taxonID	type	description	language	source
58586E20FFAD2675FF09FB73FA63FF63.taxon	type_taxon	Type species: Formicococcus cinnamomi Takahashi 1928.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	diagnosis	Diagnosis (adapted from Williams 2004 and Joshi et al. 2020). Body of adult female with anal lobe bar present. Cerarii basically numbering 18 pairs, although sometimes cerarii on head or thorax indistinct but preocular pair (C 2) always present. Anal lobe cerarii each bearing 2 or more conical cerarian setae, each seta with a pointed or flagellate tip. Anterior abdominal cerarii each usually containing more than 2 conical setae; if only 2, then auxiliary setae often present. Cerarii on head and thorax almost always present, each either with 2 or more conical or flagellate setae and a group of trilocular pores, or, if with only 2 enlarged setae, then these sometimes associated with slender auxiliary setae. Sometimes 1 or 2 cerarii on thorax obscure, occasionally reduced to a single conical or flagellate seta but always with at least 1 or 2 trilocular pores next to setal collar. Ostioles prominent. Anal ring normal, situated at apex of abdomen or slightly dorsal in position, bearing 6 or multiple ring setae. Antennae each with 6 – 8 segments, occasionally with 9. Legs well developed, hind leg with translucent pores present or absent; often tibia + tarsus shorter than trochanter + femur; claw stout, without a denticle. Circulus present or absent. Multilocular pores present, at least on venter of abdomen. Oral rim tubular ducts absent. Oral collar tubular ducts present, at least on venter of abdomen, sometimes also around margins of head and thorax.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. The adult female of Formicococcus is morphologically similar to that of Paraputo Laing but differs from the latter by presence of an anal lobe bar (Williams 2004). However, Danzig & Gavrilov-Zimin (2015) considered that the presence or absence of an anal lobe bar is a character that falls under individual variations and instead, used the presence of more than six anal ring setae as a critical generic character state of the genus Formicococcus. However, as discussed by Williams (2004) and Zhang & Wu (2017), the number of anal ring setae is not a robust genus-level character, especially as the number of setae can vary between either side of the anal ring. Herein, we have adopted the view of Williams (2004) and Zhang & Wu (2017), using the presence of an anal lobe bar as a genus-level character distinguishing Formicococcus from Paraputo. The monotypic genus Anaparaputo Borchsenius differs from other genera by having large oral collar tubular ducts on all abdominal segments and lateral to each anterior coxa. However, this character is also present in some Formicococcus species, such as F. mangiferacola Williams 2004 and F. polysperes Williams 2004, so we here consider Anaparaputo (which had been placed as a junior synonym of Paraputo by Danzig & Gavrilov-Zimin (2015 )) as a junior synonym of Formicococcus.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Guangdong: 2 ♀ ♀, Nanling, Ruyang, on Fargesia sp., 22. viii. 2010, coll. San-an Wu & Yuan Lu.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Poaceae: Bambusa sp., Fargesia sp.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Guangdong, Taiwan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. A good description and illustration of the adult female was given by Li et al. (2014).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Guizhou: 6 ♀ ♀, Suiyang, Rangshui, on leaves of Phyllostachys pubescens, 14. viii. 2010, coll. San-an Wu. Zhejiang: 9 ♀ ♀, Longquan city, on leaves of Ph. pubescens, 3. viii. 2008, coll. Jin Liu; 2 ♀ ♀, Deqing county, on the stem of Polygonatum odoratum, 11. viii. 2003, coll. Yong-sheng Chai; 13 ♀ ♀, Tianmu Mountain, on the stem of Ph. pubescens, 14. iv. 1999, coll. San-an Wu.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Asparagaceae: Polygonatum odoratum; Poaceae: Phyllostachys pubescens.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Guizhou, Zhejiang).