taxonID	type	description	language	source
A5C9A91FD80B548FA4C02A2E72748845.taxon	etymology	Etymology. Greek deilós (δειλός), timid, and Greek múkēs (μύκης), fungus, referring to the slow growth in culture and restricted production of ascomata.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
A5C9A91FD80B548FA4C02A2E72748845.taxon	type_taxon	Type species. Deilomyces minimus Guerra-Mateo, Gené & Cano	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
A5C9A91FD80B548FA4C02A2E72748845.taxon	description	Description. Mycelium composed of hyaline, septate, branched, smooth-walled hyphae. Asexual morph Chrysosporium - like. Conidia thallic, terminal, intercalary, occasionally lateral, hyaline to subhyaline, smooth-walled to tuberculate, thick-walled; terminal and lateral conidia aseptate, subglobose to obpyriform or short clavate with truncated base; intercalary conidia alternate, frequently disposed in a knuckle joint position, 0 – 1 - septate, subglobose, subcylindrical, obpyriform, or short clavate with truncated base. Sexual morph with gymnothecial ascomata. Gymnothecia superficial, single or aggregated, globose or subglobose; peridium composed of a subtle network of septate, branched, anastomosed, hyaline, smooth- and thick-walled hyphae; peridial appendages absent. Asci 8 - spored, evanescent, globose, subglobose, or pyriform. Ascospores unicellular, smooth- and thick-walled, globose, subglobose, ellipsoidal, or irregularly shaped.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
60309B22B24257AA96C5291380F88D23.taxon	description	Fig. 6	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
60309B22B24257AA96C5291380F88D23.taxon	etymology	Etymology. Latin minimus, small, referring to the small size of reproductive structures.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
60309B22B24257AA96C5291380F88D23.taxon	description	Description. Saprobic on marine sediments. Mycelium immersed and superficial, composed of hyaline, septate, branched, smooth-walled, 1.5 – 2 µm wide hyphae. Asexual morph Chrysosporium - like. Conidia thallic, terminal or intercalary, occasionally lateral on short stalks, hyaline to subhyaline, smooth-walled to tuberculate, thick-walled; terminal and lateral conidia on straight or slightly curved branches, aseptate, subglobose, obpyriform, or short clavate with truncated base, 3 – 4.5 × 2.5 – 3 µm; intercalary conidia alternate, frequently disposed in a knuckle joint position, 0 – (1) - septate, constricted at septum, subglobose, subcylindrical, obpyriform, or short clavate with truncated base, 5.5 – 8.5 (– 10.5) × 2.5 – 3 µm. Sexual morph with gymnothecial ascomata. Gymnothecia superficial, single or aggregated, initially white, turning pale yellow at maturity, globose or subglobose, 150 – 550 µm diam.; peridium composed of a subtle network of septate, branched, anastomosed, hyaline, smooth- and thick-walled, cylindrical, 1.5 – 2 µm wide hyphae; peridial appendages absent. Asci 8 - spored, evanescent, globose, subglobose, or pyriform, 4.5 – 6 × 4 – 5 µm. Ascospores unicellular, pale yellow, smooth- and thick-walled, globose, subglobose, ellipsoidal, or irregularly shaped, 2.5 × 1.5 – 2.5 µm.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
60309B22B24257AA96C5291380F88D23.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 13 – 16 mm diam., flat, predominantly submerged, aerial mycelium sparse, with scattered white (1 A 1) ascomata towards periphery, margin diffuse; reverse uncolored; producing a conspicuous urea odor. On PDA, 5 – 7 mm diam., slightly umbonate, floccose and white (1 A 1) at center, velvety and yellowish white (4 A 2) towards periphery, margin diffuse; reverse uncolored. On PYE, 3 – 4 mm diam., umbonate, woolly, white (1 A 1), margin feathery; reverse uncolored. Diffusible pigment not observed in any of the media studied.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
9AD9FE9B521F58D48C485522DAE3D345.taxon	description	Fig. 7	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
9AD9FE9B521F58D48C485522DAE3D345.taxon	etymology	Etymology. Referring to the white colonies with yellowish shades produced in culture.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
