taxonID	type	description	language	source
4A5987BAFFA49319E9CA8385FBB8F926.taxon	description	(Figs 1 A – F, 10 A, 14 A – H, 15 A – D, 25 A, 25 G – J)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA49319E9CA8385FBB8F926.taxon	materials_examined	Type material examined (1 ♂, 2 ♀♀). Lectotype (hereby designated): 1 ♂ (NHMUK) “ ♂ // Type // Kurseong, Sikkim R. P. Verschraeghen. 1911 - 218. // Kurseong Inde Verschraeghen 1904 1907 // Anomala flavonotata, Arrow Type // NHMUK 010844184 ”. Paralectotypes: 1 ♀ (NHMUK) “ ♀ // Kurseong, Sikkim R. P. Verschraeghen. 1911 - 218. // Kurseong Inde Verschraeghen 1904 ”; 1 ♀ (MNHN) “ Kurseong Inde Verschraeghen 1904 1908 // Anomala flavonotata Arr. Déterminé par Arrow 1912 // Type // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”. Additional material examined (25 ♂♂, 23 ♀♀). 1 ♀ (GSPC), “ N. E. INDIA Darjeeling ”; 1 ♂, 1 ♀ (ZMPC) “ N. E. INDIA Darjeeling ”; 1 ♂ (GSPC) “ Umgebung Darjeeling 20.8. // anom. flavonotata, arr. Compared with type. G. J. A. ”; 1 ♂ (NMBF) “ Darjeeling W. B. 2180 m, VI. 1961 leg. G. Scherer // Anomala flavonotata Arr det. G. Frey, 1962 ”; 2 ♀♀ (MHNG) “ Kurseong, Darjeeling // Coll. R. Petrovitz ”; 9 ♂♂, 8 ♀♀ (MNHN) “ Kurseong R. P. Decoly 1898 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂, 1 ♀ (ZMPC) “ Kurseong R. P. Decoly 1898 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♀ (MNHN) “ Darjeeling Inde britan. Coll. J. Clermont // MUSÉUM PARIS 1944 Coll. E. BENDERITTER ”; 1 ♂ (CZPC) “ INDIA – SIKKIM west PEELING env., 2100 m 18. - 20.7.1997 Jan Schneider lgt. ”; 1 ♂, 1 ♀ (CZPC) “ north India - west SIKKIM Khecheopari lake (1800 m. m) 4. - 10.6.1999 lgt. E. Kučera ”; 1 ♂ (ZMPC) “ north INDIA - west SIKKIM Khecheopari lake (1800 m. m) 4. - 10.6.1999 lgt. E. Kučera ”; 1 ♂ (MFPC) “ INDIA - SIKKIM Khecheopalri lake 15.6 – 22.6.2008 1800 m leg. E. Kučera ”; 1 ♂ (NMPC) “ north INDIA - west Bengal Shingalila national park Tonglu (3070 m. m) 14. - 19.6.99 lgt. E. Kučera ”; 1 ♀ (CZPC) “ north India - west SIKKIM Shingalila national park Tonglu (3070 m. m) 14. - 19.6.1999 lgt. E. Kučera ”; 1 ♀ (CZPC) “ India. SIKKIM east GANTOK env., 2000 - 2500 m FAMBONG LHO forest 8. - 15.7.1997 Jan Schneider lgt. ”; 1 ♀ (NHMUK) “ INDIA: Kurseong Div. Lat Panchar. 4,000 ft. vi. 1934. H. G. Champion. // G. C. Champion Coll. B. M. 1927 409. ”; 1 ♀ (MNHN) “ Environs de Kurseong R P. Bretaudeau // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♀ (MNHN) “ Sikkim Rhenok Eté 1894 Chasseurs Bretaudeau ”; 1 ♀ (MNHN) “ Sikkim Lanpamum // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂ (NHMUK) “ 4 // VII- 21 - 1923 El. 5000 ft. // Tendong Sikkim // NHMUK 015015957 ”; 1 ♂ (MNHN) “ Maria Basti British Bootang. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂ (MNHN) “ British Bootang Sakiȯu L. Durel // Anomala flavonotata, arr. G. J. Arrow det. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 2 ♂♂, 2 ♀♀ (MFPC) “ BHUTAN: Monger, Thebong 27 ° 17 ′ N 90 ° 20 ′ E 19. VII. 2009 H. 2273 m Viktor Siniaev leg. ”; 1 ♂ (MFPC) “ China-India border: Tawang 2500 m 05 – 10. JUL. 2013 Takeshi Maeda leg. ”; 1 ♂ (ZMPC) “ CHINA: Tibet, Shigatse, Lower Yatung, N 27 ° 15 ′ 22.38 ′′, E 89 ° 01 ′ 10.17 ′′, 1832 m, 2023. VII. 10, at light, Quan-Yu Ji leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA49319E9CA8385FBB8F926.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally brown to black, elytra sometimes lighter; shiny, with weak greenish luster, head tinged with bronze luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1, sometimes partly interrupted; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.2; subtrapezoidal, lateral margins straight and weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with very dense, large and irregular punctures. Frontoclypeal suture straight, distinct. Anterior half of frons with dense and large punctures, gradually sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin slightly excurved; with moderately dense and minute punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures, most sparse on subsutural interstice. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of sparse, short to moderately long setae from base to apical fifth. Abdomen. Propygidium with moderately dense and transverse punctures. Pygidium with dense and transverse striation, gradually breaking up into sparse and very transverse punctures towards the smooth greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 with very dense and irregular punctures; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth acute or rounded apically, extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See figs 14 A – H and 15 A – D. Female. Interocular distance / maximum head width approximately 0.7; length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, subapical tooth somewhat rounded apically; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider and shorter than upper branch. Measurement. Body length: 14.7 – 15.3 mm in males and 14.9 – 15.4 mm in females, greatest width: 8.3 – 9.0 mm in males and 8.2 – 9.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA49319E9CA8385FBB8F926.taxon	diagnosis	Differential diagnosis. Anomala flavonotata is characterized by a bifurcate upper branch of the left paramere, with a short side branch present in the apical third of the left paramere, combined with a simply pointed apex of the left paramere in lateral view. The internoapical angle is either absent or shortly produced in contrast to A. menyuensis sp. nov. Moreover, the epipleura are only sparsely covered with setae in A. flavonotata (densely covered in A. menyuensis sp. nov.).	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA49319E9CA8385FBB8F926.taxon	distribution	Distribution. China (Xizang) (new record); India (West Bengal, Sikkim); Bhutan.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	description	(Figs 2 A – C, 10 B, 15 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	materials_examined	Type material (12 ♂♂, 3 ♀♀). Holotype: ♂ (NHMB) “ China-India border: DIRANG vicinity, 1550 ± 150 m 27 ° 21 ′ - 23 ′ N 92 ° 13 ′ - 16 ′ E L. Dembický leg.; 1 - 9. vi. 2004 ”. Paratypes: 8 ♂♂, 2 ♀♀ (NHMB), “ China-India border: DIRANG vicinity, 1550 ± 150 m 27 ° 21 ′ - 23 ′ N 92 ° 13 ′ - 16 ′ E L. Dembický leg.; 1 - 9. vi. 2004 ”; 2 ♂♂, 1 ♀ (ZMPC), “ China-India border: DIRANG vicinity, 1550 ± 150 m 27 ° 21 ′ - 23 ′ N 92 ° 13 ′ - 16 ′ E L. Dembický leg.; 1 - 9. vi. 2004 ”; 1 ♂ (ZMPC) “ China-India border: East Kameng, Chayangtajo circle, Laching bagang, 2200 m 2024. VIII. 22 ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally brown to black, the elytra sometimes lighter; shiny, with weak purplish or greenish luster, head tinged with bronze luster; the elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1, sometimes partly interrupted; antennae yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.4; subtrapezoidal, lateral margins straight and weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with very dense, large and irregular punctures. Frontoclypeal suture straight, distinct. Anterior half of frons with dense and large punctures, gradually sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin nearly straight; with moderately dense and minute punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures, most sparse on subsutural interstice. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of dense, short to moderately long setae from base to apical protuberance. Abdomen. Propygidium with moderately dense and very transverse punctures. Pygidium with dense and transverse striation, gradually breaking up into sparse and transverse punctures towards the smooth greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 with very dense and irregular punctures; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth acute or rounded apically, extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 15 E – H. Female. The zigzag band on elytra broad, sometimes interrupted where reaching subsutural interstice. Interocular distance / maximum head width approximately 0.7; length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, subapical tooth somewhat rounded apically; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider and shorter than upper branch. Measurement. Body length: 15.8 – 16.1 mm in males and 15.8 – 16.1 mm in females, greatest width: 9.1 mm in males and 9.0 – 9.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	diagnosis	Differential diagnosis. Anomala menyuensis sp. nov. is closely related to A. flavonotata but can be reliably distinguished from this species by the left paramere, whose dorsal branch has an enlarged, rounded apex in lateral view with a distinct subapical protuberance, whereas in A. flavonotata, the apex is simply pointed. Additionally, the epipleura of A. menyuensis sp. nov. are densely covered with setae, in contrast to the sparse setation in A. flavonotata.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	discussion	Remarks. We examined a specimen of A. flavonotata from Tawang (only ca. 35 km northwest of the type locality of A. menyuensis sp. nov., Dirang) exhibiting morphological consistency with specimens from Sikkim and Bhutan. Despite their similar male genitalia, the stability of A. flavonotata across its distribution range suggests that it is distinct from A. menyuensis sp. nov.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA2931BE9CA8639FB25FD56.taxon	etymology	Etymology. The specific epithet menyuensis is derived from Menyu (also spelled Monyul; mā in Chinese), a historical regional name in southeastern Tibet. The species is currently known only from this region.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA09315E9CA87B4FCA9FEFF.taxon	description	(Figs 3 A – B, 10 C, 16 A – D)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA09315E9CA87B4FCA9FEFF.taxon	materials_examined	Type material (3 ♂♂, 4 ♀♀). Holotype: ♂ (NSMT) “ China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg. ” Paratypes: 1 ♀ (MFPC) “ China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg. ”; 1 ♂, 1 ♀ (ZMPC) “ China-India border: W. Siang 30. JUNE. 2011 Takeshi MAEDA-leg. ”; 1 ♂, 1 ♀ (MFPC) “ China-India border: alt. 1700 m Lower Subansiri 24 – 27. JUN. 2015 Takeshi Maeda-leg. ”; 1 ♀ (MFPC) “ China-India border: alt. 1700 m Lower Subansiri 12 – 20. AUG. 2015 Takeshi Maeda-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA09315E9CA87B4FCA9FEFF.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally brown to black, elytra sometimes lighter; shiny, with strong greenish luster; elytra with a yellow, broad zigzag band extending from primary costa 5 to 1; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.4; subtrapezoidal, lateral margins straight and moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons rugopunctate, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved; with dense and small punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, moderately dense and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of sparse, short to moderately long setae from base to apical fifth. Abdomen. Propygidium with very dense, transverse and very fine striation. Pygidium with dense and transverse striation, sparse only on the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 16 A – D. Female. Sometimes with weak purple luster. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 rather weakly developed; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than the upper one. Measurement. Body length: 14.3 – 15.2 mm in males and 13.8 – 14.7 mm in females, greatest width: 8.1 – 8.4 mm in males and 8.0 – 9.1 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA09315E9CA87B4FCA9FEFF.taxon	diagnosis	Differential diagnosis. The bifurcate upper branch of the left paramere of Anomala spiniforceps sp. nov. resembles that of A. flavonotata. However, the additional branch is situated near the middle in the new species (situated in the apical third in A. flavonotata). The lower branches of both parameres have long and acute internoapical angles in A. spiniforceps sp. nov., whereas they are distinctly shorter or absent in A. flavonotata.