taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
BBD01E0400F05F0FAAC57955AC81217F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303975	https://doi.org/10.3897/zookeys.1234.141196.figure13	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
BBD01E0400F05F0FAAC57955AC81217F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303976	https://doi.org/10.3897/zookeys.1234.141196.figure14	Figure 14. Epeorus (Caucasiron) guttatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (paratype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	Figure 14. Epeorus (Caucasiron) guttatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (paratype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
BBD01E0400F05F0FAAC57955AC81217F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303962	https://doi.org/10.3897/zookeys.1234.141196.figure1	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303964	https://doi.org/10.3897/zookeys.1234.141196.figure3	Figure 3. Epeorus (Caucasiron) himalayensis sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H, I middle leg in dorsal view J distal part of abdomen in ventral view K – M coloration of abdominal sterna.	Figure 3. Epeorus (Caucasiron) himalayensis sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H, I middle leg in dorsal view J distal part of abdomen in ventral view K – M coloration of abdominal sterna.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303965	https://doi.org/10.3897/zookeys.1234.141196.figure4	Figure 4. Epeorus (Caucasiron) himalayensis sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae and last instar larval exuvium.	Figure 4. Epeorus (Caucasiron) himalayensis sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae and last instar larval exuvium.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303967	https://doi.org/10.3897/zookeys.1234.141196.figure5	Figure 5. Epeorus (Caucasiron) himalayensis sp. nov., male imago A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg in dorsal view H penis in ventral view I titillators J male genitalia in ventral view (arrow points on shallow medio-apical emargination) K penis in dorsal view.	Figure 5. Epeorus (Caucasiron) himalayensis sp. nov., male imago A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg in dorsal view H penis in ventral view I titillators J male genitalia in ventral view (arrow points on shallow medio-apical emargination) K penis in dorsal view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303968	https://doi.org/10.3897/zookeys.1234.141196.figure6	Figure 6. Epeorus (Caucasiron) himalayensis sp. nov., female imago and egg A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G – I subgenital and subanal plates J egg K detail of micropyle. Scale bars: 5 μm (K); 50 μm (J).	Figure 6. Epeorus (Caucasiron) himalayensis sp. nov., female imago and egg A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G – I subgenital and subanal plates J egg K detail of micropyle. Scale bars: 5 μm (K); 50 μm (J).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303973	https://doi.org/10.3897/zookeys.1234.141196.figure11	Figure 11. Epeorus (Caucasiron) lineatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII (drawn from late-instar larva) F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M abdominal terga VIII – X with short spines (arrows) N abdominal terga VIII – X with well-developed spines O sternum IX of female. Drawn from late instar larvae.	Figure 11. Epeorus (Caucasiron) lineatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII (drawn from late-instar larva) F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M abdominal terga VIII – X with short spines (arrows) N abdominal terga VIII – X with well-developed spines O sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303975	https://doi.org/10.3897/zookeys.1234.141196.figure13	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303969	https://doi.org/10.3897/zookeys.1234.141196.figure7	Figure 7. Epeorus (Caucasiron) lanceolatus sp. nov., larva A head of female in dorsal view (holotype) B fore leg in dorsal view C coloration of abdominal terga D distal part of abdomen in ventral view E, F coloration of abdominal sterna II – VI (E holotype).	Figure 7. Epeorus (Caucasiron) lanceolatus sp. nov., larva A head of female in dorsal view (holotype) B fore leg in dorsal view C coloration of abdominal terga D distal part of abdomen in ventral view E, F coloration of abdominal sterna II – VI (E holotype).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303972	https://doi.org/10.3897/zookeys.1234.141196.figure10	Figure 10. Epeorus (Caucasiron) lineatus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G, H distal part of abdomen in ventral view I – M coloration of abdominal terga N – Q coloration of abdominal sterna II – VI.	Figure 10. Epeorus (Caucasiron) lineatus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G, H distal part of abdomen in ventral view I – M coloration of abdominal terga N – Q coloration of abdominal sterna II – VI.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303970	https://doi.org/10.3897/zookeys.1234.141196.figure8	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303962	https://doi.org/10.3897/zookeys.1234.141196.figure1	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303977	https://doi.org/10.3897/zookeys.1234.141196.figure15	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303978	https://doi.org/10.3897/zookeys.1234.141196.figure16	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303980	https://doi.org/10.3897/zookeys.1234.141196.figure18	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303979	https://doi.org/10.3897/zookeys.1234.141196.figure17	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
