taxonID	type	description	language	source
4E0487981A6DFFF4FF1DFF038BC2B75D.taxon	materials_examined	Type: — PHILIPPINES. Mindanao, District of Zamboanga (now Zamboanga City): Talisayan, San Ramon, Sax River (= Sas River), 14 February 1905, 240 m elevation, Williams 2167 (holotype: NY 00428854!, isotypes: NY 00428855!, US 00110084!, K 000761715!).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6DFFF4FF1DFF038BC2B75D.taxon	description	Description: — Small tree, up to 7 m tall, 12 cm in diameter at breast height, outer bark yellowish, lenticels sparse. Branchlets terete, pale green. Distal portions of the twigs with dense pale green (brownish when old) indumentum and scales; longer scales, appressed to slightly ascending, triangular to lanceolate, 3 – 6 mm long; shorter scales linear to lanceolate, up to 1.5 mm long. Leaves alternate, simple, chartaceous; petiole terete, pale green, 0.8 – 2.5 cm long, 3 – 4 mm in diameter, indumentum similar to that of distal twigs; young leaves brownish red; blade subpeltate, obovate to oblong-elliptic, 30 – 35 × 8 – 14 cm, adaxially olivaceous, glabrous except the midrib with brownish appressed acicular hairs, abaxially pale green, with appressed acicular hairs (0.8 – 1.0 mm long), denser in the midrib, secondary veins and reticulations; secondary veins 16 – 18 pairs, prominent on the lower surface, 15 – 23 mm apart along the midrib, reticulately veined, arching and fading towards the leaf margin, base truncate to subcordate, margin denticulate (hairs on each denticulation up to 1 mm long), apex acute to acuminate. Inflorescences cauliflorous, ramiflorous, borne on tubercles, fascicled cymose, many-branched, 3 – 6 cm long; flowers rotate, 7 – 15 per peduncle, 1.5 – 2.0 cm in diameter, odorless; peduncle pale green, 6 – 10 mm long, with dense, appressed, reddish, acicular hairs (0.5 – 0.8 mm long); bracts 2, green, triangular to ovate, 1.4 – 1.9 × 0.5 – 1.0 mm, with dense, appressed, reddish-maroon, acicular hairs (1 – 2 mm long); bracteoles 2, green, narrowly triangular, 3.5 – 4.0 × 0.7 – 1.0 mm, indumentum similar to that of bracts; pedicel reddish, 8 – 17 mm long, indumentum similar to that of peduncles. Sepals 5, white, pale green in fruits, outer sepals 2, elliptic-ovate, 4.2 – 4.8 × 3.0 – 3.5 mm (excluding setae), adaxial surface glabrous, abaxial surface with appressed, pinkish to red, acicular hairs (up to 1.2 mm long), margin entire, apex bluntly acute-obtuse to truncate; intermediate sepal present, partly covered with indumentum similar to that of outer sepals and partly glabrous similar to inner sepals; inner sepals 2, elliptic-ovate, 5.3 – 5.6 × 3.0 – 3.5 mm (excluding setae), adaxial surface glabrous, abaxial surface with indumentum similar to that of outer sepals but only at the median portion, margin entire, apex bluntly acute-obtuse. Petals 5, white, elliptic to ovate, 8.4 – 9.2 × 4.5 – 5.5 mm, glabrous on both surfaces, apex unequally retuse, sinus 0.5 – 0.7 mm long. Stamens 15 – 20, in two series, adnate to the base of corolla; filaments yellow, 1.0 – 1.2 mm long, glabrous; anthers dorsifixed, longitudinally dehiscent, oblong, 1.7 – 2.5 mm long, 0.1 – 0.2 mm wide, apex divided into 2 lobes, each lobe 0.15 – 0.20 mm long. Ovary white, ovoid, 2.0 – 2.2 × 3.0 – 3.5 mm, glabrous; locules 3, placentation axile, ovules numerous. Styles 3 (rarely 4), white, turning reddish in fruit, filiform, proximal 10 – 15 % fused, upper arms free, 4.5 – 5.0 mm long; stigma capitate, minutely corniculate. Fruits baccate, translucent green, globose to subglobose, 5.5 – 6.5 × 7.0 – 8.6 mm (styles excluded), glabrous, styles persistent. Seeds numerous, brown to black, irregularly rhomboid to oblong, 0.5 – 0.7 × 0.2 – 0.7 mm, embedded in mucilaginous matrix.	