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA09315E9CA87B4FCA9FEFF.taxon	etymology	Etymology. The name spiniforceps is derived from the Latin words spina (spine) and forceps. It refers to the peculiar parameres, which bear prominent spine-like projections.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAE9317E9CA809CFCC4FD6B.taxon	description	(Figs 3 C – F, 10 E, 17 A – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAE9317E9CA809CFCC4FD6B.taxon	materials_examined	Type material (20 ♂, 16 ♀♀). Holotype: ♂ (SCAU) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ” Paratypes: 6 ♂♂, 8 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 5 ♂♂, 5 ♀♀ (ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 5 ♂♂, 2 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill., 1768 m, 29.21315309 ° N, 95.17111894 ° E, 2023. VII. 13 - 14, Hao-Yi Liu leg. ”; 1 ♂ (MFPC) “ China-India border: Upper Siang 1850 m 04 – 10. AUG. 2013 Takeshi Maeda-leg. ”; 2 ♂♂, 1 ♀ (MFPC) “ China-India border: alt. 1500 m Dibang Valley 21 – 30. JUN. 2017 Takeshi Maeda-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAE9317E9CA809CFCC4FD6B.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally dark brown to black, the elytra usually lighter; shiny, with weak to strong greenish luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins straight and moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and coarsely rugopunctate. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons rugopunctate, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.65. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved; with dense and small punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a secondary stria, all intervals bearing moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of dense, moderately long setae from base to apical protuberance, often partly abraded. Abdomen. Propygidium with very dense, transverse and very fine striation. Pygidium with dense and transverse striation, sparse only on the greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 each with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 17 A – H. Female. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral margin of pronotum more excurved. Lateral carina of elytra slightly wider than in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than the upper one. Measurement. Body length: 13.7 – 15.5 mm in males and 15.0 – 16.2 mm in females, greatest width: 7.5 – 8.9 mm in males and 8.2 – 9.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAE9317E9CA809CFCC4FD6B.taxon	diagnosis	Differential diagnosis. The parameres of Anomala luoyuensis sp. nov. are most similar to those of A. spiniforceps sp. nov. in general structure. In A. luoyuensis, the upper branch of the left paramere is distinctly thickened and unbranched, and the lower branches of both parameres are completely narrowed in ventral view, rather long, unequal in length, and parallel. In contrast, the lower branches are broad in A. spiniforceps, with only the internoapical angle narrowed and protruding, equal in length and overlapping.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAE9317E9CA809CFCC4FD6B.taxon	etymology	Etymology. Luoyu (also spelled Lhoyul; Ḇā in Chinese) is a regional name that has historically been used to refer to an area in southeastern Tibet, roughly corresponding to present-day Motuo County. The specific epithet refers to the distribution of the new species in this region.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAC9311E9CA8275FD1CFE12.taxon	description	(Figs 4 A – D, 10 D, 18 A – D)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAC9311E9CA8275FD1CFE12.taxon	materials_examined	Type material (33 ♂♂, 54 ♀♀). Holotype: ♂ (CAU) “ CHINA: Tibet, Nyingchi City, Medog County, 1106 m 29 ° 19 ′ 41.01 ″ N, 95 ° 19 ′ 55.25 ″ E 2022. VIII. 16 light trap Yu-Chen Zheng leg. ” Paratypes: 1 ♂ (CCPC) “ CHINA, Xizang (Tibet), Motuo Beibeng, Jiangxincun 705 m, 2019, VII- 24, Leg. X. - D. YANG CCCC ”; 1 ♂ (ZMPC) “ CHINA: Tibet, Nyingchi City, Medog County, 1143 m 29 ° 19 ′ 22.62 ″ N, 95 ° 20 ′ 1.65 ″ E 26. VII. 2022 (at light) Chuan-Tao Qu leg. ”; 1 ♀ (ZMPC) “ CHINA: Tibet, Nyingchi City, Medog County, 1143 m 29 ° 19 ′ 22.62 ″ N, 95 ° 20 ′ 1.65 ″ E 20. VII. 2022 (at light) Chuan-Tao Qu leg.; 2 ♀♀ (ZMPC) “ CHINA Xizang Nyingchi City Medog County Town 2015 - VIII- 16 Zhuo Chen leg. ”; 1 ♂ (ZMPC) “ Xizang, Medog Viewing Platform 2020 - VII- 29 light Yi-Ming Zhang leg. ”; 1 ♂, 1 ♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Beibeng, 864 m, 29.24807910 ° N, 95.18061479 ° E, 2023. VII. 12, Hao-Yi Liu leg. ”; 5 ♂♂, 13 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill., 1768 m, 29.21315309 ° N, 95.17111894 ° E, 2023. VII. 13 - 14, Hao-Yi Liu leg. ”; 1 ♂, 2 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., 96 k Marangkang vill., 1305 m, 29.56287354 ° N, 95.46563172 ° E, 2023. VII. 17, Hao-Yi Liu leg. ”; 10 ♂♂, 9 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 8 ♂♂, 8 ♀♀ (ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 1 ♂, 2 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi, Motuo, Beibeng Township, Gelin Vill. 1560 m, 21. VIII. 2023, Zhi-Hao Qi, Chuan Liu, Peng-Xu Mu leg. ”; 1 ♂, 2 ♀♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 20, 29.3431 ° N, 95.17 ° E, light trap, Hong-Bin Liang et al. leg. // IOZ (E) 2083634 – 636 ”; 1 ♂, 7 ♀♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 19, 29. 3431 ° N, 95.17 ° E, light trap, Hong-Bin Liang, Zheng-Zhong Huang leg. // IOZ (E) 2083637 – 644 ”; 1 ♀ (IZAS) “ Xizang, North of Medog County, 1005 m, 2015. VIII. 14 N, 29.3319 ° N, 95.3397 ° E, light trap, Hong-Bin Liang, Zheng-Zhong Huang leg. // IOZ (E) 2083645 ”; 1 ♀ (IZAS) “ Xizang, North of Medog County, 1005 m, 2015. VIII. 16, 29.3319 ° N, 95.3397 ° E, light trap, Hong-Bin Liang et al. leg. // IOZ (E) 2083646 ”; 1 ♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 20, 29.3431 ° N, 95.17 ° E, light trap, Hong-Bin Liang leg. // IOZ (E) 2083647 ”; 1 ♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 23, 29.3431 ° N, 95.17 ° E, light trap, Hong-Bin Liang, Zheng-Zhong Huang leg. // IOZ (E) 2083648 ”; 1 ♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 23, 29.3431 ° N, 95.17 ° E, light trap, Jian Yao leg. // IOZ (E) 2083649 ”; 1 ♀ (IZAS) “ Xizang, Medog County, Beibeng Township, 799 m, 2015. VIII. 24, 29.3431 ° N, 95.17 ° E, light trap, Jian Yao leg. // IOZ (E) 2083650 ”; 1 ♂, 1 ♀ (CZPC) “ CHINA, Tibet Linzhi pref., Motuo co. 80 k, June 2019 local collector leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAC9311E9CA8275FD1CFE12.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally black; shiny, with weak greenish luster; elytra with a yellow, narrow to very broad zigzag band extending from primary costa 5 to 1; protibial teeth and antennae reddish brown, club darker; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margin almost straight and weakly convergent anteriad, anterior margin slightly convex, anterior corner obsolete and rounded; anterior margin weakly reflexed; surface very densely and finely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior half of frons rugopunctate, other portions with dense and large punctures, gradually sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.7. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad and weakly curved. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved; with dense and minute punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, moderately dense and large punctures, interstice II with a secondary stria, all intervals bearing moderately dense and minute punctures. Humeral protuberance moderately prominent, apical protuberance strongly prominent. Lateral carina weakly expanded from base to apical two fifths. Epipleuron with a row of rather sparse and short setae in basal third. Abdomen. Propygidium with very dense and transverse striation. Pygidium with dense and transverse striation, sparse on the greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally, concave between ventrites 5 and 6 in profile; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin .. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 18 A – D. Female. The zigzag band on elytral narrow to broad, sometimes interrupted where reaching the subsutural interstice. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral margin of pronotum more excurved. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially; laterally flat between ventrites 5 and 6. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider and shorter than upper branch. Measurement. Body length: 14.8 – 17.0 mm in males and 14.9 – 17.3 mm in females, greatest width: 8.3 – 10.0 mm in males and 9.2 – 10.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAC9311E9CA8275FD1CFE12.taxon	diagnosis	Differential diagnosis. Anomala latifasciata sp. nov. can be distinguished from the similar A. flavonotata by the unbranched and apically rounded upper branch of the left paramere. The lower branches of both parameres have a straight lateral margin, which is distinctly bend preapically with the internoapical angle not produced.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAC9311E9CA8275FD1CFE12.taxon	etymology	Etymology. The name latifasciata comes from the Latin words latus (broad) and fasciatus (banded), referring to the broadest elytral band present in the species.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAB9313E9CA81A0FDEAFEDA.taxon	description	(Figs 4 E, 10 F, 18 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAB9313E9CA81A0FDEAFEDA.taxon	materials_examined	Type material. Holotype: ♂ (NSMT) “ China-India border: alt. 2500 m West Kameng 29. JUN. – 03. JUL. 2015 Takeshi Maeda-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAB9313E9CA81A0FDEAFEDA.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally black; shiny, with weak bronze luster; elytra dark brown, with a yellow, broad to very broad zigzag band extending from primary costa 5 to 1; protibial teeth and antennae reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins almost straight and subparallel, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface very densely and finely rugopunctate. Frontoclypeal suture straight, distinct. Anterior third of frons finely rugopunctate, other portions including vertex with moderately dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, weakly curved at middle. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved; with dense and small punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, moderately dense and large punctures, interstice II with a somewhat irregular secondary stria, all intervals bearing moderately dense and minute punctures. Humeral and apical protuberances both strongly prominent. Lateral carina weakly expanded from base to middle. Epipleuron with a row of rather sparse and short setae in basal third. Abdomen. Propygidium with very dense and transverse striation. Pygidium with dense and transverse striation, gradually sparser apicad, breaking up into sparse and transverse punctures only near the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, both equal in length; lower margin of lower branch of inner protarsal claw not convex, forming a notch internobasally; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 18 E – H. Female. Unknown. Measurement. Body length: 16.2 mm, greatest width: 8.2 mm.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAB9313E9CA81A0FDEAFEDA.taxon	diagnosis	Differential diagnosis. Anomala incurva sp. nov. is very similar to A. latifasciata sp. nov. But the upper branch of the left paramere bears a widely expanded, flat and incurved apex. The lower branches of both parameres have straight lateral margins.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFAB9313E9CA81A0FDEAFEDA.taxon	etymology	Etymology. The species name derives from the Latin adjective incurvatus, meaning curved or bent, referring to the special shape of left paramere.