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. This species was transferred to Paraputo, as P. bambusus (Wu), by Danzig & Gavrilov-Zimin (2015) because it has an anal ring with six setae. However, the number of anal ring setae is not a robust genus-level character, especially as the number of setae can vary between the two sides of the anal ring. As it has an anal lobe bar on each anal lobe, this species is here restored to Formicococcus. Good descriptions and illustrations of the adult females were given by Wu (2001) and Li et al. (2014).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Sichuan: 4 ♀ ♀, Ya’an city, Sichuan Agricultural University Campus, under bark crack of Platanus sp., 27. vii. 2014, coll. Jiang-tao Zhang & Xu-bo Wang. Guangxi: 5 ♀ ♀, Xing’an county, undetermined host, 27. vi. 1999, coll. San-an Wu.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Ebenaceae: Diospyros sp.; Lauraceae: Cinnamomum camphora; Platanaceae: Platanus sp.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Guangxi, Hubei, Sichuan, Taiwan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Wu (2001) also reported this species from Hubei, Xianfeng county. Good descriptions and illustrations of the adult females were given by Ezzat & McConnell (1956) and Tu et al. (1988).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Rutaceae: Citrus sp.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Fujian).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. A good description and illustration of the adult female was given by Tang (1992).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	description	(Fig. 1)	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. Holotype Adult ♀ on a slide, China: Yunnan, Xishuangbanna / on Delonix regia (Fabaceae), ix. 2015 / specimens provided by Yi-wei Fang; BFUC. (The label is written in Chinese.) Paratypes: 4 ♀ ♀ (on four slides), same date and locality as holotype; 2 in BFUC, 2 in CFJAU.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	description	Description Appearance in life: Not recorded. Slide-mounted adult female (n = 5). Body of adult female (Fig. 1) broadly oval, 1.80 – 2.55 mm long and 1.15 – 2.00 mm wide. Anal lobes slightly prominent, ventral surface of each lobe with an apical seta, each 215 – 232 μm long, ratio of lengths of apical setae to anal ring setae 1: 0.37 – 0.51, and with a well-developed anal lobe bar. Antennae each 333 – 360 μm long, 7 - or 8 - segmented; apical segment longest, bearing 4 fleshy setae. Legs well developed, stout; hind leg measurements in μm: coxa 88 – 110 long; trochanter + femur 270 – 300 long, tibia + tarsus 209 – 237 long; claw stout, 35.0 – 42.5 long, without denticle; claw digitules knobbed, longer than claw. Ratio of lengths of hind tibia + tarsus to hind trochanter + femur 1: 1.26 – 1.31. Ratio of lengths of hind tibia to tarsus 1: 0.79 – 0.88. Translucent pores numerous on hind coxa. Eye oval, located at body margin posterior to antennal base. Labium 235 – 242 μm long, longer than clypeolabral shield. Circulus present, about 115 μm long and 113 μm wide, situated between abdominal segments III and IV, divided by intersegmental line. Ostioles well developed, each lip with inner edge sclerotized and with 4 – 12 short setae and many trilocular pores. Anal ring situated on dorsum about half its length from apex of abdomen, 87 – 97 μm long and 75 – 89 μm wide, bearing 6 long setae, each 100 – 132 μm long, longer than diameter of anal ring. Cerarii basically numbering 18 pairs, cerarian setae sometimes with flagellate tips. Anal lobe cerarii (C 18) each containing 6 – 8 conical setae, each 21.5 – 26.0 μm long and 7 – 8 μm wide at base, and a group of trilocular pores. Posterior abdominal cerarii (C 15 – C 17) each containing 3 – 6 conical setae; more anterior cerarii each usually containing 2 conical setae, but on thorax sometimes reduced to a single conical seta or sometimes cerarius not discernible. Dorsal surface with short, stiff setae, each 10 – 15 μm long on head and 17.5 – 27.0 μm long on abdomen. Trilocular pores, each 3.5 – 4.0 μm wide, evenly distributed. Multilocular disc pores and oral collar tubular ducts absent. Discoidal pores, each smaller than a trilocular pore, few, scattered. Ventral surface with slender flagellate setae, each 51.5 – 75.0 μm long on head and 57.5 – 80.0 