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6DFFF4FF1DFF038BC2B75D.taxon	distribution	Distribution and Ecology: — Endemic to the Philippines. Prior to its rediscovery, Saurauia denticulata was only known from its type specimen, which was collected more than a century ago in the lowland primary forest (240 m elevation) of Barangay Talisayan, Zamboanga City. Recently, new populations of S. denticulata were discovered in Barangay Tinuyop, Municipality of Leon B. Postigo. This species was observed to be thriving along rivers and streams in secondary tropical lowland forest at 350 – 700 m elevation. Notably, it has been recorded to be flowering and fruiting from January to April. Provisional IUCN conservation assessment: — The area of occupancy (AOO) of Saurauia denticulata is estimated to be 176.00 km 2 (Bachman et al. 2011). Based on our field observations, there are only about 10 – 20 mature individuals at the type locality (Barangay Talisayan, Zamboanga City), and about 5 – 10 mature individuals in Barangay Tinuyop, Leon B. Postigo, Zamboanga del Norte. At the type locality, the habitat is facing the threat of ecotourism and water facilities development. Moreover, agricultural expansion and small-scale mining operations pose significant risks to the newly discovered populations in Barangay Tinuyop, Leon B. Postigo. Due to the limited distribution and existing threats to its habitat, we categorized it as Endangered (EN B 2 b (iii )), in accordance with IUCN red list categories and criteria (IUCN, 2022). Notes: — In Zamboanga Peninsula, Saurauia denticulata resembles S. zamboangensis in having indumentum and scales on the distal twigs, cauliflorous inflorescence, and ovoid and 3 - locular ovary. However, S. denticulata can be easily recognized by its dense pale green (vs. reddish) indumentum and scales, longer scales being triangular to lanceolate (vs. narrowly lanceolate to triangular), shorter scales being linear to lanceolate (vs. acicular to linear) and up to 1.5 mm (vs. 2 mm) long, subpeltate (vs. non-peltate) and obovate to oblong-elliptic (vs. lanceolate) leaves, truncate to subcordate leaf bases (vs. acute), 16 – 18 pairs of secondary veins (vs. 12 – 14 pairs), 4.5 – 5.0 mm long styles that are 10 – 15 % proximally fused (vs. 3.5 – 4.0 mm long and 25 – 28 % proximally fused), and smaller fruits (5.5 – 6.5 × 7.0 – 8.6 mm vs. 8.5 – 10.0 × 7.5 – 9.5 mm).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6DFFF4FF1DFF038BC2B75D.taxon	description	Additional specimen examined: PHILIPPINES. Mindanao, Zamboanga del Norte Province, Municipality of Leon B. Postigo, Barangay Tinuyop, 8 ° 3 ’ 49.39 ” N, 122 ° 55 ’ 48.03 ” E, elevation 322 m, 6 February 2021, K. R. F. Mazo 15 (PNH 258578!; CMUH!)	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6BFFF6FF1DFED98BBBB75D.taxon	materials_examined	Type: — Philippines, Mindanao, Zamboanga, Mount Tubuan, 30 October 1919, elevation 500 m, BS 36607 Ramos & Edaño (lectotype (designated here): A 00098377!, isolectotype: US 00110167!).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6BFFF6FF1DFED98BBBB75D.taxon	description	Description: — Shrub to small tree, up to 5 m tall, 10 cm in diameter at breast height, outer bark greenish brown, lenticels sparse. Branchlets terete, green. Distal portions of the twigs with dense reddish (brownish when old) scales; longer scales appressed to ascending, narrowly lanceolate to triangular, 2.5 – 6.0 mm long; shorter scales acicular to linear, up to 2 mm. Leaves alternate, simple, chartaceous; petiole terete, pale green, 1.3 – 2.6 cm long, 4 – 5 mm in diameter, indumentum similar to that of distal twigs; young leaves green; blade lanceolate, 24 – 34 × 4.0 – 9.5 cm, adaxially olivaceous, glabrous except the midrib with sparse acicular scales; abaxially pale green, with appressed acicular hairs (0.5 – 2.0 mm long), denser in the midrib, secondary veins and reticulations; secondary veins 12 – 14 pairs, prominent on the lower surface, 22 – 35 mm apart along the midrib, reticulately veined, arching and fading towards the leaf margin, base acute, margin spinose (hairs up to 1.7 mm long), apex sub-caudate to acuminate. Inflorescences cauliflorous, borne on tubercle, fascicle cymose, many-branched, 3 – 7 cm long; flowers rotate, 5 – 12 per peduncle, 1.5 – 2.0 cm in diameter, odorless; peduncle brownish-red, 5 – 11 mm long, with dense appressed, reddish, acicular hairs (1.5 mm long); bracts 2, red, triangular, 1.4 – 1.5 × 0.5 – 0.7 mm, with appressed reddish, acicular hairs (0.7 – 1.3 mm long); bracteoles 2, red, triangular, 1.5 – 1.6 × 0.5 – 0.7 mm, indumentum similar to the bracts; pedicels brownish-red, 9 – 22 mm long, indumentum similar to that of peduncles. Sepals 5, white, pinkish-red in fruits; outer sepals 2, white, oblong-ovate, 6.0 – 7.5 × 2.5 – 4.3 mm (excluding setae), adaxial surface glabrous, abaxial surface with red, appressed, linear to lanceolate scales (up to 1.3 mm long), margin entire, apex acute to obtuse; intermediate sepal present, partly covered with hairs similar to outer sepals and partly glabrous similar to inner sepals; inner sepals 2, white, oblong to ovate, 6.3 – 8 × 2.6 – 4.5 mm (excluding setae), adaxial surface glabrous, abaxial surface with indumentum similar to that of outer sepals but scale only present at the median base of the inner sepals, margin entire, apex bluntly acute-obtuse. Petals 5, white, oblong-elliptic, 7.5 – 9.0 × 4.5 – 6.0 mm, glabrous on both surfaces, apex unequally retuse, sinus 0.4 – 0.8 mm long. Stamens 13 – 18, yellow, in two series, adnate to the base of the corolla; filaments yellow, 1.0 – 1.3 mm long, glabrous; anthers 3.0 – 3.2 mm long × 0.3 – 0.4 mm wide, oblong, dorsifixed, longitudinally dehiscent, apex divided into 2 lobes, lobes 0.1 – 0.2 mm long. Ovary white, ovoid, 2.0 – 2.2 × 3.0 – 3.5 mm, glabrous; locules 3, placentation axile, ovules numerous. Styles 3, white (reddish in fruit), proximal 25 – 28 % fused, filiform, 3.5 – 4.0 mm long, upper arms free; stigma capitate, minutely corniculate. Fruits baccate, translucent white, globose to subglobose, 8.5 – 10.0 × 7.5 – 9.5 mm (styles excluded), glabrous, styles persistent. Seeds numerous, brown to black, irregular triangular to oblong, 0.3 – 0.5 × 0.1 – 0.4 mm, embedded in mucilaginous matrix.	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6BFFF6FF1DFED98BBBB75D.taxon	distribution	Distribution and Ecology: — Endemic to the Philippines. In the Barangay Talisayan, Zamboanga City, this species is found in a primary forest at 500 – 1000 m elevation. In Barangay Tinuyop, Leon B. Postigo, populations were seen thriving in a secondary evergreen lowland tropical forest at 200 – 350 m elevation. Saurauia zamboangensis has been observed flowering and fruiting from February to August. Local name and Uses: — Locals in Zamboanga del Norte call Saurauia zamboangensis as himag-pula whereas it is known as sasaua for the Subanen, an indigenous group in the Zamboanga peninsula (Merrill, 1925). Its fruits are consumed as a forest snack. Provisional IUCN conservation assessment: — The area of occupancy (AOO) of Saurauia zamboangensis is estimated to be less than 500 km 2 (296.00 km 2; Bachman et al. 2011). Additionally, only 50 – 100 and 30 – 50 mature individuals were observed in Barangay Talisayan, Zamboanga City, and Barangay Tinuyop, Leon B. Postigo, Zamboanga del Norte, respectively. The habitats of both populations face the threats of infrastructure development for ecotourism and water supply, agricultural expansion, and unregulated small-scale mining. We therefore categorized S. zamboangensis as Endangered (EN B 2 b (iii )) following IUCN red list categories and criteria (IUCN, 2022).