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA89312E9CA80B0FB4CFA06.taxon	description	(Figs 5 A – C, 11 E, 19 A – D)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA89312E9CA80B0FB4CFA06.taxon	materials_examined	Type material (12 ♂♂, 29 ♀♀). Holotype: ♂ (SYSU, En- 419555) “ Xizang, Linzhi region, Bomi County, Tongmai Town 30 ° 6 ′ 8 ″ N 95 ° 6 ′ 2 ″ E 2088 m 2018. VII. 8 Zu-Long Liang & Shi-Shuai Wang leg. ” Paratypes: 1 ♂, 1 ♀ (SYSU, En- 419553 – 419554) “ Xizang, Linzhi region, Bomi County, Tongmai Town 30 ° 6 ′ 8 ″ N 95 ° 6 ′ 2 ″ E 2088 m 2018. VII. 8 Zu-Long Liang & Shi-Shuai Wang leg. ”; 1 ♂, 3 ♀♀ (ZMPC) “ CHINA: Nyingchi, Bomê, Tangmai, 2150 m, 2020. VIII. 5, Yu-Chen Zheng leg. ”; 1 ♂, 7 ♀♀ (ZMPC) “ CHINA: Nyingchi, Bomê, Tangmai, 2150 m, 2020. VIII. 6, Yu-Chen Zheng leg. ”; 1 ♂, 3 ♀♀ (GSPC) “ CHINA - Tibet Linzhi, Bomi county 2000 m, VII. 2020 ”; 1 ♀ (NMPC) “ CHINA, E-TIBET 2000 - 2400 m N of BRAHAMAPUTRA great bend 30 ° 00 ′ - 07 ′ / 94 ° 52 ′ - 95 ° 09 ′ 16. - 20.7.92 L. + R. BUSINSKÝ lgt. ”; 1 ♂, 2 ♀♀ (CZPC) “ CHINA, Tibet, Linzhi Bomi county, 2200 m VII. 2023, loc. coll. leg. ”; 1 ♂, 1 ♀ (CZPC) “ CHINA, Tibet Linzhi area, Bomi co. 2000 m, V-VI. 2021 local collector leg. ”; 3 ♀♀ (CZPC) “ CHINA, Tibet Linzhi area, Bomi co. 2000 m, VIII-IX. 2019 local collector leg. ”; 2 ♂♂, 5 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi, Lulang, Pailong, Shentang Hot Spring, 30 ° 0 ′ 49 ″ N, 94 ° 59 ′ 24 ″ E, 2060 m, 2025. VII. 26, Chuan-Tao Qu leg. ”; 1 ♂, 2 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Xu-Hong-Yi Zheng ”; 1 ♂ (SCAU) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Zhi-Teng Chen leg. // ABM 1 ”; 1 ♂, 1 ♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Zhi-Teng Chen leg. ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA89312E9CA80B0FB4CFA06.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally reddish brown, elytra lighter, rarely generally dark brown; dull, with strong greenish sheen; elytra with a yellow, narrow to broad zigzag band extending from primary costa 3 to 1, and a yellow spot on primary costa 4, rarely connected; antennae reddish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with dense, large and coarse punctures. Frontoclypeal suture slightly bisinuate, distinct. Anterior third of frons and area along inner margin of eye punctate as clypeus, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.65. Length of antennal club slightly longer than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin feebly incurved in anterior third or posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle right-angled and rounded apically. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly excurved; with dense and small punctures, covered by dense and short mesonotal setae along base. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a secondary stria, all intervals with dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of dense and short setae from base to the level of apical protuberance, usually partly abraded. Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse rugostriation, also dense on the greatest tumidity; with dense and long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 each with transverse and incompletely annulated punctures, short and moderately dense medially, gradually transverse and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to anterior margin of protarsomere 2. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an indistinct basal protuberance. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial, also with some irregular and long setae sublaterally; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 19 A – D. Female. Generally dull black with strong greenish sheen, rarely elytra reddish brown around scutellum; the elytral band sometimes formed by three separate small spots. Length of antennal club approximately 0.9 times as long as combined length of antennomeres 1 – 6. Lateral margin of pronotum more excurved at middle, straight at anterior and posterior third; posterior angle obtuse. Lateral carina of elytra slightly wider than in male; setae along lateral margin longer. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to middle of protarsomere 3; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of pro- and metatarsomere 5 acute, that of metatarsomere rather small, mesotarsomere without denticle, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw as long as its upper one and the lower margin not convex. Measurement. Body length: 14.0 – 14.6 mm in males and 14.0 – 15.3 mm in females, greatest width: 7.7 – 8.1 mm in males and 7.5 – 8.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA89312E9CA80B0FB4CFA06.taxon	diagnosis	Differential diagnosis. Anomala bomiensis sp. nov. can be distinguished from all other species of the A. flavonotata - subgroup (see discussion below) by the dull, but slightly iridescent surface of the body. The upper branch of the left paramere is horizontally protruding at middle.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA89312E9CA80B0FB4CFA06.taxon	etymology	Etymology. This new species is named after the type locality, Bomi County in Xizang.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA9930CE9CA84D4FEE1F846.taxon	description	(Figs 5 D – F, 11 A, 19 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA9930CE9CA84D4FEE1F846.taxon	materials_examined	Type material (19 ♂♂, 9 ♀♀). Holotype: ♂ (SCAU) “ TIBET, Nyingchi, Medog, Bari, Renqinbeng, 1942 m, 2014 - VII- 29 small lamp trap, Xiao-Dong Yang Leg. 14 Y 0324 CCCC ”. Paratypes: 1 ♀ (CCPC) “ TIBET, Nyingchi, Medog, Bari, Renqinbeng, 1942 m, 2014 - VIII- 7 small lamp trap, Xiao-Dong Yang Leg. 14 Y 0415 CCCC ”; 1 ♂ (CCPC) “ TIBET, Nyingchi, Medog, Bari, Renqinbeng, 1942 m, 2014 - VIII- 7 small lamp trap, Xiao-Dong Yang Leg. 14 Y 0416 CCCC ”; 1 ♂ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Renqinbeng, 2023. VII. 22, Zhi-Teng Chen leg. ”; 1 ♂ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog, 1117 m, 29.33116683 ° N, 95.33869959 ° E 2025. VI. 28, Ke-Dian Chen leg. ”; 1 ♂ (ZMPC) “ CHINA: TIBET, Nyingchi City, Bolonggong (80 k), 2243 m, 29.65540554 ° N, 95.49091649 ° E 2023. VII. 10, Hao-Yi Liu leg. // Voucher ACC 1 ”; 3 ♂♂, 1 ♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 3 ♂♂, 1 ♀ (ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 3 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill., 1768 m, 29.21315309 ° N, 95.17111894 ° E, 2023. VII. 13 - 14, Hao-Yi Liu leg. ”; 5 ♂♂ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, S of Bolonggong, 2023. VII. 9, Xu-Hong-Yi Zheng ”; 2 ♂♂, 2 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Zhi-Teng Chen leg. ”; 1 ♀ (SCAU) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Zhi-Teng Chen leg. // ACC 2 ”; 1 ♂ (ZMPC) “ CHINA: TIBET, Nyingchi City, S of Medog Co., 1894 m 29.32160167 N. 95.36277584 E 2023. VII. 10 Zhi-Teng Chen lgt. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA9930CE9CA84D4FEE1F846.taxon	description	Description. Male. Body shape elongated ovoid, weakly convex. Color. Generally dark reddish brown to black; shiny, with strong bronze or greenish luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1; antennae yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons punctate as clypeus, other portions with dense and large punctures, slightly sparser and smaller posteriad, sparse on vertex. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, curved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly to moderately excurved; with dense and large punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of dense and moderately long setae from base to near apex, often partly abraded. Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse rugostriation, sparse around the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to anterior margin of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equal in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres distinctly thickened, each tarsomere 5 with an internomedial denticle, that of protarsomere 5 largest and triangular, others small and acute, metatarsomere 5 with an additional rounded basal protuberance. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 19 E – H. Female. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small and acute, basal protuberance of metatarsomere 5 weak; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw as long as its upper one and the lower margin not convex. Measurement. Body length: 11.9 – 14.0 mm in males and 12.2 – 14.0 mm in females, greatest width: 6.4 – 8.0 mm in males and 7.1 – 7.9 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA9930CE9CA84D4FEE1F846.taxon	diagnosis	Differential diagnosis. Anomala lhoba sp. nov. can be distinguished from all other species of the A. flavonotata - subgroup by the shortest upper branch of left paramere, as well as the simple lower branches of both parameres, which are not bifurcate in lateral view.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFA9930CE9CA84D4FEE1F846.taxon	etymology	Etymology. The new species is named after the Lhoba people, an ethnic group traditionally residing in the Motuo region.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB4930EE9CA81A0FBAEFD57.taxon	description	(Figs 6 A – C, 11 D, 20 A – D)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB4930EE9CA81A0FBAEFD57.taxon	materials_examined	Type material (11 ♂♂, 6 ♀♀). Holotype: ♂ (SCAU) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan, 2019. IV – VI, at light Wei-Zong Yang leg. ” Paratypes: 9 ♂♂, 6 ♀♀ (ZMPC) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan, 2019. IV – VI, at light Wei-Zong Yang leg. ”; 1 ♂ (CZPC) “ CHINA: Yunnan, Dehong, Yingjiang, Tongbiguan, 2019. IV – VI, at light Wei-Zong Yang leg. ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB4930EE9CA81A0FBAEFD57.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally dark brown to black; shiny, with weak greenish luster; elytra with a yellow, narrow zigzag band extending from primary costa 5 to 1, usually broken up into three spots; antennae dark brown; protibial teeth reddish brown only at extremity; abdominal ventrites 2 – 4 sometimes with some vague reddish spots; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. An inverted triangular area on anterior third of frons with dense and coarse punctures, other portions with dense and large punctures, slightly sparser and smaller posteriad, sparse on vertex. Interocular distance / maximum head width approximately 0.6. Length of antennal club equally as long as combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior and posterior third, curved at middle. Anterior angle right-angled and protruding, posterior angle obtuse. Posterior marginal line indistinct and broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly to moderately excurved; with dense and large punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of moderately dense and short setae from base to near apex, often partly abraded. Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse striation, sparse around the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 moderately densely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth rounded. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional rounded basal protuberance. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 20 A – D. Female. The elytral band always formed by two or three separate small spots, sometimes very indistinct. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to middle of protarsomere 3, preapical tooth acute; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small and acute, basal protuberance of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than its upper one and the lower margin not convex. Measurement. Body length: 14.7 – 16.0 mm in males and 15.3 – 15.5 mm in females, greatest width: 8.3 – 9.2 mm in males and 9.0 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB4930EE9CA81A0FBAEFD57.taxon	diagnosis	Differential diagnosis. Anomala dilatostilus sp. nov. can be distinguished from all other species of the A. flavonotata - subgroup by the longest upper branch of the left paramere, which is abruptly widened in the distal section. The lower branches of both parameres are long, with an almost straight inner margin.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB4930EE9CA81A0FBAEFD57.taxon	etymology	Etymology. The name dilatostilus is derived from the Latin adjective dilatus (expanded or widened) and noun stilus (stem). It describes the abruptly widening of the left paramere in the distal section.