μm long on abdomen. Cisanal and obanal setae present, each 100 – 132 μm long. Multilocular disc pores, each about 8 μm in diameter, present posterior vulva in single or double transverse rows across median area of abdominal segments IV – VII; sometimes also 1 or 2 on marginal areas of abdominal segment VII. Trilocular pores evenly distributed, fewer than on dorsum. Oral collar tubular ducts of 2 sizes present: (i) a large type, each 9 – 10 μm long and 4 μm wide, present across posterior medial areas of abdominal segments V – VI, sometimes a few also on medial areas of abdominal segments IV or VII, and forming groups on margins of posterior abdominal segments V – VIII; some also present on head; and (ii) a small type, each 7 – 8 μm long and 2.5 – 3.0 μm wide, present across posterior medial areas of abdominal segments V – VII, also 1 or 2 present on abdominal segment VIII. Discoidal pores few, scattered.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Fabaceae: Delonix regia.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Yunnan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	etymology	Etymology. The species name is based on a combination of the host plant genus name, Delonix, with the Latin suffix “ - icola ” meaning dweller, and is used as a noun in apposition.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Formicococcus delonicola is most similar to F. latens Williams 2004 in possessing oral collar tubular ducts on the head and in marginal groups on the posterior abdominal segments. However, it differs from the latter by the following features (condition of F. latens given in parentheses): (i) dorsal setae on abdominal segment VIII short, about same length as other dorsal setae (noticeably longer than other dorsal setae); (ii) anal ring setae longer than diameter of anal ring (shorter than diameter of anal ring or same length); and (iii) cerarian setae sometimes with flagellate tips (conical, without flagellate tips).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Rosaceae: Eriobotrya japonica.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Fujian).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. This species was transferred to Paraputo, as P. eriobotryae (Wang), by Danzig & Gavrilov-Zimin (2015) because it has an anal ring with six setae. However, the number of anal ring setae is not a robust genus-level character, especially as the number of setae can vary between the two sides of the anal ring. As it has an anal lobe bar on each anal lobe, this species is here restored to Formicococcus. A good description and illustration of the adult female was given by Wang (1985).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Arecaceae: Areca catechu; Cyperaceae: Cyperus rotundus; Poaceae: Oryza sativa, Saccharum officinarum, Sorghum bicolor. Distributions. China (Guangdong), India, Indonesia, Malaysia, Thailand.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Descriptions and illustrations of the adult females were given by Ferris (1954), Williams (1970) and Williams (2004).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Guizhou: 4 ♀ ♀, Xingyi city, under bark crack of Ficus virens, 8. v. 2017, coll. Jiangtao Zhang & Ming Zhao. Yunnan: 1 ♀, Jinghong city, on Ficus sp., 25. iv. 1957, coll. N. S. Borchsenius.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Moraceae: Ficus sp., F. virens.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Guizhou, Yunnan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. As it has an anal lobe bar on each anal lobe, Anaparaputo liui Borchsenius is here transferred to Formicococcus as Formicococcus liui (Borchsenius) comb. n. A good description and illustration of the adult female was given by Borchsenius (1962).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Ebenaceae: Diospyros philippinensis; Euphorbiaceae: Macaranga sinensis, M. tanarius, Mallotus sp., Ma. japonicus; Fabaceae: Pueraria montana (= P. lobata); Moraceae: Ficus microcarpa.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Taiwan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Tsai (2011) also reported this mealybug on Diospyros philippinensis in Taiwan. A good description and illustration of the adult female was given by Tu et al. (1988).