9AD9FE9B521F58D48C485522DAE3D345.taxon	description	Description. Saprobic on marine sediments. Mycelium superficial and immersed, composed of hyaline, septate, branched, smooth-walled, 2.5 – 3 µm wide hyphae. Asexual morph with straight, branched, cylindrical, 2.5 – 3 µm wide fertile hyphae, giving rise to arthroconidia in a random disposition. Arthroconidia enteroarthric, 0 – 1 - septate, hyaline, smooth- and thin-walled, cylindrical to subcylindrical, barrel- or T-shaped, campaniform, with conspicuous frills, (2.5 –) 3.5 – 6 (– 9) × 2.5 – 3 µm; secession rhexolytic. Racket hyphae present. Sexual morph not observed.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
9AD9FE9B521F58D48C485522DAE3D345.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 24 – 26 mm diam., flat with sparse aerial mycelium, white (1 A 1), margin entire and submerged; reverse uncolored. On PDA, 30 mm diam., crateriform, slightly elevated, radially sulcate, fasciculate in center to velvety towards periphery, white (1 A 1) with light yellow (3 A 4) exudate in center, margin entire; reverse uncolored. Diffusible pigment light yellow (3 A 4). On PYE, 36 mm diam., slightly elevated, radially sulcate, woolly or floccose, yellowish grey (4 B 3) to white (1 A 1), margin fimbriate; reverse radially sulcate, olive brown (4 D 8) to white (1 A 1).	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
C258B588FEF6566486FA9D03CFBA416E.taxon	description	Fig. 8	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
C258B588FEF6566486FA9D03CFBA416E.taxon	etymology	Etymology. Greek para - (παρα -), to resemble, referring to the morphological similarity to the sexual and asexual morphs of M. filamentosa.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
C258B588FEF6566486FA9D03CFBA416E.taxon	description	Description. Saprobic on marine sediments. Mycelium superficial and immersed, composed of hyaline, septate, branched, smooth-walled, 1.5 – 2 µm wide hyphae. Asexual morph with well-differentiated fertile hyphae, arising laterally from vegetative hyphae, branched, straight, sinuous, or contorted, forming intercalary and terminal arthroconidia randomly, 1.5 – 2 µm wide. Arthroconidia enteroarthric, 0 – 1 - septate, hyaline, smooth- and thin-walled, cylindrical to subcylindrical or T-shaped, 3.5 – 4.5 (– 7.5) × 1.5 – 2 µm; secession rhexolytic. Sexual morph heterothallic (observed in FMR 20151 × IHEM 28255 on OA). Gymnothecia superficial, surrounded by asexual fertile hyphae, single or aggregated, orange-brown, globose to subglobose, 200 – 310 µm diam. (excluding appendages); peridium composed of a conspicuous network of hyphae, septate, branched, hyaline to brownish orange, finely asperulate, thick-walled, cylindrical, 2.5 – 4 (– 4.5) µm wide, with short and long lateral appendages; short appendages arising at acute angles, spine-like, with subacute ends, orange-brown, finely asperulate, 14 – 25 (– 30) × 3 – 4 µm; long appendages arising from peridium at acute and subacute angles, unbranched, straight or curved, cylindrical, progressively tapering terminally, orange-brown, finely asperulate and thick-walled with a basal knuckle-joint towards the base, smooth terminally, with a rounded or subacute curved apex, 200 – 560 × 3 – 4 µm long. Asci 8 - spored, evanescent, irregularly disposed, globose to subglobose, 7 × 5 – 6 µm. Ascospores unicellular, pale yellow, yellow in mass, reticulate (reticulation regular with smooth polygonal meshes under SEM), thick-walled, subglobose, 3 × 2.5 µm.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
C258B588FEF6566486FA9D03CFBA416E.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 15 – 20 mm diam., flat, floccose, white (1 A 1), submerged towards periphery, margin fimbriate; reverse white (1 A 1). On PDA, 20 – 22 mm diam., umbonate, cottony, white (1 A 1), margin entire and diffuse; reverse orange-white (5 A 2) to white (1 A 1). On PYE, 25 – 27 mm diam., convoluted, slightly radially sulcate at periphery, cottony, white (1 A 1), margin entire; reverse uncolored. Diffusible pigment not observed in any of the media studied.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