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB59308E9CA8224FAE4FB3A.taxon	description	(Figs 6 D – F, 11 B, 20 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB59308E9CA8224FAE4FB3A.taxon	materials_examined	Type material (3 ♂♂, 3 ♀♀). Holotype: ♂ (NSMT) “ China-India border: alt. 1500 m Dibang Valley 21 – 30. JUN. 2017 Takeshi Maeda-leg. ” Paratypes: 1 ♀ (MFPC) “ China-India border: alt. 1500 m Dibang Valley 21 – 30. JUN. 2017 Takeshi Maeda-leg. ”; 1 ♂ (ZMPC) “ China-India border: alt. 1500 m Dibang Valley 21 – 30. JUN. 2017 Takeshi Maeda-leg. ”; 1 ♂, 1 ♀ (MFPC) “ China-India border: Upper Siang 1850 m 04 – 10. AUG. 2013 Takeshi Maeda-leg. ”; 1 ♀ (ZMPC) “ China-India border: Upper Siang 1850 m 04 – 10. AUG. 2013 Takeshi Maeda-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB59308E9CA8224FAE4FB3A.taxon	description	Description. Male. Body shape elongated ovoid, weakly convex. Color. Generally dark reddish brown to black; shiny, with strong bronze or greenish luster; elytra with a yellow, narrow to broad zigzag band extending from primary costa 5 to 1; antennae reddish or yellowish brown, club darker; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margin straight and moderately convergent anteriad, anterior margin slightly convex, anterior corner obsolete and rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture straight, distinct. Anterior third of frons punctate as clypeus, other portions including vertex with moderately dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club equally as long as combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, curved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly to moderately excurved; with dense and large punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with extensive, irregular and large punctures, interstice II with a secondary stria, all intervals with moderately dense punctures, which are slightly or distinctly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of dense and moderately long setae from base to posterior margin, often partly abraded. Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse striation, sparse only on the greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually elongated, denser laterad, frequently confluent at the lateralmost area, ventrite 6 very densely and finely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres distinctly thickened, each tarsomere 5 with an acute internomedial denticle, that of protarsomere 5 largest, blunt and situated proximally, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of sparse and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 20 E – H. Female. Length of antennal club approximately 0.8 times as long as combined length of antennomeres 1 – 6. Lateral margin of pronotum more excurved. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, that of protarsomere situated at middle and acute, basal denticle of metatarsomere 5 absent; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw as long as its upper one and the lower margin not convex. Measurement. Body length: 13.2 – 14.1 mm in males and 13.2 – 13.9 mm in females, greatest width: 7.5 – 7.8 mm in males and 7.5 – 7.9 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB59308E9CA8224FAE4FB3A.taxon	diagnosis	Differential diagnosis. The parameres of Anomala maedai sp. nov. are somewhat similar to those of A. luoyuensis sp. nov. But the upper branch of the left paramere is thinner. The lower branches of the parameres are broad, with only the internoapical angle protruding (the whole lower branches narrowed and rather long in A. luoyuensis sp. nov.). It is also different from other members of the A. flavonotata - subgroup by internomedial denticle of protarsomere 5, which is situated proximally in males, whereas it is situated at the middle in other species.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB59308E9CA8224FAE4FB3A.taxon	etymology	Etymology. This species is named after Takeshi Maeda (Ibaraki, Japan) who collected the type series.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB39305E9CA85D0FE55FEDA.taxon	description	(Figs 4 F, 7 A – D, 11 C, 21 A – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB39305E9CA85D0FE55FEDA.taxon	materials_examined	Type material (60 ♂♂, 33 ♀♀). Holotype: ♂ (SCAU) “ CHINA: Yunnan prov., Baoshan City Longyang dist., Mts. Gaoligongshan Baihualing, Hanlongzhai 25 ° 17 ′ 39.6 ″ N, 98 ° 48 ′ 10.87 ″ E ~ 25 ° 18 ′ 28.46 ″ N, 98 ° 47 ′ 32.13 ″ E 2020. VI. 20 – 23, 1400 – 1900 m, Lu Qiu lgt. ” Paratypes: 8 ♂♂, 7 ♀♀ (ZMPC) “ CHINA: Yunnan prov., Baoshan City Longyang dist., Mts. Gaoligongshan Baihualing, Hanlongzhai 25 ° 17 ′ 39.6 ″ N, 98 ° 48 ′ 10.87 ″ E ~ 25 ° 18 ′ 28.46 ″ N, 98 ° 47 ′ 32.13 ″ E 2020. VI. 20 – 23, 1400 – 1900 m, Lu Qiu lgt. ”; 7 ♂♂, 1 ♀ (ZMPC) “ CHINA: Yunnan, Baoshan, Mts. Gaoligong, Baihualing, 1800 m 2020. V. 10 - 15 at light Yu-Chen Zheng leg. ”; 9 ♂♂ (ZMPC) “ CHINA: Yunnan, Baoshan, Mts. Gaoligong, Baihualing, 2000 m 2020. V. 17 at light Yu-Chen Zheng leg. ”; 4 ♂♂, 5 ♀♀ (CCPC) “ CHINA, Yunnan, Baoshan, Tengchong, Wuhe Town, Xiaodifang vill. 2118 m 2020. VII. 22 - 26 mercury lamp Leg. Y. - H. Li CCCC ”; 4 ♂♂, 3 ♀♀ (ZMPC) “ CHINA, Yunnan, Baoshan, Tengchong, Wuhe Town, Xiaodifang vill. 2118 m 2020. VII. 22 - 26 mercury lamp Leg. Y. - H. Li CCCC ”; 1 ♂, 1 ♀ (ZMPC) “ CHINA: Yunnan, Nujiang pref., Gongshan Co., Dulongjiang Tp. near Maku, 1862 m 27.678995 ° N, 98.299091 ° E 2023. VII. 19, Zhi-Chao Zhang leg. ”; 1 ♂ (ZMPC) “ Yunnan, Nujiang, Dulongjiang, Maku Village, Yueliang Waterfall 2019. VIII. 22, 1172.64 m 27.687347 ° N 98.276295 ° E Zi-Hao Sheng leg. (at light) ”; 4 ♂♂ (CCPC) CHINA: Yunnan, Nujiang Pref. Gongshan Co., Dulongjiang Tp., Sanxiang 1470 m 2009. VI. 5 Xiao-Yu Zhu leg. ”; 4 ♂♂, 5 ♀♀ (CZPC) “ CHINA, Yunnan, Fugong co. border to Myanmar N Gaoligongshan 2000 m vii. 2020, local collector leg ”; 2 ♂♂, 1 ♀ (ZMPC) “ MYANMAR: Kachin Area Nanse Town [Nase, in Kawnglangphu] (N of Gaoligongshan) 2000 m, 6 – 7.2020 ”; 1 ♂ (CZPC) “ Myanmar, Kechin state, Nanse vill., 06. 2018 h = 1500 m ”; 2 ♂♂ (CZPC) “ NE - MYANMAR Kachin state, Nanse vill. 1500 - 2000 m, vi. 2018 local collector lgt. ”; 1 ♀ (MFPC) “ nr. Putao N. Kachin BURMA AUG. 1999 ”; 1 ♂ (MFPC) “ E. Kachin N. MYANMAR JUN. 2001 ”; 3 ♂♂, 1 ♀ (MFPC) “ N. MYANMAR E. Kachin county Chudu-Razi [in Kawnglangphu] 2006. VI ”; 1 ♀ (MFPC), “ N. MYANMAR E. Kachin county Chudu-Razi 2007. VI ”; 1 ♀ (MFPC) “ N. MYANMAR E. Kachin county Chudu-Razi 2007. IX ”; 1 ♂, 1 ♀ (MFPC) “ 2700 m Dehtingzen N. E. Kachin MYANMAR 10 – 25. JUN. 2000 ”; 4 ♂♂, 4 ♀♀ (MFPC) “ 1800 m nr. China N. E. Kachin N. MYANMAR SEP. 2000 ”; 2 ♂♂, 1 ♀ (MFPC) “ Chudu Razi NE. Kachin stat. N. MYANMAR AUG. 2005 ”; 1 ♂ (MFNB) “ Kambaiti 7000 ’ 22.06.34. // Anomala flavonotata Arr. [Ohaus’s handwriting] ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB39305E9CA85D0FE55FEDA.taxon	description	Description. Male. Body shape elongated ovoid, moderately convex. Color. Generally reddish brown to black, sometimes elytra distinctly lighter; shiny, with weak greenish luster; elytra with a yellow, narrow to very broad zigzag band extending from primary costa 5 to 1, rarely from 4 to 1; antennae yellowish or reddish brown, club darker; protibial teeth reddish brown only at extremity; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.4; subtrapezoidal, lateral margins straight and moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture slightly bisinuate, distinct. An inverted triangular area on anterior third of frons punctate as clypeus, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third or posterior third, curved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea strongly impressed. Disc with moderately dense and small punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly excurved; with moderately dense and minute punctures. Elytra. Intervals slightly elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with irregularly doubled secondary stria, interstice II with a secondary stria, all intervals with moderately dense punctures, which are distinctly smaller than strial ones. Humeral protuberance moderately prominent, apical protberance strongly prominent. Lateral carina expanded from base to apical two fifths. Epipleuron with a row of dense and moderately long setae from base to near apex, sometimes partly abraded. Abdomen. Propygidium with very dense, transverse and fine striation. Pygidium with dense and transverse striation, sparse around the greatest tumidity; with some long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 3 strongly carinate laterally, ventrite 3 slightly weaker; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and sparse medially, gradually more transverse and denser laterad, frequently confluent at the lateralmost area, ventrite 6 moderately densely rugostriolate; ventrites 2 – 5 each with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and rather long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 5.0. Legs. Protibia bidentate, both teeth distinct; apical tooth extending to middle of protarsomere 2, preapical tooth acute. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches two times wider than their upper branches, lower branch longer in protarsal claw and equally as long as upper branch in mesotarsal claw; lower margin of lower branch of inner protarsal claw convex distally next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 small, metatarsomere 5 with an additional triangular basal denticle. Ventral surface of mesofemur with a transverse row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with a transverse row of sparse and long setae laterally, submedial row scarcely bearing seta; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 21 A – H. Female. The elytral band normally narrow, usually broken up at middle or into three small spots on each elytron. Length of antennal club approximately 0.7 times as long as combined length of antennomeres 1 – 6. Lateral margin of pronotum more excurved. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to middle of protarsomere 3, preapical tooth acute; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, that of metatarsomere largest, basal denticle of metatarsomere 5 slightly weaker; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw as long as its upper one and the lower margin not convex. Measurement. Body length: 12.5 – 14.8 mm in males and 13.5 – 15.1 mm in females, greatest width: 7.0 – 8.8 mm in males and 7.6 – 9.0 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB39305E9CA85D0FE55FEDA.taxon	diagnosis	Differential diagnosis. Anomala suavis sp. nov. is very similar to A. maedai sp. nov. However, its upper branch of the left paramere is straighter and has an acute denticle at the apex. Additionally, the ventral plate is wider between the two parameres in A. suavis sp. nov. The body of A. suavis sp. nov. is also more convex laterally.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB39305E9CA85D0FE55FEDA.taxon	etymology	Etymology. The Latin adjective suavis refers to the delicate sheen of the species, which gives it a soft and attractive appearance.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBE9304E9CA80A5FEE6FBAE.taxon	description	(Figs 7 E – F, 11 F, 16 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBE9304E9CA80A5FEE6FBAE.taxon	materials_examined	Type material (3 ♂♂, 7 ♀♀). Holotype: ♂ (MHNG) “ MYANMAR – 50 km East of Putao, Nan Thi, 950 m, 8. V. 1998, leg. S. Murzin ”. Paratypes: 4 ♀♀ (GSPC) “ MYANMAR – 50 km East of Putao, Nan Thi, 950 m, 8. V. 1998, leg. S. Murzin ”; 1 ♂, 2 ♀♀ (ZMPC) “ MYANMAR – 50 km East of Putao, Nan Thi, 950 m, 8. V. 1998, leg. S. Murzin ”; 1 ♂ (CZPC) “ MYANMAR N (Burma) 21 km E Putao, H- 550 m Nan Sa Bon vill., 1 - 5.5.98 leg. S. Murzin & V. Siniaev ”; 1 ♀ (CZPC) “ MYANMAR N (Burma) 65 km NW Putao, h- 1250 m Zi Yar Dam vill., 18. - 21.5.98 leg. S. Murzin & V. Siniaev ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBE9304E9CA80A5FEE6FBAE.taxon	description	Description. Male. Body shape ovoid, moderately convex. Color. Generally dark brown; shiny, with strong bronze luster; elytra with a yellow, broad zigzag band extending from primary costa 5 to 1; antennae brown; protibial teeth reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.2; subtrapezoidal, lateral margins moderately convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface densely and coarsely rugopunctate. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior third of frons punctate as clypeus, other portions with dense and large punctures, slightly sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.6. Length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third, slightly incurved in posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse and rounded. Posterior marginal line absent, other marginal lines complete. Lateral fovea strongly indistinct. Disc with moderately dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin weakly excurved; with moderately dense and small punctures. Elytra. Intervals almost flat, only slightly elevated near apical protuberance, subsutural interstice widest; strial punctures dense and large, subsutural interstice with irregularly doubled secondary stria, interstice II with a secondary stria, all intervals with moderately dense punctures, which are distinctly smaller than strial ones. Humeral and apical protuberance moderately prominent. Lateral carina expanded from base to apical two fifths. Epipleuron without obvious setae. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with broadly rounded apex, disc with moderately dense and transverse striation, even on the greatest tumidity; with some long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites not at all carinate laterally; ventrites 2 – 5 each with transverse and incompletely annulated punctures, mostely equal in width, sparse medially, gradually denser laterad, frequently confluent at the lateralmost area, ventrite 6 minutely striolate in anterior half; ventrites 2 – 5 with a transverse and sparse row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron longitudinally striolate, with moderately dense and long setae. Ventral mesothoracic surface with dense, small and confluent punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser and shorter setae. Width of mesofemur / narrowest extremity of mesosternum approximately 6.0. Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to middle of protarsomere 2. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, lower branch of inner protarsal claw with rounded apex, longer and three times wider than its upper branch, lower branch of outer mesotarsal claw as long as and two times wider than its upper branch; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres distinctly thickened, each tarsomere 5 with an internomedial denticle, that of protarsomere 5 largest, blunt and situated proximally, that of mesotarsomere 5 rather small, and that of metatarsomere 5 acute, metatarsomere 5 without basal denticle. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; metafemur with two rows of moderately dense and short setae, one sublateral and the other submedial, usually abraded; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 16 E – H. Female. Length of antennal club approximately 0.7 times as long as combined length of antennomeres 1 – 6. Pronotum longer, lateral margin of pronotum more excurved at middle. Lateral carina of elytra slightly wide than in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to base of protarsomere 3, preapical tooth acute; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of each tarsomere 5 small, that of protarsomere 5 situated at middle, that of metatarsomere 5 largest; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower branch of protarsal claw shorter than its upper one. Measurement. Body length: 13.6 mm in males and 13.2 mm in females, greatest width: 8.1 mm in males and female.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBE9304E9CA80A5FEE6FBAE.taxon	diagnosis	Differential diagnosis. Anomala murzini sp. nov. is externally similar to species of the A. flavonotata - subgroup, but the lower branch of the inner protarsal claw is rounded at the apex (vs. acute in the A. flavonotata - subgroup) and the abdominal ventrites are not carinate laterally (vs. ventrites 1 – 3 strongly carinate). It also has a slender and trisinuate upper branch of the left paramere, which is strongly reduced in species of the A. flavonotata - subgroup. The ventral branches of the parameres are trisinuate in profile with a hook-liked apex.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBE9304E9CA80A5FEE6FBAE.taxon	etymology	Etymology. The new species is named after Sergey Murzin (Moscow, the Russian Federation), who collected the type series.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBF9306E9CA85B1FEF1FE86.taxon	description	(Figs 8 A – C, 12 A – B, 22 A – H, 25 B)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBF9306E9CA85B1FEF1FE86.taxon	materials_examined	Type material examined. Lectotype (hereby designated): ♂ (NHMUK) “ Type H. T. // Assam. W. F. Badgley. 1906 – 185. // Anomala flavovaria, Type Arrow // LECTOTYPUS Anomala flavovaria ARROW, 1917 des. Zorn, 2011 // NHMUK 015015956 ”. Additional material examined (3 ♂♂, 1 ♀). 1 ♂ (MFPC) “ N. INDIA: alt. 1700 m Nagaland state Naga hills 28. V. – 21. VI. 2016 Takeshi Maeda-leg. ”; 1 ♂ (ZMPC) “ N. INDIA: alt. 1700 m Nagaland state Naga hills 28. V. – 21. VI. 2016 Takeshi Maeda-leg. ”; 1 ♀ (MFPC) “ E. N. INDIA: Naga State Naga hills 02 – 15. JUN. 2019 ”; 1 ♂ (SMNS) “ Popilia Asam // coll. A. Greb SMNS 1996 ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBF9306E9CA85B1FEF1FE86.taxon	description	Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex. Color. Generally black, with variable yellowish areas as following: a broad and irregular lateral band of pronotum abruptly stretching inward posteriorly (rarely not so), with encircled round black area anteriorly, hypomeron (rarely black), a square marking on lateral metasternum (rarely black), each abdominal ventrite except for narrow posterior area (usually reduced to a spot on lateralmost area), propygidium except for posterior area (usually completely black), pygidium except for a narrow anterior area (usually only one or two spots on each anterior angle), and the elytral band yellow; shiny, with rather weak greenish sheen; elytra not distinctly lighter, with a narrow to broad zigzag band extending from interstice V to primary costa 1, sometimes breaking up into three spots; antenna, also distal sections of tibiae, femora and protarsus brown (usually black); setae yellowish brown, robust setae black. Head. Greatest width / length of clypeus approximately 2.3; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and broadly rounded; anterior margin weakly reflexed; surface with dense, large and coarse punctures. Frontoclypeal suture moderately incurved, distinct. Frons with very dense and large punctures, gradually sparser and smaller posteriad. Interocular distance / maximum head width approximately 0.63. Length of antennal club 0.9 times the combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third and posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with dense and large punctures. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved, apex somewhat rounded; with dense and small punctures. Elytra. Intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, annulated, subsutural interstice with an irregular secondary stria, doubled near scutellum, interstices II – IV each with a secondary stria, that of interstice III ending at middle, all intervals with moderately dense punctures, which are distinctly smaller than strial ones. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical fourth. Epipleuron with few short setae. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with moderately dense and transverse striation, not sparser on the greatest tumidity; with sparse and long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 4 strongly carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually more transverse and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron with sparse and small punctures, moderately dense and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with moderately dense and small punctures, setae slightly shorter. Metacoxa punctate as lateral metasternum but bearing sparser setae. Width of mesofemur / narrowest extremity of mesosternum approximately 5.0. Legs. Protibia bidentate, both teeth distinct; apical tooth slender and extending to anterior margin of protarsomere 2, preapical tooth acute. Inner spur inserted at the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw slightly convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Pro- and metatarsomere 5 with an acute and large internomedial denticle, mesotarsomere 5 only slightly convex internomedially. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 22 A – D. Female. Generally black, each abdominal ventrite with a yellow spot at lateral area, antennomeres 1 – 6 yellowish brown; yellow and zigzag band on each elytron formed by three small spots. Antenna shorter, length of antennal club approximately 0.7 times the combined length of antennomeres 1 – 6. Lateral carina of elytra broadest around middle, at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to base of protarsomere 3; inner spur inserted at mid-length of protibia; protarsomeres thinner, pro- and mesotarsomere with small and acute internomedial denticle; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch, lower margin of lower branch of inner protarsal claw slightly convex next to the internobasal notch;. Measurement. Body length: 13.1 – 13.4 mm in males and 12.6 mm in female, greatest width: 8.1 – 8.3 mm in males and 7.7 mm in female.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBF9306E9CA85B1FEF1FE86.taxon	diagnosis	Differential diagnosis. The shiny and black general appearance of Anomala flavovaria is resembling species of the A. flavonotata - subgroup. However, A. flavovaria has additional clear markings on thorax and abdomen as well as a strongly carinate abdominal ventrite 4. The upper branch of the right paramere reaches the middle of the whole paramere length. The upper branch of the left paramere is slender in the apical half.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBF9306E9CA85B1FEF1FE86.taxon	discussion	Remarks. The black male from Assam (figs 22 E – H) exhibits a narrower membranous area between the parameres compared to the lectotype and males from Nagaland. Its dorsal sclerotized branches are also narrower. It is unclear whether this difference is taxonomically significant without additional specimens and more precise locality data.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	description	(Figs 8 D – F, 12 C, 23 A – D)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	materials_examined	Type material. See Zorn (2005). Lectotype from Kurseong, paralectotype from Gantok. Additional material examined (20 ♂♂, 31 ♀♀). 1 ♂ (NMPC) “ NEPAL SE 28.6. - 1.7.2012 Shivalaya env. E. Kučera lgt. Nep / B / 01 ”; 1 ♀ (NHMB) “ Phul Choki 2000 m 7.6. // Nepal, 1977 W Wittmer ”; 2 ♂♂, 1 ♀ (NHMUK) “ NEPAL, 5800 ' Kathmandu Dist. Godawari, at light 5 / vi / 1983 // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♀ (NHMB) “ Kali-G. Khola Kopchepani 1600 m 18. VI. 1986 // W-Nepal Dhawalagiri C. Holzschuh ”; 1 ♂ (NHMB) “ Kali-G. Khola Kopchepani-Gasa 1600 – 2000 m 19. VI. 1986 // W-Nepal Dhawalagiri C. Holzschuh ”; 1 ♀ (NHMB) “ Kali-G. Khola Gasa-Kalopani 2000 – 2500 m 20. VI. 1986 // W-Nepal Dhawalagiri C. Holzschuh ”; 1 ♂ (NHMB) “ Kali-G. Khola Kalopani 25 / 2800 m 21 – 25. VI. 1986 // W-Nepal Dhawalagiri C. Holzschuh ”; 1 ♂ (NHMB) “ NEPAL: Kosi - # 13 Depitar 27 ° 27 ′ N / 87 ° 17 ′ E to Barabishe 27 ° 26 ′ N / 87 ° 18 ′ E 1250 - 560 m, 12. vi. 01 // NHMB Basel expedition to Nepal, 2001 ”; 1 ♂ (NMBF) “ NEPAL Likhu Khola Tal 1700 m 4. VI. 62 leg. G. Ebert // Anomala flaviventris Arr. ”; 1 ♀ (NMBF) “ Nepal, Kathmandu Valley Godavari 16 - 1800 m 1. VI. 1967 Dierl-Forster-Schacht // Anomala flaviventris Arr. det. G. Frey, 1973 ”; 1 ♀ (NMBF) “ Nepal, Kathmandu Valley Godavari 16 - 1800 m 5. VI. 1967 Dierl-Forster-Schacht // Anomala flaviventris Arr. det. G. Frey, 1973 ”; 1 ♀ (CZPC) “ NEPAL-HIMALAYA Annapurna Geb. - Lamjung H. lg. Schmidt 1994 // Chamje-Tal 10.6. 