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Rosaceae: Prunus mume.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Zhejiang).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. This species was transferred from Formicococcus back to Planococcus as P. mumensis Tang by Danzig & Gavrilov-Zimin (2015) because it has cerarii without multiple setae and an anal ring with six ring setae. However, as auxiliary setae are present in all the cerarii and the anal lobe cerarii each contain three conical setae, here we follow Cox (1989) in replacing it under Formicococcus. Good descriptions and illustrations of the adult females were given by Tang (1977, 1992).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Guangxi: 3 ♀ ♀, Jingxi city, on the root of Panax notoginseng, 30. xi. 2015, coll. Jun-jing Zhou. Yunnan: 2 ♀ ♀, Ruili city, Nanmao Lake Park, on the root of Ophiopogon sp., 22. x. 2016, coll. Xu-bo Wang & Yao-guang Qin; 2 ♀ ♀, Jinghong city, Binhe Park, under bark crack of Ficus sp., 12. x. 2016, coll. Xu-bo Wang & Yao-guang Qin; 2 ♀ ♀, Linchang, on the root of Amomum villosum, 21. v. 2016, coll. Zi-lin Yang; 2 ♀ ♀, Dehong, Coffea sp., 17. viii. 2013, coll. Fu-zhong Wu; 1 ♀, Baoshan city, Eriobotrya japonica, 3. viii. 2012; 1 ♀, Jinghong city, Artocarpus heterophyllus, 20. x. 2008, coll. Kai-ping Ji; 1 ♀, Jinghong city, on the root of Musa acuminata, 3. iv. 2008, coll. Jin Liu, San-an Wu & Shan-shan Wang; 15 ♀ ♀, Jinghong city, Xishuangbanna Tropical Flowers and Plants Garden, on the root of Strelitzia reginae, 2. iv. 2008, coll. Jin Liu, San-an Wu & Shan-shan Wang; 1 ♀, Jinghong city, Xishuangbanna Tropical Flowers and Plants Garden, on the root of Dypsis lutescens, 2. iv. 2008, coll. Jin Liu, San-an Wu & Shan-shan Wang; 3 ♀ ♀, Jinghong city, Xishuangbanna Tropical Flowers and Plants Garden, Puccinellia distans, 1. iv. 2008, coll. Jin Liu, San-an Wu & Shan-shan Wang.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Araceae: Amorphophallus paeoniifolius; Araliaceae: Panax notoginseng; Arecaceae: Areca catechu, Cocos nucifera, Dypsis lutescens, Rhapis excelsa; Asparagaceae: Ophiopogon sp.; Asteraceae: Ageratum conyzoides, Synedrella nodiflora; Cyperaceae: Rhynchospora colorata (= Cyperus kyllingia); Euphorbiaceae: Balakata baccata (= Sapium baccatum), Macaranga conifera, M. triloba; Fabaceae: Arachis hypogaea, Delonix regia, Erythrina; Lamiaceae: Clerodendrum infortunatum; Meliaceae: Lansium parasiticum (= L. domesticum); Moraceae: Artocarpus heterophyllus, Ficus sp.; Musaceae: Musa acuminata; Phyllanthaceae: Phyllanthus niruri; Piperaceae: Piper betle, P. nigrum; Poaceae: Puccinellia distans; Rubiaceae: Coffea sp., C. arabica; Rosaceae: Eriobotrya japonica; Solanaceae: Physalis minima; Strelitziaceae: Strelitzia reginae; Urticaceae: Urtica parviflora; Zingiberaceae: Amomum villosum, Curcuma longa, Zingiber sp., Z. officinale. Distributions. China (Guangxi, Yunnan), India, Malaysia, Philippines, Thailand, Vietnam.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Zhang et al. (2015) reported that this species formed fungus-insect galls with Phlebopus portentosus (Berk. and Broome) Boedijn 1951 (Fungi: Boletales: Boletinellaceae) on the roots of Coffea arabica and Delonix regia in Yunnan. Good descriptions and illustrations of the adult and second instar females were provided by Williams (2004).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Acanthaceae: Crossandra sp.; Anacardiaceae: Lannea coromandelica, Mangifera indica; Annonaceae: Annona squamosa; Apocynaceae: Tabernaemontana sp.; Arecaceae: Phoenix dactylifera; Asteraceae: Conyza bonariensis (= C. ambigua); Cucurbitaceae: Cucurbita maxima, Cu. pepo; Fabaceae: Acacia nilotica (= A. arabica), Albizia lebbeck, Erythrina sp., E. variegata (= E. indica), Prosopis cineraria (= P. spicigera), Vachellia nilotica; Gnetaceae: Gnetum luofuense; Moraceae: Artocarpus heterophylla, Ficus benghalensis, F. religiosa, Morus sp.; Myrtaceae: Psidium guajava, Syzygium cumini; Papaveraceae: Argemone mexicana; Rhamnaceae: Ziziphus sp.; Rubiaceae: Coffea arabica; Rutaceae: Citrus sp.; Vitaceae: Vitis vinifera. Distributions. China (Yunnan), Bangladesh, India, Iran, Pakistan.