taxonID	type	description	language	source
830106F4986C5C3EACC30877509FDA7D.taxon	etymology	Etymology. The genus name is derived from the Ancient Greek γλυφή (gluphe), meaning " a carving, " and Greek γλῶσσα (glossa), meaning " tongue. "	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
830106F4986C5C3EACC30877509FDA7D.taxon	distribution	Distribution. From south-western China across Indochina to Myanmar, Thai-Malay Peninsula, islands of Sumatra and Borneo (Fig. 1).	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
045DB91529295F338B0E0D7FF04A2884.taxon	etymology	Etymology. The genus name is derived from the Greek μικρός (mikros), meaning " small, " and " Hylas " (for origin of this name see above).	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
045DB91529295F338B0E0D7FF04A2884.taxon	distribution	Distribution. Frogs of the genus Microhyla are widely distributed across the East (southern China, including Taiwan and Hainan islands, and Ryukyu Archipelago of Japan), Southeast (Myanmar and Indochina, Malayan Peninsula, Sumatra, Java, Bali, and Borneo), and South Asia (Bangladesh, Nepal, Indian subcontinent to north-eastern Pakistan in the west and Sri Lanka in the south) (Fig. 1).	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
C444A1B69DF558E6B6CCBE2DA94E5A81.taxon	etymology	Etymology. The genus name is derived from the Greek νᾶνος (nanos), meaning " dwarf ", " pygmy ", and the mythological figure, Hylas (Ancient Greek: Ὕλας), which is probably derived from the Ancient Greek verb " ὕλαω " meaning " to bark " (Bourret 1942). In classical mythology, Hylas, son of King Theiodamas, was a youth who served as Heracles' companion, lover, and servant. Heracles took Hylas with him on the Argonauts' expedition, during which Hylas was kidnapped by nymphs of the spring in Pegae, Mysia, and turned into an echo. Heracles left the ship and was searching for Hylas for a great length of time, calling his name: " His adjunxit Hylan nautae quo fonte relictum / Clamassent ut littus Hyla! Hyla! omne sonaret " (" The mariners cried on Hylas till the shore / Then Re-echoed Hylas! Hylas! soothed ... "; Virgil 1916, Ecl. 6, 43). The genus name refers to the small body size (<25 mm) of all known Nanohyla species, while maintaining resemblance to its sister genus Microhyla, from which it is separated herein. The new genus name is feminine in gender.	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
C444A1B69DF558E6B6CCBE2DA94E5A81.taxon	diagnosis	Diagnosis. The new genus is assigned to the subfamily Microhylinae on the basis of phylogenetic affinities and the following combination of morphological character states: vomers small, confined to the anterior and medial margins of choanae; clavicles and, in most cases, procoracoids absent, maxillary arcade edentate (Parker 1934). Nanohyla gen. nov. differs from other Microhylinae genera by the following combination of osteological character states: (1) frontoparietals fused with exoccipitals; (2) exoccipitals fused with each other (incomplete fusion in N. pulchella); (3) neopalatines present; (4) sphenethmoids completely fused with parasphenoid (incomplete fusion in N. pulchella); (5) crista parotica entirely cartilaginous; (6) otic ramus of squamosal well-developed; (7) tympanic annulus well-developed; (8) transverse processes of presacral vertebrae with the following orientation: IV and V posterolaterally, II, VII and VIII anterolaterally, III and VI at right angle to body axis; (9) clavicles absent; (10) omosternum present, cartilaginous; (11) prehallux cartilaginous; (12) terminal phalanges of longest fingers and toes T-shaped. The combination of diagnostic external morphological characters includes: (13) small to extremely small frogs (adult SVL 11.8 - 25.8 mm); (14) snout rounded or pointed in profile; (15) supratympanic fold present; (16) ridge on posterior sides of choanae absent; (17) first finger (FI) length less than 1 / 2 FII or reduced to a nub; (18) finger discs present, at least on FII-FIV; (19) dorsal median longitudinal grooves on finger discs generally present (with the exception of N. perparva); (20) toes dorsolaterally flattened, prominent discs present; (21) dorsal median longitudinal grooves on toe discs present; (22) metatarsal tubercle single (inner metatarsal tubercle present, outer absent); (23) dorsomedial line absent; (24) superciliary tubercles absent; (25) tibiotarsal articulation of adpressed hindlimb reaching well beyond snout; (26) toe webbing well-developed (at least one-half webbed); (27) skin on dorsum feebly granular to tubercular; (28) tympanum externally distinct at least in males (N. annamensis, N. annectens, N. arboricola, N. marmorata, N. nanapollexa, N. pulchella) or barely distinct (N. hongiaoensis, N. perparva, N. petrigena); (29) terrestrial or scansorial semi-arboreal microhabitat preference.	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
C444A1B69DF558E6B6CCBE2DA94E5A81.taxon	distribution	Distribution. The distribution area of Nanohyla gen. nov. covers montane forests of the Annamite (Truong Son) Mountains in Vietnam, eastern Laos, and north-eastern Cambodia, the Titiwangsa Mountain Range in the southernmost Thailand and peninsular Malaysia, mountains of Borneo (including Sabah and Sarawak of Malaysia, Brunei, and Kalimantan of Indonesia) and the Sulu Archipelago of the Philippines (see Fig. 1). The occurrence of Nanohyla gen. nov. in Cardamom Mountains in eastern Thailand (the record of " M. annamensis " from Khao Sebab by Taylor [1962], see Fig. 1) is questionable (see Poyarkov et al. 2014, 2020 a).	en	Gorin, Vladislav A., Scherz, Mark D., Korost, Dmitriy V., Poyarkov, Nikolay A. (2021): Consequences of parallel miniaturisation in Microhylinae (Anura, Microhylidae), with the description of a new genus of diminutive South East Asian frogs. Zoosystematics and Evolution 97 (1): 21-54, DOI: http://dx.doi.org/10.3897/zse.97.57968, URL: http://dx.doi.org/10.3897/zse.97.57968