1500 - 1700 m ”; 1 ♂, 4 ♀♀ (ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu, 1900 m, 2020. VII. 29, Yu-Chen Zheng leg. ”; 5 ♀♀ (ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu, Yingbin Road 2000 m, 2020. VII. 29, Yu-Chen Zheng leg. ”; 3 ♂♂, 3 ♀♀ (ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu Town, 1970 m 27.971656 ° N, 85.966972 ° E, light trap, 2023. VII. 1 Zu-Qi Mai leg. ”; 1 ♂ (ZFMK) “ Pedong bei Darjeeling 1935 Sig. R. Oberthür Eing. Nr. 4, 1956. // Anomala flaviventris Arr. det. G. Frey, 1967 / 68 ”; 1 ♂ (MNHN) “ Pedong A. Desgodins // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂, 2 ♀♀ (MNHN) “ Phedong R. P. Desgodins // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♀ (MNHN) “ British Bootang L. Durel 1898 // Anomala flavofasciata, a. Compared with type. G. J. A. // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂ (MNHN) “ British Bootang Maria Basti 1899 // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂ (MNHN) “ British Bootang Maria Basti L. Durel ”; 1 ♀ (NMBF) “ 8272 18 // 1913 345 // Annandale Kurseong E. Himalayas alt. 4700 - 5000 ft. 28. VI- 10. // Anomala flavofasciata Arrow ”; 2 ♀♀ (CZPC) “ INDIA – SIKKIM west PEELING env., 2100 m 18. - 20.7.1997 Jan Schneider lgt. ”; 1 ♂ (CZPC) “ north INDIA-West Bengal Shingalila national park Shirikhola 18. - 28.5.1999 lgt. E. Kučera (2600 m. m) ”; 1 ♂, 2 ♀♀ (CZPC) “ north INDIA – west SIKKIM Khecheopari lake (1800 m. m) 04. - 10.6.1999 lgt. E. Kučera ”; 1 ♀ (CZPC) “ INDIA – SIKKIM Khecheopari Lake 21.6. – 26.6.2003 lgt. E. Kučera ”; 2 ♀♀ (MFPC) “ INDIA - SIKKIM Khecheopalri lake 15.6 – 22.6.2008 1800 m leg. E. Kučera ”; 1 ♀ (NMBF) “ 21 km O Wangdi Phodr. [Wangdue Phodrang] 1700 – 2000 // Nat. - Hist. Museum Basel - Bhutan Expedition 1972 // Anomala spiloptera det. G. Frey, 1973 ”; 2 ♂♂ (GSPC) “ 21 km O Wangdi Phodr. 1700 – 2000 // Nat. - Hist. Museum Basel - Bhutan Expedition 1972 // Anomala spiloptera det. G. Frey, 1973 ”.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	description	Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex. Color. Generally yellowish brown, distal and proximal sections of pro- and mesotibiae darker, the head, disc of pronotum except for a narrow lateral area, meso- and metathoracic surface, pygidium in anterior two thirds, abdominal ventrites 2 – 5 (in posterior half), metatibia and all tarsomeres reddish brown; dull, with weak iridescent sheen; elytra lighter than pronotum, with a yellowish brown, narrow to broad zigzag band extending from primary costa 4 to 1, sometimes very pale or partly interrupted; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.1; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with dense, large and coarse punctures. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance / maximum head width approximately 0.65. Length of antennal club 0.8 times the combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third and posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with dense, rather large and transverse punctures, smaller and rounded along lateral and posterior margins. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved, or semielliptical; with dense and large punctures as well as several erect and moderately long setae. Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, annulated, subsutural interstice with irregularly doubled secondary stria, interstices II – IV each with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical third. Epipleuron with a row of moderately dense and moderately long setae from base to apical two fifths, usually partly abraded. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with moderately dense and transverse striation, sparse only on the greatest tumidity; with sparse and long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites not carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron with sparse and small punctures, moderately dense and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with smaller punctures, density and length of setae as lateral area. Metacoxa punctate as lateral metasternum but bearing sparser setae. Very narrow at the extremity of mesosternum. Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to middle of protarsomere 2. Inner spur inserted shortly behind the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 smaller. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and rather long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial, also with some irregular and long setae sublaterally; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 23 A – D. Female. Generally reddish or blackish brown, lateral area of abdominal ventrites, mesofemur, marginal areas of metafemur, as well as antenna yellowish brown, protibial teeth reddish brown; elytra not paler than pronotum, zigzag band narrower, sometimes breaking up into three small spots. Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1 – 6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of protarsomere 5 distinctly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch. Measurement. Body length: 12.6 – 14.2 mm in males and 12.9 – 14.4 mm in females, greatest width: 7.1 – 7.8 mm in males and 6.9 – 8.2 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	diagnosis	Differential diagnosis. The ventral surface of the parameres is gradually changing from sclerotized to membranous in the apical half in Anomala flavofasciata. Compared to similar species, the parameres are also most slender.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	discussion	Remarks. The record of A. spiloptera Burmeister, 1855 from Wangdi Phodrong, Bhutan, published by Frey (1975), was shown to be A. flavofasciata after examination of the original specimens (NMBF and GSPC, with Frey’s label).	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFBD9300E9CA8059FB6AF8D6.taxon	distribution	Distribution. China (Xizang) (new record); India (West Bengal, Sikkim); Nepal; Bhutan.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB89302E9CA81A0FC6AFA96.taxon	description	(Figs 9 A – B, 12 D, 23 E – H)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB89302E9CA81A0FC6AFA96.taxon	materials_examined	Type material (14 ♂♂, 21 ♀♀). Holotype: ♂ (SCAU) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ” Paratypes: 3 ♂♂, 4 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 2 ♂♂, 5 ♀♀ (ZYPC) “ CHINA: TIBET, Nyingchi City, Beibeng Township, Gelin Vill., 1753 m, 29.214870 ° N, 95.169272 ° E, 18 – 30. VII. 2023 (at light), Yao-Nan Zhang leg. ”; 1 ♂, 4 ♀♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Beibeng, Gelin Vill., 1768 m, 29.21315309 ° N, 95.17111894 ° E, 2023. VII. 13 - 14, Hao-Yi Liu leg. ”; 1 ♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., 96 k Marangkang vill., 1305 m, 29.56287354 ° N, 95.46563172 ° E, 2023. VII. 17, Hao-Yi Liu leg. ”; 1 ♂, 4 ♀♀ (ZMPC) “ CHINA: Tibet, Nyingchi City, Medog County, 1143 m 29 ° 19 ′ 22.62 ″ N, 95 ° 20 ′ 1.65 ″ E 26. VII. 2022 (at light) Chuan-Tao Qu leg. ”; 1 ♂ (ZMPC) “ CHINA: Tibet, Nyingchi City, Medog County, 1143 m 29 ° 19 ′ 22.62 ″ N, 95 ° 20 ′ 1.65 ″ E 20. VII. 2022 (at light) Chuan-Tao Qu leg.; 1 ♀ (ZMPC) “ CHINA: TIBET, Nyingchi City, Medog Co., Ribeng Valley Bridge, 29.62527885 ° N, 95.48554466 ° E, 1681 m, 2023. VII. 9, Xu-Hong-Yi Zheng ”; 2 ♂♂, 1 ♀ (MFPC) “ China-India border: alt. 1700 m Lower Subansiri 18. V. – 01. VI. 2017 Takeshi Maeda-leg. ”; 1 ♀ (MFPC) “ China-India border: Lower Subansiri 9 – 19. AUG. 2018 Takeshi Maeda-leg. ”; 3 ♂♂ (MFPC) “ China-India border: W. Siang 03. JULY. 2011 Takeshi MAEDA-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB89302E9CA81A0FC6AFA96.taxon	description	Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex. Color. Generally yellowish brown, the head, antennal club, disc of pronotum except for a narrow lateral area, scutellum, mesosternum, metasternum (except for a square marking laterally), pygidium in anterior half, abdominal ventrites 1 – 5 (except for lateral portion), meso- and metatibiae as well as all tarsomeres dark brown, protibia only blackish brown at dorsal face, protibial teeth reddish brown; in lighter form, abdominal ventrites 2 – 5 darkened only in posterior half, pygidium only darkened at narrow anterior line, pro- and mesotibia yellowish brown; dull, with weak iridescent sheen; elytra lighter than pronotum, with a yellowish brown, narrow to broad zigzag band extending from primary costa 5 to 1, rarely from interstice V, sometimes very pale or interrupted; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.0; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface dense, large and coarse punctures. Frontoclypeal suture moderately incurved, distinct. An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance / maximum head width approximately 0.65. Length of antennal club 0.9 times the combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third, slight concave in posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line absent, other marginal lines complete. Lateral fovea indistinct. Disc with dense, rather large and transverse punctures, smaller and rounded along lateral and posterior margins. Lateral margin with several long setae. Scutellum triangular, lateral margin moderately excurved, or semielliptical; with dense and small punctures, as well as a few erect and short setae. Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, annulated, subsutural interstice with irregularly doubled secondary stria, interstices II – IV each with a somewhat irregular secondary stria, all intervals with moderately dense punctures, which are slightly or distinctly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical third. Epipleuron with a row of moderately dense and short setae from base to apical two fifths, usually partly abraded. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium with moderately dense and transverse striation, sparse only on the greatest tumidity; with sparse and long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites not carinate laterally; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron with sparse and small punctures, moderately dense and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with moderately dense and small punctures, setae slightly shorter. Metacoxa punctate as lateral metasternum but bearing sparser setae. Width of mesofemur / narrowest extremity of mesosternum approximately 8.0. Legs. Protibia bidentate, both teeth distinct; apical tooth slender and extending to anterior margin of protarsomere 2, preapical tooth acute. Inner spur inserted at the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Each tarsomere 5 with an acute internomedial denticle, that of mesotarsomere 5 smaller. Ventral surface of mesofemur with a row of dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional dense and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one lateral and the other submedial, also with some irregular and long setae sublaterally; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 23 E – H. Female. Usually entirely black, a vague lateral area of pronotum orange brown, each abdominal ventrite with a yellow spot at lateralmost area, antenna yellowish brown; elytra not paler than pronotum, zigzag band usually narrower or breaking up into three small spots. Rarely generally with similar color pattern like male. Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1 – 6. Lateral carina of elytra at most 1.5 times as wide as in male. Pygidium more triangular and flat preapically; ultimate abdominal ventrite 1.5 times as long as that in male medially. Apical protibial tooth spatulate, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial denticle of protarsomere 5 distinctly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch. Measurement. Body length: 14.6 – 15.3 mm in males and 14.3 – 16.0 mm in females, greatest width: 7.8 – 8.9 mm in males and 7.5 – 9.4 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB89302E9CA81A0FC6AFA96.taxon	diagnosis	Differential diagnosis. Anomala fenestrata sp. nov. is very similar to A. flavofasciata but larger. The parameres are stouter and have larger membranous areas laterally and ventrally. The ventral surface of the parameres is becoming abruptly membranous in the apical half.