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A6BFFF6FF1DFED98BBBB75D.taxon	description	Notes: — Saurauia zamboangensis is unique among all Philippine Saurauia species by its lanceolate, slender, subcaudate-acuminate leaves that are adaxially glabrous and conspicuously appressed setose on the midrib. In Barangay Tinuyop, S. zamboangensis occasionally grows sympatrically with S. denticulata. Additional specimen examined: — PHILIPPINES. Mindanao, Zamboanga del Norte Province, Municipality of Leon B. Postigo, Barangay Tinuyop, 8 ° 3 ’ 52.46 ” N, 122 ° 55 ’ 44.66 ” E, elevation 316 m, 17 February 2021, K. R. F. Mazo 17 (PNH 258580!; CMUH!)	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A69FFF8FF1DFEDB8B9FB779.taxon	materials_examined	Type: — Philippines, Visayas, Leyte, Palo, January 1906, elevation 200 m, Elmer 7530 (lectotype (designated here): NY 00428870!, isolectotype: E 00683159!).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A69FFF8FF1DFEDB8B9FB779.taxon	description	Description: — Shrub to small tree, 2.5 m high, 10 cm in diameter at breast height, bark greenish-brown, lenticels sparse. Branchlets terete, green. Distal portions of the twigs with dense, erect and spreading bristle-like hairs; hairs 5 – 7 mm long, greenish on middle portion, drying brown at the tip. Leaves alternate, simple chartaceous; petiole green, terete, 10 – 16 mm long, 3.0 – 4.6 mm in diameter, with indumentum similar to that of distal twigs; young leaves light green; blade green, oblanceolate, 13.5 – 27.0 × 3 – 5 cm, adaxially colliculate, uniformly with bristle-like thin and erect hairs (up to 1.5 mm long except the larger hairs on the midrib); abaxially pale green, pubescent, hairs denser on the midrib, secondary veins and reticulations; secondary veins 12 – 14 pairs, prominent on the lower surface, 5 – 14 mm apart along the midrib, reticulately veined, arching and fading towards the leaf margin, base rounded, margin serrulate and ciliate (hairs up to 2.5 mm long), apex caudate-acuminate. Inflorescences axillary, terminal, solitary or few-branched cyme, 4 – 6 cm long; flowers rotate, 1 – 4 per peduncle, 2.0 – 2.3 cm in diameter, odorless; peduncle pale green, 1.3 – 3.0 cm long, with indumentum similar to that of distal twigs (hairs 3.0 – 4.5 mm long); bracts 2, green, linear, 10 – 15 × 1.5 – 2.0 mm, with indumentum similar to that of distal twigs (hairs 1 – 2 mm long); bracteoles 2, green, linear, 6 – 9 × 1.0 – 1.5 mm, with indumentum similar to that of bracts (hairs 2.5 – 4.0 mm long); pedicels light green, 10 – 15 mm long, with indumentum similar to that of peduncle (hairs 5 – 6 mm long). Sepals 5, white, pale green in fruits; outer sepals 2, translucent white, ovate, 7.2 – 8.4 × 4.2 – 4.5 mm (excluding setae), adaxial surface glabrous, abaxial surface with pale green, appressed and slightly spreading bristle-like hairs (up to 4 mm long), margin entire, apex bluntly acute-obtuse; intermediate sepal present, partly covered with hairs similar to outer sepals and partly glabrous similar to inner sepals; inner sepals 2, translucent white, oblong-elliptic to ovate, 5.2 – 6.1 × 3.3 – 3.8 mm (excluding setae), adaxial surface glabrous, abaxial surface with indumentum similar to that of outer sepals but only at the median portion, margin entire, apex bluntly acute-obtuse. Petals 5, white, oblong, 10 – 12 × 4.5 – 5.6 mm, glabrous on both surfaces, apex strongly unequally retuse, sinus 