691B9F9C6D9B5720A5740E797BF2EC19.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303971	https://doi.org/10.3897/zookeys.1234.141196.figure9	Figure 9. Epeorus (Caucasiron) lanceolatus sp. nov., female imago (wing and head not fully molted from subimago) A habitus in dorsal view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg H, I subgenital and subanal plates J egg K detail of micropyle L texture on the surface of egg. Scale bars: 50 μm (J); 5 μm (K, L).	Figure 9. Epeorus (Caucasiron) lanceolatus sp. nov., female imago (wing and head not fully molted from subimago) A habitus in dorsal view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg H, I subgenital and subanal plates J egg K detail of micropyle L texture on the surface of egg. Scale bars: 50 μm (J); 5 μm (K, L).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303969	https://doi.org/10.3897/zookeys.1234.141196.figure7	Figure 7. Epeorus (Caucasiron) lanceolatus sp. nov., larva A head of female in dorsal view (holotype) B fore leg in dorsal view C coloration of abdominal terga D distal part of abdomen in ventral view E, F coloration of abdominal sterna II – VI (E holotype).	Figure 7. Epeorus (Caucasiron) lanceolatus sp. nov., larva A head of female in dorsal view (holotype) B fore leg in dorsal view C coloration of abdominal terga D distal part of abdomen in ventral view E, F coloration of abdominal sterna II – VI (E holotype).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303970	https://doi.org/10.3897/zookeys.1234.141196.figure8	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303971	https://doi.org/10.3897/zookeys.1234.141196.figure9	Figure 9. Epeorus (Caucasiron) lanceolatus sp. nov., female imago (wing and head not fully molted from subimago) A habitus in dorsal view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg H, I subgenital and subanal plates J egg K detail of micropyle L texture on the surface of egg. Scale bars: 50 μm (J); 5 μm (K, L).	Figure 9. Epeorus (Caucasiron) lanceolatus sp. nov., female imago (wing and head not fully molted from subimago) A habitus in dorsal view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G middle leg H, I subgenital and subanal plates J egg K detail of micropyle L texture on the surface of egg. Scale bars: 50 μm (J); 5 μm (K, L).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303977	https://doi.org/10.3897/zookeys.1234.141196.figure15	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303978	https://doi.org/10.3897/zookeys.1234.141196.figure16	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303980	https://doi.org/10.3897/zookeys.1234.141196.figure18	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303979	https://doi.org/10.3897/zookeys.1234.141196.figure17	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
98D3F3C8D9755D23BBCCE07E72D7E3F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303968	https://doi.org/10.3897/zookeys.1234.141196.figure6	Figure 6. Epeorus (Caucasiron) himalayensis sp. nov., female imago and egg A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G – I subgenital and subanal plates J egg K detail of micropyle. Scale bars: 5 μm (K); 50 μm (J).	Figure 6. Epeorus (Caucasiron) himalayensis sp. nov., female imago and egg A habitus in lateral view B head in dorsal view C head in lateral view D abdomen in dorsal view E abdomen in lateral view F abdomen in ventral view G – I subgenital and subanal plates J egg K detail of micropyle. Scale bars: 5 μm (K); 50 μm (J).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303972	https://doi.org/10.3897/zookeys.1234.141196.figure10	Figure 10. Epeorus (Caucasiron) lineatus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G, H distal part of abdomen in ventral view I – M coloration of abdominal terga N – Q coloration of abdominal sterna II – VI.	Figure 10. Epeorus (Caucasiron) lineatus sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F middle leg in dorsal view G, H distal part of abdomen in ventral view I – M coloration of abdominal terga N – Q coloration of abdominal sterna II – VI.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303973	https://doi.org/10.3897/zookeys.1234.141196.figure11	Figure 11. Epeorus (Caucasiron) lineatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII (drawn from late-instar larva) F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M abdominal terga VIII – X with short spines (arrows) N abdominal terga VIII – X with well-developed spines O sternum IX of female. Drawn from late instar larvae.	Figure 11. Epeorus (Caucasiron) lineatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII (drawn from late-instar larva) F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M abdominal terga VIII – X with short spines (arrows) N abdominal terga VIII – X with well-developed spines O sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303964	https://doi.org/10.3897/zookeys.1234.141196.figure3	Figure 3. Epeorus (Caucasiron) himalayensis sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H, I middle leg in dorsal view J distal part of abdomen in ventral view K – M coloration of abdominal sterna.	Figure 3. Epeorus (Caucasiron) himalayensis sp. nov., larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H, I middle leg in dorsal view J distal part of abdomen in ventral view K – M coloration of abdominal sterna.