightly broader than uncus. Posterior bilateral projection bayonet-like. Vinculum moderately broad, nearly the same width of the tegumenal lobe. Juxta rounded, plate-like and bearing two blade-like processes dorsally. Saccus broad U-shaped. Valva broad (ca. 5 × uncus length) and short (ca. 2.5 × uncus length), costa stripe-like, simple; valvula membranous and fan-shaped, with setae sparsely scattered; a weakly sclerotized oval plate presented above the ventral margin of valvula; sacculus short (about 1.75 × uncus length) with its basal section bearing a L-like process. Phallus cylindrical and narrowed towards its distal end, with coecum well-developed. Female is unknown.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	distribution	Distribution. Malaysia (Borneo).	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	etymology	Etymology. The specific epithet zhaoi is named after Mr Mingzhi Zhao (ZFMK, Bonn, Germany), who helped the first author acquire the specimen. It is a noun in the genitive case.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
366BA2148350BE0E07BBFECBC36DFA5E.taxon	discussion	Remarks. The placement of this species in the genus Docleomorpha is temporary. Although the wing patterns and shape resemble those of typical Docleomorpha, the details of juxta and valva are very different as already described above, and it is possible that this species should be transferred into a different genus. Nevertheless such assumption requires the evidence from molecular methods.	en	Huang, Si-Yao, Horie, Kiyoshi, Espeland, Marianne (2025): A review of the genus Docleomorpha Hering (Lepidoptera, Zygaenidae, Chalcosiinae) with descriptions of four new species. Zootaxa 5689 (1): 155-164, DOI: 10.11646/zootaxa.5689.1.6, URL: https://doi.org/10.11646/zootaxa.5689.1.6