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Wu (2009) reported this mealybug from Yunnan on Coffea spp. Good descriptions and illustrations of the adult females were provided by Ezzat & McConnell (1956) and Williams (2004).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Theaceae: Schima noronhae.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Taiwan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. A good description and illustration of the adult female was given by Tu et al. (1988).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Juglandaceae: Engelhardia sp.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Yunnan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. A good description and illustration of the adult female was given by Borchsenius (1962).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plant. Theaceae: Camellia oleifera (= C. oleosa).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Hunan).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. Good descriptions and illustrations of the adult females were given by Wang (1982 a, 1982 b).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	materials_examined	Material examined. CHINA: Beijing: 2 ♀ ♀, Beijing Forestry University Campus, under bark crack of Platanus occidentalis, 20. ix. 2016, coll. Jiang-tao Zhang; 1 ♀, Beijing Forestry University Campus, under bark crack of Platanus occidentalis, 19. ix. 2016, coll. Jiang-tao Zhang; 1 ♀, Beijing Forestry University Campus, under bark crack of Platanus occidentalis, 16. ii. 2016, coll. Jiang-tao Zhang & Xu-Bo Wang; 6 ♀ ♀, Beijing Botanical Garden, Koelreuteria paniculata, 20. ix. 2008, coll. De-Ping Xiong. Jiangxi: 5 ♀ ♀, Ganzhou city, Yangmeidu Park, under bark crack of Acer palmatum, 23. viii. 2016, coll. Jiang-tao Zhang & Qin-gang Dong; 3 ♀ ♀, Yingtan city, Yingtan Park, under bark crack of Platanus sp., 14. viii. 2016, coll. Jiang-tao Zhang & Qin-gang Dong; 4 ♀ ♀, Yingtan city, under bark crack of Platanus sp., 14. viii. 2016, coll. Jiang-tao Zhang & Qin-gang Dong. Hebei: 1 ♀, Xingtai city, under bark crack of Platanus sp., 8. x. 2016, coll. Jiang-tao Zhang. Shaanxi: 18 ♀ ♀, Northwest A & F University Campus, Prunus cerasifera var. atropurpurea, 2. ix. 1999, coll. San-an Wu.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	biology_ecology	Host plants. Platanaceae: Platanus sp., P. occidentalis; Rosaceae: Prunus cerasifera var. atropurpurea; Sapindaceae: Acer palmatum, Koelreuteria paniculata.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	distribution	Distribution. China (Beijing, Hebei, Jiangxi, Shaanxi).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Remarks. A good description and illustration of the adult female was given by Wu & Zheng (2001). Notes. The species Pedronia acanthodes Wang 1979 was described as new by Wang (1979), subsequently transferred to Indococcus by Tang (1992), and then transferred (doubtfully) to Formicococcus by Danzig & GavrilovZimin (2015). It is not included in this study since, according to Wang’s descriptions (1982 a, 2001), it possesses quinquelocular pores on the venter and lacks bars on the anal lobes; we therefore judge that it does not fall within our understanding of the morphology of Formicococcus. Another species, Drymococcus rhizophilus Borchsenius 1962, was transferred to Formicococcus, as F. rhizophilus (Borchsenius 1962), by Danzig & Gavrilov-Zimin (2015) because it has an anal ring with multiple ring setae. However, the number of anal ring setae is not a robust genus-level character, especially as the number of setae can differ between the two sides of the same anal ring. As the species lacks anal lobe bars, here it is not included in Formicococcus.	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
58586E20FFAD2675FF09FB73FA63FF63.taxon	discussion	Key to adult females of Formicococcus species in the Palaearctic and Oriental Regions (adapted and modified from Wu & Zheng 2001, Williams 2004, Joshi et al. 2020 and Tanaka 2021).	en	Zhang, Jiang-Tao, Wu, San-An (2022): A review of the genus Formicococcus Takahashi 1928 (Hemiptera: Coccomorpha Pseudococcidae) of China, with description of a new species. Zootaxa 5188 (2): 157-168, DOI: 10.11646/zootaxa.5188.2.5