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB89302E9CA81A0FC6AFA96.taxon	etymology	Etymology. The name is derived from the Latin adjective fenestratus (having a window), alluding to the broad, translucent membranous structure present in the sclerotized parameres.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB9933CE9CA8464FC64F822.taxon	description	(Figs 9 C – D, 12 E, 13 A, 13 C, 24 A – D, 25 C, 25 E, 25 K – N)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB9933CE9CA8464FC64F822.taxon	materials_examined	Type material examined (1 ♂, 2 ♀♀). Lectotype (hereby designated): 1 ♂ (NHMUK) “ Type // Figured for “ Fauna of India. ” // Kurseong, Sikkim R. P. Verschraeghen. 1911 - 218. // Kurseong Inde Verschraeghen 1904 // Anomala flaviventris, Arrow Type ”. Paralectotypes: 2 ♀♀ (NHMUK) “ IND. MUS. 3091 16 // 1913 - 345 // Kurseong alt. 5000 ft E. Himalayas Lynch, c. // NHMUK 015015929 – 30 ”. Additional material examined (88 ♂♂, 54 ♀♀). 1 ♂, 1 ♀ (NHMUK) “ Gopaldhara, Darjeeling. 1914. – 414. // NHMUK 015015931, NHMUK 015015945 ”; 1 ♂ (NHMUK) “ Gopaldhara, Darjeeling. 1914. – 414. // Gopaldhara, Br. Sikkim. W. K. Webb // NHMUK 015015932 ”; 1 ♂ (NHMUK) “ Figured for “ Fauna of India. ” // Gopaldhara, Rungbong Vall., Sikkim. H. Stevens. 1916 – 218 // NHMUK 015015933 ”; 5 ♂♂, 2 ♀♀ (NHMUK) “ Gopaldhara, Rungbong Vall., Sikkim. H. Stevens. 1916 – 218 // NHMUK 015015935 – 37, NHMUK 015015941 – 44 ”; 1 ♂ (NHMUK) “ Gopaldhara, Br. Sikkim. W. K. Webb // Andrewes Bequest. B. M. 1922 - 221 // Anomala flaviventris Arr. // NHMUK 015015938 ”; 1 ♀ (NHMUK) “ Gopaldhara 1911 // Andrewes Bequest. B. M. 1922 - 221 // NHMUK 015015940 ”; 3 ♂♂ (NHMUK) “ Gopaldhara, Rungbong Vall., Sikkim H. Stevens ”; 3 ♀♀ (NHMUK) “ Sikkim Gopaldhara, Rungbong Vall. H. Stevens 1922 - 307 ”; 1 ♀ (MNHN) “ Gopaldhara, Br. Sikkim. W. K. Webb // Anomala flaviventris Arrow // MUSÉUM PARIS 1952 COLL. R. OBERTHÜR ”; 1 ♂, 3 ♀♀ (MNHN) “ Himalaya Kurseong 13.6. India // Anomala flaviventris arr. Det. Tesař ”; 1 ♂ (NMBF) “ Himalaya Kurseong 7.6. India // flaviventris Arrow ”; 1 ♀ (NMBF) “ Himalaya Kurseong 13.6. India ”; 2 ♂♂, 2 ♀♀ (MNHN) “ Darjeeling Inde britan. Coll. J. Clermont // MUSÉUM PARIS 1944 Coll. E. BENDERITTER ”; 1 ♀ (MNHN) “ Inde // MUSÉUM PARIS 1930 Coll. SICARD // anomala sp. ”; 1 ♀ (MNHN) “ Sikkim // MUSÉUM PARIS 1930 Coll. SICARD ”; 3 ♂♂, 2 ♀♀ (ZMPC) “ NEPAL: N of Kathamandu, W-Shivapuri Nagarjun N. P. 1670 m 2025. VII. 5 - 6 light trap ”; 1 ♂ (NHMB) “ C. NEPAL 1992 GODAVARI 11. vi. 1500 - 1700 m Leg. Jiří Moravec ”; 1 ♂, 1 ♀ (NHMUK) “ NEPAL, 5800 ' Kathmandu Dist. Godawari, at light 5 / vi / 1983 // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♂ (NHMUK) “ NEPAL: 1500 m Kathmandu Secondary veg. 15 June [19] 84 Lt. Col. M. G. Allen // BMNH {E} 1983 - 254 M. G. Allen ”; 1 ♂ (NHMB) “ Sheopuri 27.6. N Kathmandu 2400 - 2500 m // O Nepal 1980 W. Wittmer // Anomala flaviventris Arr. det. Sabatinelli 1981 ”; 4 ♂♂ (GSPC) “ NEPAL, Kathmandu val., Mulkharka, 20.5.91 1600 m leg. Sabatinelli ”; 2 ♂♂ (ZMPC) “ NEPAL, Kathmandu val., Mulkharka, 20.5.91 1600 m leg. Sabatinelli ”; 1 ♂, 4 ♀♀ (NHMUK) “ NEPAL: 6800 ', Kathmandu Distr., Kakani, 1 – 2. vi. 1983 // At light // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♂, 1 ♀ (ZMPC) “ NEPAL: 6800 ', Kathmandu Distr., Kakani, 1 – 2. vi. 1983 // At light // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♀ (NHMUK) “ NEPAL: 6800 ’, Kathmandu Dist., Kakani, // 15. vii. 1984 // Lt. Col. M. G. Allen Brit. Mus. 1983 - 254 ”; 3 ♂♂, 1 ♀ (NHMUK) “ NEPAL, 6800 ’ Kathmandu Dist. Kakani 15 / vi / 1983 Malaise Trap // M. J. D. Brendell BMNH {E} 1983 - 222 ”; 2 ♂♂, 2 ♀♀ (NHMUK) “ NEPAL, Kakani 2070 m, 3 / vii / 1983 Pine & Oak Forest Coll, M G Allen // BMNH {E} 1983 - 254 M. G. Allen ”; 2 ♂♂, 3 ♀♀ (ZMPC) “ NEPAL: Bagmati zone, Lalitpur dist., Bhardev Village, 1850 m, at light 2025. VI. 29 ”; 3 ♂♂ (GSPC) “ NEPAL, Helambu Pati Bhanijang, 19.5. [19] 91 1750 m leg. Sabatinelli ”; 3 ♂♂ (ZMPC) “ NEPAL, Helambu Pati Bhanijang, 19.5. [19] 91 1750 m leg. Sabatinelli ”; 10 ♂♂, 5 ♀♀ (NHMB) “ Nuwakot Pati Bhanjyang 1900 m 16 - 18. VI. 89 // C-Nepal Bagmati C. Holzschuh ”; 1 ♂ (NHMB) “ Nuwakot Gul Bh. - Pati Bhanjyang 2300 - 1900 m 16. VI. 89 // Nepal Bagmati M. Brancucci ”; 1 ♀ (NHMB) “ Dapkakharka-Manegero 2100 - 2500 m 12. VI. 89 M. Brancucci // Nepal Bagmati Sindhupalchok ”; 1 ♂ (NHMB) “ Tadzhiskistan Ramit Schutzgebiet // 4 - 5. V. 90 1250 m W. Wittmer ”; 7 ♂♂, 2 ♀♀ (NHMB) “ Nagarkot 8. VI. 1990 // Nepal G. Sabatinelli ”; 4 ♂♂, 2 ♀♀ (GSPC) “ NEPAL, Bagmati distr. Nagarkot, 1800 m 3.6.90 leg. Sabatinelli ”; 1 ♂, 1 ♀ (CZPC) “ Nepal nr. Nagarcot 2000 m VII. 95 O. Ammosov leg. ”; 5 ♂♂ (NHMB), 1 ♂ (GSPC) “ Manigow 10. VI. 1200 - 1900 m // Nepal 1978 Bhakta B, Ch. // Anomala flaviventris Arr. det. Sabatinelli 1981 ”; 1 ♂ (NMBF) “ Nepal Prov. Nr. 3 East Jubing 1600 m 9. V. 1964 leg. v. Dierl // Anomala flaviventris Arr. det. G. Frey, 1973 ”; 4 ♂♂, 3 ♀♀ (CCPC) “ CHINA, Xizang (Tibet), Jilongxian, Xinjiangcun, 1785 m, 2019, VI- 25, Leg. X. - D. YANG CCCC ”; 3 ♂♂, 2 ♀♀ (ZMPC) “ CHINA, Xizang (Tibet), Jilongxian, Xinjiangcun, 1785 m, 2019, VI- 25, Leg. X. - D. YANG CCCC ”; 1 ♂ (ZMPC) “ China: Tibet, Shigatse, Nyalam co. [Nielamu], Zhangmu Town, 1970 m 27.971656 ° N, 85.966972 ° E, light trap, 2023. VII. 1 Zu-Qi Mai leg. ”; 3 ♂♂, 3 ♀♀ (ZMPC) “ China: Tibet, Shigatse, Gyirong co. [Jilong], Gyirong Town (port), 1879 m 28.289865 ° N, 85.347493 ° E, light trap, 2023. VII. 5 Zu-Qi Mai leg. ”; 3 ♂♂, 4 ♀♀ (ZMPC) “ China: Tibet, Shigatse, Gyirong co., Gyirong Town (port), 1879 m 28.289865 ° N, 85.347493 ° E, light trap, 2023. VII. 7 - 8 Zu-Qi Mai leg. ” Literature record. Siliguri (Arrow 1912).	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB9933CE9CA8464FC64F822.taxon	description	Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex. Color. Generally yellowish brown, clypeus, disc of pronotum except for a narrow to broad lateral area, elytra, small central area of metasternum, anterior part of pygidium (varying from a narrow line to the whole two thirds), abdominal ventrites (in posterior half), metatibia and all tarsomeres dark brown; frons and vertex blackish brown; dull, with weak greenish sheen; elytra usually distinctly lighter than pronotum, with a yellow, narrow to very broad and oblique band extending from primary costa 4 to 1, sometimes only area behind the band dark; antennae yellowish or reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.1; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with dense, large and coarse punctures. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance / maximum head width approximately 0.57. Antenna long, length of antennal club 1.1 times the combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third, slightly incurved in posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea indistinct. Disc with moderately dense and large punctures. Lateral margin with several moderately long setae. Scutellum triangular, lateral margin strongly excurved; with dense and large punctures. Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with irregularly doubled secondary stria, interstices II and III each with a secondary stria, all intervals with dense punctures, which are distinctly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical two fifths. Epipleuron with a row of moderately dense and short setae from base to apical two fifths, usually partly abraded. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium broad and strongly tumid, with broadly rounded apex, disc with dense, transverse and incompletely annulated punctures, gradually confluent near the greatest tumidity; with dense and long setae preapically and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 4 strongly carinate laterally, ventrite 4 more or less flat, without two separated protuberances medially, not particularly concave between ventrites 4, 5 and 6 in profile; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron with sparse and small punctures, sparse and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with sparse and small punctures, setae short. Metacoxae punctate as lateral metasternum but bearing sparser setae. Very narrow at the extremity of mesosternum. Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to middle of protarsomere 2. Inner spur inserted at the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres thin, pro- and mesotarsomere 5 each with a small internomedial protuberance slightly behind middle, mesotarsomere 5 with a large and acute internomedial denticle. Ventral surface of mesofemur with a row of moderately dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional sparse and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one sublateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 24 A – D. Female. Elytral band broad to very broad, sometimes the whole anterior two thirds of elytra yellow; abdominal ventrites and pygidium (except for a narrow anterior line) yellowish brown, metatibia and all tarsomeres darker than in male. Lateral margin of pronotum more excurved. Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1 – 6. Pygidium more triangular, less tumid preapically; ultimate abdominal ventrite 2.0 times as long as that in male medially, ventrites flat in profile, without protuberance. Apical protibial tooth longer and apex rounded, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial protuberance of protarsomere 5 absent, that of metatarsomere slightly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch. Measurement. Body length: 11.3 – 13.2 mm in males and 11.3 – 13.0 mm in females, greatest width: 6.1 – 7.0 mm in males and 6.0 – 7.4 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB9933CE9CA8464FC64F822.taxon	diagnosis	Differential diagnosis. Anomala flaviventris is very similar to A. brancuccii Sabatinelli, 1991. We found them sympatric in Godawari, Shivapuri Nagarjun, Mulkharka and Gopaldhara. The lower branches of the parameres are slightly longer than or as long as the upper branches in A. flaviventris (smaller and shorter than the upper branches in A. brancuccii). The upper branches have nearly straight anterior margins in A. flaviventris, whereas they are convex in A. brancuccii. A more distinct external feature is the two protuberances present on abdominal ventrite 4 in male of A. brancuccii, which are absent in male of A. flaviventris. The abdominal ventrites of female A. brancuccii are also more convex in profile than in A. flaviventris. The protibia, protarsomeres and protarsal claws in male of A. brancuccii are longer, and the protarsal claws are also more curved than in males of A. flaviventris.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFFB9933CE9CA8464FC64F822.taxon	distribution	Distribution. China (Xizang); India (West Bengal, Sikkim); Nepal.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	description	(Figs 9 E – F, 12 F, 13 B, 13 D, 24 E – H, 25 D, 25 F, 25 O – Q)	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	materials_examined	Type material examined (5 ♂♂, 6 ♀♀). Paratypes: 2 ♂♂, 5 ♀♀ (NHMB) “ Lambobagar Gao 1400 m 28. - 31.5. // O. Nepal 1980 W. Wittmer // PARATYPUS Anomala brancuccii n. sp. det. Sabatinelli 1990 ”; 1 ♀ (NHMB) “ Hatiya-Lambobagar Gao 2.6. 1550 - 1000 m // O. Nepal 1980 W. Wittmer // PARATYPUS Anomala brancuccii n. sp. det. Sabatinelli 1990 ”; 2 ♂♂ (GSPC) “ NEPAL, Kasaki distr. Dhampus, 1650 m 24. V. 90 leg. Sabatinelli // PARATYPUS Anomala brancuccii n. sp. det. Sabatinelli 1990 ”; 1 ♂ (ZMPC ex GSPC) “ Dandapakhar [in Dolakha] 1700 m 7.6.1976 // Nepal, W. Wittmer C. Baroni U. 1976 // PARATYPUS Anomala brancuccii n. sp det. Sabatinelli 1990 // SAB ”; Additional material examined (62 ♂♂, 68 ♀♀). 