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303965	https://doi.org/10.3897/zookeys.1234.141196.figure4	Figure 4. Epeorus (Caucasiron) himalayensis sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae and last instar larval exuvium.	Figure 4. Epeorus (Caucasiron) himalayensis sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae and last instar larval exuvium.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303970	https://doi.org/10.3897/zookeys.1234.141196.figure8	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	Figure 8. Epeorus (Caucasiron) lanceolatus sp. nov., larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII (holotype) G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill VII (flattened on slide) K gill plate VII in ventral view L, M abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) N sternum IX of female. Drawn from late instar larva and last instar larval exuvium.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303975	https://doi.org/10.3897/zookeys.1234.141196.figure13	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	Figure 13. Epeorus (Caucasiron) guttatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F, G coloration of abdominal terga H coloration of abdominal sterna II – VI I middle leg in dorsal view J distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303962	https://doi.org/10.3897/zookeys.1234.141196.figure1	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303978	https://doi.org/10.3897/zookeys.1234.141196.figure16	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303980	https://doi.org/10.3897/zookeys.1234.141196.figure18	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
FC5293A0C6C857C3A1CDA74F77CA2AF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303979	https://doi.org/10.3897/zookeys.1234.141196.figure17	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
C44E9428B608507EB0A88C493969FF85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303977	https://doi.org/10.3897/zookeys.1234.141196.figure15	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	Figure 15. Epeorus (Caucasiron?) suspicatus, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view (holotype) E head of female in dorsal view F middle leg in dorsal view G, H coloration of abdominal terga (G holotype) I – J coloration of abdominal sterna II – VI (I holotype) K, L distal part of abdomen in ventral view (K holotype).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
C44E9428B608507EB0A88C493969FF85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303978	https://doi.org/10.3897/zookeys.1234.141196.figure16	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	Figure 16. Epeorus (Caucasiron?) suspicatus, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII (flattened on slide) K gill plate VII in ventral view L abdominal segments VIII – X in lateral view (arrow shows indistinct posterolateral projection of tergum X) M sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
C44E9428B608507EB0A88C493969FF85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303962	https://doi.org/10.3897/zookeys.1234.141196.figure1	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
8339538B878F522FB270B9D2A9E0B11F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303979	https://doi.org/10.3897/zookeys.1234.141196.figure17	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	Figure 17. Epeorus (Caucasiron?) psi, larva A habitus in dorsal view B habitus in ventral view C habitus in lateral view D head of male in dorsal view E head of female in dorsal view F legs in dorsal view (arrows point to elongated and pointed dorso-apical projections) G coloration of abdominal terga H coloration of abdominal sterna II – VI I distal part of abdomen in ventral view.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
8339538B878F522FB270B9D2A9E0B11F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303980	https://doi.org/10.3897/zookeys.1234.141196.figure18	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	Figure 18. Epeorus (Caucasiron?) psi, larva A labrum, left half in dorsal view, right half in ventral view (black dots correspond to setae along antero-lateral margin) B incisors of left mandible C incisors of right mandible (both flattened on slide; dashed polygons correspond to area covered by setae) D setae on dorsal surface of femora E setae on surface of tergum VII F surface and posterior margin of abdominal tergum VII G gill I H gill III (arrow shows projection on costal margin) I gill plate VI in dorsal view J gill plate VII in ventral view K gill plate VII (flattened on slide) L abdominal segments VIII – X in lateral view (arrow shows posterolateral projection of tergum X) M detail of dorso-apical projection of femora (middle leg) N sternum IX of female. Drawn from late instar larvae.	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
8339538B878F522FB270B9D2A9E0B11F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1303962	https://doi.org/10.3897/zookeys.1234.141196.figure1	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	Figure 1. Distribution of studied Epeorus spp. across Asian mountains. Triangles refer to localities in the literature; our sampling sites are marked with red dots. A species name marked with an asterisk indicates the unclear systematic position of the species within the genus Epeorus. Dashed line represents the frame of the detailed map in Fig. 2. Boundaries of mountain ranges were adopted from Snethlage et al. (2022).	2025-04-09	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška		Zenodo	biologists	Hrivniak, Ľuboš;Sroka, Pavel;Godunko, Roman J.;Martynov, Alexander V.;Palatov, Dmitry M.;Bojková, Jindřiška			