D566DC646B985C6D9A4680EBCD554333.taxon	description	Fig. 9	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
D566DC646B985C6D9A4680EBCD554333.taxon	etymology	Etymology. Latin sedimentum, settling, and Latin - cola, to inhabit, referring to the species’ preference for soil and sediment substrates.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
D566DC646B985C6D9A4680EBCD554333.taxon	description	Description. Saprobic on marine sediments. Mycelium superficial and immersed, composed of septate, branched, hyaline, smooth-walled, 1.5 – 2.5 µm wide hyphae. Asexual morph with intercalary or terminal fertile hyphae growing directly from the vegetative hyphae, often anastomosed, straight, slightly branched, branches short and sinuous, forming arthroconidia randomly, 1.5 – 2.5 µm wide. Arthroconidia enteroarthric, intercalary or terminal, 0 – 1 - septate, hyaline, smooth- and thin-walled, cylindrical to subcylindrical or T-shaped, (2.5 –) 4.5 – 5.5 (– 7.5) × 1.5 – 2.5 µm; secession rhexolytic. Sexual morph homothallic. Gymnothecia observed on OA, superficial, single or aggregated, orange-brown, globose to subglobose, 350 – 450 µm diam. (excluding appendages); peridium composed of a conspicuous network of hyphae, septate, branched, hyaline to brownish orange, asperulate, thick-walled, cylindrical, 2.5 – 3 µm wide, with short and long lateral appendages; short appendages arising at acute angles, spine-like, with subacute to truncated ends, orange-brown, asperulate, 15 – 40 µm long; long appendages arising from peridium at acute and subacute angles, unbranched, straight or curved, cylindrical, progressively tapering terminally, orange-brown, asperulate and thick-walled toward the base, paler and smooth terminally, with a rounded or subacute curved apex and a basal knuckle-joint, 500 – 850 µm long. Asci 8 - spored, evanescent, irregularly disposed, globose, subglobose, or pyriform, 7.5 – 9 × 5.5 – 6 µm. Ascospores unicellular, pale yellow, smooth-walled to slightly echinulate (reticulation regular with smooth polygonal meshes under SEM), thick-walled, globose, 2.5 – 3 µm diam.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
D566DC646B985C6D9A4680EBCD554333.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 40 mm diam., flat, glabrous at center with abundant ascomata, white (1 A 1), producing sparse aerial mycelium towards periphery; reverse uncolored. On PDA, 46 – 50 mm diam., slightly umbonate, cottony, light yellow (4 A 4) to white at periphery, margin entire and diffuse; reverse reddish orange (7 B 8) to light orange (5 A 4), white towards periphery. On PYE, 47 mm diam., slightly umbonate, cottony or glabrous (CBS 319.61), white (1 A 1), with light yellow patches at center, margin entire and diffuse; reverse uncolored. Diffusible pigment not observed in any of the media studied.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
84078856EA015B2D80A295BA4FC19A09.taxon	description	Fig. 10	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
84078856EA015B2D80A295BA4FC19A09.taxon	etymology	Etymology. Latin semi -, half, and Latin nūda, naked, referring to the ascomata with slightly differentiated peridial hyphae.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
84078856EA015B2D80A295BA4FC19A09.taxon	description	Description. Saprobic from marine environment. Mycelium superficial and immersed, composed of hyaline, septate, branched, smooth-walled, 2.5 – 3 µm wide hyphae. Asexual morph with straight, branched, cylindrical, 2.5 – 3 µm wide fertile hyphae, giving rise to arthroconidia in a random disposition. Arthroconidia enteroarthric, 0 – 1 - septate, hyaline, smooth- and thin-walled, cylindrical to subcylindrical or T-shaped, (4 –) 12 – 13 (– 14.5) × 2.5 – 3 µm; secession rhexolytic. Sexual morph homothallic. Gymnothecia observed in all media tested, superficial, aggregated, white to pale yellow, turning pale orange-brown in old cultures (8 weeks), globose to subglobose, 350 – 500 µm diam.; peridium composed of a subtle network of hyphae, septate, branched, hyaline to pale yellow, verruculose to verrucose (tuberculate under SEM), 2.5 – 3 µm wide. Asci 8 - spored, evanescent, irregularly disposed, subglobose or pyriform, 11.5 – 14 × 9 – 10.5 µm. Ascospores unicellular, pale yellow, reticulate (reticulation regular with polygonal meshes and conspicuous ridges under SEM), thick-walled, globose, 4 – 5 µm diam.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