1 ♂ (NMBF) “ Himalaya Expedition India Kumaon- Himalaya Distr. NainiTal Bhim Tal 1500 m 18.6.1971 leg. de Freina // Anomala flaviventris Arr. det. G. Frey, 1967 / 68 ”; 1 ♂ (NMBF) “ Himalaya Expedition India Kumaon-Himalaya Distr. NainiTal Bhim Tal 1500 m 27.6.1971 leg. de Freina // Anomala flaviventris Arr. det. G. Frey, 1967 / 68 ”; 1 ♂ (NMBF) “ Himalaya Expedition India Kumaon-Himalaya Distr. NainiTal Bhim Tal 1500 m 07.6.1971 leg. de Freina // Anomala flaviventris Arr. det. G. Frey, 1967 / 68 ”; 1 ♂ (NMBF) “ Himalaya Expedition India Kumaon-Himalaya Distr. NainiTal Bhim Tal 1500 m 22.6.1971 leg. de Freina ”; 2 ♂♂ (NMBF) “ Himalaya Expedition India Kumaon-Himalaya Distr. NainiTal Bhim Tal 1500 m 20.6.1971 leg. de Freina // Anomala flaviventris Arr. det. G. Frey, 1967 / 68 ”; 2 ♂♂, 5 ♀♀ (CZPC) “ Nepal, Ghorka-Dist., Ganesh Himal Abuthum Lekh Burang 1500 m 22. V. 1996, leg. J. Kulbe ”; 4 ♂♂, 7 ♀♀ (CZPC) “ Nepal S-Ganesh Himal Ankhu Khola Burang 1500 m 22.5.1996 lg. Ahrens, Kulbe, Rulik ”; 2 ♂♂, 1 ♀ (CZPC) “ Ost-Nepal Rolwaling Himal // Dolakha 1000 m 12.05.2000 leg. A. Kleeberg ”; 2 ♂♂, 2 ♀♀ (ZFMK) “ Nepal: Ganesh Himal; Ankhu Khola valley, Borang village, 1720 m, 19. vi. 2000 leg. J. Schmidt ”; 1 ♀ (HNHM) “ NEPÁL, Ganesh Himal 3 km W of Godland, 2800 m 2004. VII. 18. Leg: Szécsényi Lajos ”; 1 ♂ (ZFMK) “ NEPAL: Annapurna Himal, Modi Khola vall. Dhinnu vill. 2700 m 15. VI. 2000 lg. Schmidt ”; 1 ♀ (CZPC) “ NEPAL-HIMALAYA Annapurna Geb. - Lamjung H. lg. Schmidt 1994 // zw. Syange und Chamje 1300 m 12.6. ”; 1 ♀ (CZPC) “ NEPAL, Annapurna Himal, near Talopani 1300 m, 03, 20. VI. 1996 leg. Gy. M. Lázló and G. Ronkay ”; 3 ♀♀ (ZFMK) “ Nepal: Dhaulagiri Himal, s- slope; Jhi N Beni, 1750 m, 16. vi. 1998, leg. J. Schmidt ”; 1 ♂, 1 ♀ (MTD) “ NEPAL, Dhaulagiri-Himal, e-slope, Jhi (upp. Beni), N 28 ° 25 ′; E 83 ° 30 ′, 1500 m, 16.06.1998, Jäger ”; 1 ♂ (MSPC) “ NEPAL, 10.6.1990 Langtang Nat. Park. Syabru, 2000 m S. Bily leg. ”; 1 ♂, 2 ♀♀ (NMPC) “ NEPAL SE 28.6. - 1.7.2012 Shivalaya env. E. Kučera lgt. Nep / B / 01 ”; 1 ♂ (ZMPC) “ NEPAL: Bagmati zone, Rasuwa distr., Thulogaun village 28 ° 01 ′ 39.74 ″ N, 85 ° 10 ′ 41.57 ″ E 1436 m 2024. VIII. 16 at light ”; 1 ♀ (NHMB) “ C-NEPAL 19 - 20.7.2000 Bagmati prov., NAGARJUN forest 27.45 N 85.17 E; 1387 m [GPS] Jan Farkač lgt. (at light) // NEPAL Expedition, Jan Farkač, David Král & Jan Schneider, 2000 ”; 1 ♀ (NHMB) “ Nepal Bagmati Sindhupalchok Dubhachaur-Sarmatang // 1600 - 2500 m 3. VI. 89 M. Brancucci ”; 2 ♂♂, 1 ♀ (GSPC) “ NEPAL, Kathmandu val., Mulkharka, 20.5. [19] 91 1600 m leg. Sabatinelli ”; 2 ♂♂ (ZMPC) “ NEPAL, Kathmandu val., Mulkharka, 20.5. [19] 91 1600 m leg. Sabatinelli ”; 4 ♂♂, 20 ♀♀ (NHMB) “ Nepal Kathmandu V. Balaju 1400 m // 18. V. 1989 M. Brancucci ”; 1 ♀ (NMBF) “ Nepal, Kathmandu Valley Godavari 16 - 1800 m 8. VI. 1967 Dierl-Forster-Schacht ”; 2 ♂♂ (NMBF) “ Staatsslg. München Nepal Kathmandu Valley Godavari 1600 m 31. V. 1964 leg. Loffler (9) ”; 1 ♂ (NHMB) “ C. NEPAL 1992 GODAVARI 11. vi. 1500 - 1700 m Leg. Jiří Moravec ”; 1 ♂, 1 ♀ (NHMB) “ C. NEPAL 1992 GODAVARI 16 - 18. v. 1500 - 1900 m Leg. Jiří Moravec ”; 1 ♀ (CZPC) “ NEPAL Centr., Bagmati Zone, Kathmandu valley, Lalitpur Distr., Godawari Phulchoki, 2200 - 2700 m, 1 - 7. vi. 1996, P. Cechovsky lgt. ”; 4 ♂♂ (MSPC) “ NEPAL 18. V. 1989 Kathmandu valley GODAVARI C. Holzschuh leg. ”; 7 ♂♂, 5 ♀♀ (ZMPC) “ NEPAL: N of Kathamandu, W-Shivapuri Nagarjun N. P. 1670 m 2025. VII. 5 - 6 light trap ”; 2 ♂♂ (NHMUK) “ NEPAL: 1550 M Godaveri, 6. v. 1984 Mixed forest. Lt. Col M. G. Allen Brit Mus 1983 - 254 ”; 1 ♂, 1 ♀ (NHMUK) “ NEPAL, 5800 ' Kathmandu Dist. Godawari, at light 5 / vi / 1983 // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♂ (NHMUK) “ NEPAL: 4500 ', Kathmandu, British Embassy, 20. v. – 23. vi. 1983 // At light // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♂, 1 ♀ (NHMUK) “ NEPAL: 4500 ', Kathmandu, Gokarna, 7. vi. 1983 // At light // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♀ (NHMUK) “ NEPAL, 4500 ' Kathmandu. 31 / vi / 1983 At MV light // M. J. D. Brendell. BMNH {E} 1983 - 222. ”; 1 ♂, 1 ♀ (NHMB) “ C. NEPAL 1992 GORKHA 27 - 30. v. Dorandi Khola Leg. Jiří Moravec ”; 1 ♂, 5 ♀♀ (NHMB) “ Parahang 1700 m 9. VI. 89 M. Brancucci // Nepal Bagmati Sindhupalchok ”; 1 ♂ (NME) “ NEPAL, P: Gandaki; D: Manang way from Dharapani to Tal 28 ° 31 ′ 14 ” N, 84 ° 21 ′ 23 ″ E, 1920 m NN to 28 ° 28 ′ 06 ″ N, 84 ° 22 ′ 24 ″ E, 1685 m NN, 28. V. 2013, leg. A. Kopetz # 29 ”; 1 ♂ (GSPC) “ NEPAL, Mechi zone Taplejung 1700 m 22. V. 93 G. Sabatinelli leg. ”; 2 ♂♂ (NMBF) “ NEPAL Likhu Khola Tal 1700 m 4. VI. 62 leg. G. Ebert // Anomala flaviventris Arr. ♂ det. Machatschke 1969. ”; 1 ♀ (NMBF) “ NEPAL Likhu Khola Tal 1700 m 4. VI. 62 leg. G. Ebert // Anomala flaviventris det. G. Frey, 1963 // Anomala flaviventris Arr. ♀ det. Machatschke 1969. ”; 1 ♂ (NMBF) “ NEPAL Likhu Khola Tal 1700 m 4. VI. 62 leg. G. Ebert // Anomala flaviventris det. G. Frey, 1963 ”; 1 ♂ (NHMB) “ NEPAL: Kosi - # 11 Num 27 ° 33 ′ N / 87 ° 17 ′ E, 1550 m, 8 / 11. vi. 01 // NHMB Basel expedition to Nepal, 2001 ”; 1 ♂ (NMBF) “ NEPAL Sun Khosi Tal 2150 m 2. V. 62 leg. G. Ebert // Anomala flaviventris det. G. Frey, 1963 ”; 1 ♂ (NMBF) “ NEPAL Sun Khosi Tal 2150 m 2. V. 62 leg. G. Ebert // Anomala flaviventris Arr. ♀ det. Machatschke 1969. ”; 1 ♀ (ZMPC) “ China: Tibet, Shigatse, Nyalam co., Zhangmu, Yingbin Road, 2000 m, 2020. VII. 29, Yu-Chen Zheng leg. ”; 1 ♂ (NHMUK) “ Gopaldhara, Rungbong Vall., Sikkim. H. Stevens. 1916 – 218 // NHMUK 015015934 ”; 1 ♂ (NHMUK) “ Gopaldhara 1911 // Andrewes Bequest. B. M. 1922 - 221 // NHMUK 015015939 ”; 1 ♀ (NHMB) “ Changra 18 km S Tongsa 1900 m 22 / 6 // Nat. - Hist. Museum Basel - Bhutan Expedition 1972 // Anomala flaviventris Arr. det. G. Frey, 1974 ”; 1 ♀ (NHMB) “ 21 km O Wangdi Phodr. 1700 - 2000 // Nat. - Hist. Museum Basel - Bhutan Expedition 1972 // Anomala flavopicta Arr. det. G. Frey, 1973 ”; 1 ♂ (MFPC) “ BHUTAN: Dung Dung Nyelsa 27 ° 32 ′ N 90 ° 11 ′ E. 01 – 03. VII. 2009. H. 2970 m Viktor Siniaev-leg. ”	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	description	Description. Male. Body shape elongated ovoid, somewhat subparallel, moderately convex. Color. Generally yellowish brown, clypeus, disc of pronotum except for a broad lateral area, elytra, broad central area of metasternum, pygidium, abdominal ventrites (in posterior half), metatibia and all tarsomeres chocolate brown; frons and vertex blackish brown; dull, with weak greenish sheen; elytra with a yellow, narrow to broad and oblique band extending from primary costa 4 to 1; antennae yellowish or reddish brown; setae yellowish brown, robust setae reddish brown. Head. Greatest width / length of clypeus approximately 2.1; subtrapezoidal, lateral margins weakly convergent anteriad, anterior margin slightly convex, anterior corners obsolete and rounded; anterior margin weakly reflexed; surface with dense, large and coarse punctures. Frontoclypeal suture slightly incurved, distinct. An inverted triangular area on anterior half of frons punctate as clypeus, other portions with very dense and large punctures, smaller posteriad. Interocular distance / maximum head width approximately 0.57. Antenna long, length of antennal club slightly shorter than combined length of antennomeres 1 – 6. Inner margin of eye with a few moderately long setae. Pronotum. Sides strongly convergent anteriad, lateral margin straight in anterior third, slightly incurved in posterior third, moderately excurved at middle. Anterior angle acute and protruding, posterior angle obtuse. Posterior marginal line broadly interrupted before scutellum, other marginal lines complete. Lateral fovea indistinct. Disc with moderately dense and large punctures. Lateral margin with several moderately long setae. Scutellum triangular, lateral margin strongly excurved; with dense and large punctures. Elytra. Surface weakly wrinkled, intervals moderately elevated, subsutural interstice widest; strial punctures dense and large, subsutural interstice with irregularly doubled secondary stria, interstices II and III each with a secondary stria, all intervals with dense punctures, which are distinctly smaller than strial ones, each bearing a minute seta. Humeral and apical protuberance moderately prominent. Lateral carina slightly expanded from base to apical two fifths. Epipleuron with a row of moderately dense and short setae from base to apical two fifths, usually partly abraded. Abdomen. Propygidium with dense, transverse and fine striation. Pygidium broad and strongly tumid, with broadly rounded apex, disc with dense, transverse and incompletely annulated punctures, gradually confluent near the greatest tumidity; with dense and long setae preapically, and a transverse row of long setae along posterior margin. Abdominal ventrites 1 – 4 strongly carinate laterally, ventrite 4 with two separated protuberances medially, concave between ventrites 4, 5 and 6 in profile; ventrites 2 – 5 with transverse and incompletely annulated punctures, short and moderately dense medially, gradually elongated and denser laterad, frequently confluent at the lateralmost area, ventrite 6 densely rugostriolate; ventrites 2 – 5 each with a transverse and moderately dense row of long setae, dense laterally, the row of ventrite 6 present at posterior margin. Ventral thoracic surface. Hypomeron with sparse and small punctures, sparse and long setae. Ventral mesothoracic surface with dense and small punctures each bearing a short seta. Ventral metathoracic surface with dense, large and annulated punctures, gradually denser and confluent laterad, with dense and long setae, medial area with sparse and small punctures, setae short. Metacoxa punctate as lateral metasternum but bearing sparser setae. Very narrow at the extremity of mesosternum. Legs. Protibia bidentate, both teeth distinct and acute; apical tooth extending to middle of protarsomere 2. Inner spur inserted at the level of the preapical tooth. Inner protarsal claw and outer mesotarsal claw split apically, both lower branches equally as long as and two times wider than their upper branches; lower margin of lower branch of inner protarsal claw not convex next to the internobasal notch; outer metatarsal claw slightly longer than inner one. Protarsomeres thin, pro- and mesotarsomere 5 each with a small internomedial protuberance slightly behind middle, mesotarsomere 5 only with a large and acute internomedial denticle. Ventral surface of mesofemur with a row of moderately dense and long setae at lateral margin, a row of long and slightly robust setae submedially, additional sparse and long setae between rows; ventral surface of metafemur with two rows of moderately dense and long setae, one sublateral and the other submedial; meso- and metatibiae each bearing three transverse groups of robust setae emerging from a carina, with the basal fourth featuring some randomly arranged and shorter robust setae. Male genitalia. See fig. 24 E – H. Female. Abdominal ventrites and pygidium (except for a narrow anterior line) yellowish brown, metatibia and all tarsomeres darker than in male. Antenna shorter, length of antennal club approximately 0.8 times the combined length of antennomeres 1 – 6. Pygidium more triangular, less tumid preapically; ultimate abdominal ventrite 2.0 times as long as that in male medially, ventrites only feebly convex in profile, without protuberance. Apical protibial tooth longer and apex rounded, extending to anterior margin of protarsomere 2; inner spur inserted at mid-length of protibia; protarsomeres thinner, internomedial protuberance of protarsomere 5 absent, that of metatarsomere slightly smaller than in male; lower branch of inner protarsal claw and outer mesotarsal claw slightly wider than upper branch. Measurement. Body length: 13.4 – 14.4 mm in males and 13.5 – 14.5 mm in females, greatest width: 6.6 – 7.8 mm in males and 6.7 – 7.5 mm in females.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	diagnosis	Differential diagnosis. See that of Anomala flaviventris.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	discussion	Remarks. The examination of the specimen from Changra (NHMB, with Frey’s label) recorded as A. flaviventris by Frey (1975) confirmed that it actually belongs to A. brancuccii.	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
4A5987BAFF849338E9CA81A0FE5EFCC6.taxon	distribution	Distribution. China (Xizang) (new record); India (Uttarakhand, West Bengal, Sikkim) (new record); Nepal; Bhutan (new record).	en	Zhao, Ming-Zhi, Fujioka, Masayuki, Zorn, Carsten (2025): Taxonomic study of species related to Anomala flavonotata Arrow, 1912, A. flavofasciata Arrow, 1912, and A. flavovaria Arrow, 1917 from the Himalaya and adjacent regions (Coleoptera: Scarabaeidae: Rutelinae). Zootaxa 5693 (3): 301-356, DOI: 10.11646/zootaxa.5693.3.1, URL: https://doi.org/10.11646/zootaxa.5693.3.1