0.5 – 0.7 mm long. Stamens 20 – 25, yellow, in three series, adnate to the corolla; filaments yellow, 1.2 – 2.0 mm long, glabrous; anthers oblong, 2.0 – 2.5 mm long × 0.5 – 0.7 mm wide, dorsifixed, longitudinally dehiscent, apex divided into 2 lobes, each lobe 0.5 – 1.3 mm long. Ovary white, ovoid, 1.5 – 2.0 × 2.5 – 2.7 mm, glabrous; locules 3, placentation axile, ovules numerous. Styles 3, white, (pale green in fruit), proximal 30 % fused, filiform, 2.8 – 3.5 mm long, stigma capitate, minutely corniculate. Fruits baccate, translucent white, globose to subglobose, 7 – 8 × 8.5 – 10.5 mm (styles excluded), glabrous, styles persistent. Seeds numerous, brown, irregularly triangular to oblong, 0.4 – 0.8 × 0.3 – 0.6 mm, embedded in mucilaginous matrix.	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A69FFF8FF1DFEDB8B9FB779.taxon	distribution	Distribution and Ecology: — Endemic to the Philippines. Saurauia merrillii is distributed in the islands of Leyte, Samar, and in the province of Zamboanga del Norte. In Zamboanga Peninsula, S. merrillii is found growing in the secondary lowland evergreen rainforest near a river or stream at 400 – 750 m elevation. It has been observed flowering in December and fruiting in May. Local Name and Uses: — Similar to other Saurauia species in Zamboanga del Norte, S. merrillii is also known as himag. The mature twigs and stems are dried and used as an ointment for wounds. Provisional IUCN conservation assessment: — Saurauia merrillii is proposed here as Vulnerable (V D 2) following IUCN red list categories and criteria (IUCN, 2022). The extent of occurrence (EOO) is less than 20,000 km 2 (7, 734 km 2; Bachman et al. 2011). During fieldwork in Barangay Tinuyop, Leon B. Postigo, Zamboanga del Norte, only three individuals of S. merrillii were observed. The habitat of S. merrillii in Barangay Tinuyop is not protected, and threats such as small-scale mining, forest road construction, and land conversion were recorded.	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A69FFF8FF1DFEDB8B9FB779.taxon	description	Notes: — Saurauia merrillii resembles S. erythrotricha Elmer (1915: 2614) and S. clementis Merrill (1906: 208) in having densely bristly distal twigs, leaves, and sepals, axillary and terminal inflorescences, 3 - locular, and glabrous ovoid ovary. However, it differs from S. erythrotricha in having greenish-brown bristle-like hairs (vs. reddish-brown to purplish), longer petioles (10 – 16 mm vs. 5 – 10 mm), oblanceolate leaves (vs. subovate to obovate, elliptic), 12 – 14 pairs of secondary veins (vs. 5 – 8 pairs), smaller sepals (5.5 – 7.5 × 3.0 – 4.2 mm vs. 8 × 6 mm), larger petals (10 – 12 × 4.5 – 5.6 mm vs. 8 × 2.5 – 3.5 mm), and shorter styles (2.8 – 3.5 mm vs. 4 mm). On the other hand, S. merrillii can be easily differentiated from S. clementis Merr. in having oblanceolate leaves (vs. elliptic to oblong-obovate), 12 – 14 pairs of secondary veins (7 – 8 pairs), and oblong petals (vs. obovate).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
4E0487981A69FFF8FF1DFEDB8B9FB779.taxon	materials_examined	Additional specimens examined: — Saurauia merrillii Elmer. PHILIPPINES. Mindanao, Zamboanga del Norte Province, Municipality of Leon B. Postigo, Barangay Tinuyop, 8 ° 2 ’ 50.69 ” N, 122 ° 56 ’ 32.72 ” E, elevation 337 m, 2 December 2021, K. R. F. Mazo 62 (PNH 258600!; CMUH!); Visayas, Samar, Loquilocon Wright, April – May 1948, Sulit 2099 (PNH 6114!).	en	Mazo, Kean Roe F., Aribal, Lowell G. (2025): Rediscoveries, lectotypifications and new species record of the genus Saurauia (Actinidiaceae) from southwestern Philippines. Phytotaxa 704 (1): 69-78, DOI: 10.11646/phytotaxa.704.1.6, URL: https://doi.org/10.11646/phytotaxa.704.1.6