84078856EA015B2D80A295BA4FC19A09.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 30 mm, flat, with sparse white aerial mycelium at center and glabrous towards periphery, margin diffuse, ascomata abundant and densely aggregated in a concentric ring; reverse uncolored. On PDA, 37 mm diam., slightly floccose, with concentrical rings, greenish yellow (1 A 6), margin entire and diffuse, ascomata abundant arranged in concentric rings; reverse brownish yellow (5 C 8) at center, yellowish brown (5 E 8) and yellowish white (1 A 2) at periphery. On PYE, 44 mm diam., slightly raised with concentrical rings, radially sulcate, floccose, white (1 A 1) to greenish yellow (1 A 6), margin entire, submerged in the medium, ascomata abundant arranged in concentric rings; reverse uncolored. Diffusible pigment not observed in any of the media studied.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
B3D0C136EC33524DB5D54E2D8E0E268A.taxon	description	Fig. 11	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
B3D0C136EC33524DB5D54E2D8E0E268A.taxon	etymology	Etymology. Latin sexus, sex, referring to the exclusive presence of the sexual morph in the type strain.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
B3D0C136EC33524DB5D54E2D8E0E268A.taxon	description	Description. Saprobic on marine sediments. Mycelium superficial and immersed, composed of hyaline, septate, branched, smooth-walled, 1.5 – 2 µm wide hyphae. Racket hyphae present. Asexual morph not observed. Sexual morph homothallic. Gymnothecia observed on OA, superficial, single or confluent, pale yellow, globose to subglobose, 250 – 420 µm diam. (excluding appendages); peridium composed of a conspicuous network of hyphae, septate, branched, hyaline to orange-brown, asperulate, thick-walled and cylindrical, 3 – 4.5 µm wide, with short and long lateral appendages; short appendages arising at acute angles, spine-like, with subacute to rounded ends, orange-brown, finely asperulate, 15 – 25 µm long; long appendages arising from peridium at acute and subacute angles, unbranched, straight or curved, cylindrical, progressively tapering terminally, orange-brown, asperulate and thick-walled toward the base, paler and smooth terminally, with a rounded or subacute curved apex and mostly with a basal knuckle-joint, 150 – 340 µm long. Asci 8 - spored, evanescent, irregularly disposed, globose, subglobose, or pyriform, 6 – 8 × 4.5 – 5.5 µm. Ascospores unicellular, pale yellow, smooth-walled to slightly echinulate (reticulation regular with smooth polygonal meshes under SEM), thick-walled, globose, 2 – 2.5 µm diam.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
B3D0C136EC33524DB5D54E2D8E0E268A.taxon	description	Culture characteristics (after 14 days at 25 ° C). Colonies on OA reaching 20 mm diam., flat with sparse aerial mycelium, yellowish white (4 A 2) at initial stages, brownish yellow (5 C 7) when mature, margin entire, ascomata abundant and densely aggregated in center; reverse orange (5 B 8) to pale orange (5 A 3). On PDA, 22 – 26 mm diam., crateriform, radially sulcate at periphery, light orange to orange, velvety, margin entire to slightly lobate; reverse brownish orange (7 C 8) at center, deep yellow (4 A 8) towards periphery. On PYE, 26 – 28 mm diam., elevated, slightly crateriform, radially sulcate, velvety, deep yellow (4 A 8) at center, light orange (5 A 5) towards periphery, margin entire, slightly lobate; reverse brownish orange (7 C 8) to deep yellow (4 A 8). Diffusible pigment not observed in any of the media studied.	en	Guerra-Mateo, Daniel, Gené, Josepa, Becker, Pierre, Cano-Lira, José F. (2025): Onygenales from marine sediments: diversity, novel taxa, global distribution, and adaptability to the marine environment. IMA Fungus 16: e 158470, DOI: 10.3897/imafungus.16.158470
