taxonID	type	description	language	source
5F0C87E2163BFFBEFE2AEA40FB6CFE5C.taxon	description	Figs 1 A – B, 2 A – B, 3 – 7	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2163BFFBEFE2AEA40FB6CFE5C.taxon	description	This species has been studied extensively (see synonymy) and we do not have new material. However, it is included here because two very similar species newly described below require an updated diagnosis, and because a few details were missing in previous studies. In addition, some specimens previously assigned tentatively to this species are now considered to belong to other species (see Distribution below).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2163BFFBEFE2AEA40FB6CFE5C.taxon	diagnosis	Diagnosis Leaf-dwelling, long-legged pholcid with dark pattern on carapace (Figs 1 – 2). Easily distinguished from most known congeners (except M. wayuu sp. nov. and M. uca sp. nov.) by strongly asymmetric male palps (Figs 3 – 4; including asymmetry of femur and tibia), by male chelicerae with pair of strong lateral protrusions (Fig. 5 A), and by female external and internal genitalia (Figs 5 C, 6 – 7; epigynum with posterior semicircular process, or ‘ scape’; internal genitalia with complex system of pouches, ducts, and folds). Distinguished from both M. wayuu and M. uca by male chelicerae with rounded rather than pointed distal apophyses (Fig. 5 A), by main branch of left procursus with distinctive ventral indentation (bold arrow in Fig. 4 C), by hair-like process on right procursus (arrow in Fig. 4 D), and by less pronounced palpal asymmetry, i. e., absolutely and relatively smaller right palp (e. g., right / left tibia diameter <2.5 vs> 2.6; see also Fig. 22). Further distinguished from M. uca by different color pattern on carapace (both in males and females; Fig. 2), by smaller size and shorter legs (e. g., male tibia 1 <5.0 vs> 6.5; female tibia 1 <4.0 vs> 5.5), and by female external genitalia (epigynal scape without asymmetric groove; compare Fig. 6 B with Fig. 16 B). Description (amendments; see Huber 2000, 2004) Right procursus with hair-like process on retrolateral side (arrow in Fig. 4 D). Prolateral trichobothrium absent on tibia 1, present on other leg tibiae. Female tibia 1 length (N = 50): 3.2 – 3.8 (mean 3.5). Shape of pore plates slightly variable (Fig. 7 B, D). Barcodes We sequenced five specimens from two localities (geographic distance: 270 km) (Table 1; Fig. 23). Within localities, distances were 0.0 %; between localities, distances ranged from 8.4 – 9.1 % (Table 2). Distances to the other three species treated herein ranged from 15.5 to 20.2 %.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2163BFFBEFE2AEA40FB6CFE5C.taxon	distribution	Distribution Known from several localities in eastern Venezuela (Sucre, Bolívar) and northwestern Brazil (Roraima) (Fig. 24). The females from Falcón listed in Huber & Villarreal (2020) are here assigned to the newly described M. wayuu sp. nov. (see below). Specimens from Peru listed and illustrated in Huber (2000) are considered to represent a distinct, formally undescribed species (or more than one species).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	description	Figs 1 C – D, 2 C – D, 8 – 12	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	diagnosis	Diagnosis Easily distinguished from most known congeners (except M. mariguitarensis and M. uca sp. nov.) by strongly asymmetric male palps (Fig. 8), by male chelicerae with pair of strong lateral protrusions (Fig. 10 A), and by female external and internal genitalia (Figs 11 – 12; epigynum with posterior semicircular process, or ‘ scape’; internal genitalia with complex system of pouches, ducts, and folds). Distinguished from M. mariguitarensis by male chelicerae with pointed rather than rounded distal apophyses (Fig. 10 A), by main branch of left procursus without distinctive ventral indentation (compare Fig. 4 C with Fig. 9 C), by absence of hair-like process on right procursus (compare Fig. 4 D with Fig. 9 D), and by stronger palpal asymmetry, i. e., absolutely and relatively bigger right palp (e. g., right / left tibia diameter> 2.6 vs <2.5; see also Fig. 22). Further distinguished from M. uca by different color pattern on carapace (both in males and females; Fig. 2), by smaller size and shorter legs (e. g., male tibia 1 <5.0 vs> 6.5; female tibia 1 <4.0 vs> 5.5), and by female genitalia (epigynal scape without asymmetric groove; compare Fig. 11 B with Fig. 16 B).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	etymology	Etymology The species name honors the Wayuu, an indigenous ethnic group of the Guajira Peninsula in northernmost Colombia and northwestern Venezuela.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	materials_examined	Type material Holotype COLOMBIA – La Guajira • ♂; Tomarrazón; 11.0701 ° N, 72.9357 ° W; 310 m a. s. l.; 19 Sep. 2022; B. A. Huber leg.; degraded forest along river; MUSENUV Ar 3531. Paratypes COLOMBIA – La Guajira • 1 ♂, 2 ♀♀; same collection data as for holotype; MUSENUV Ar 3532 • 2 ♂♂, 2 ♀♀; same collection data as for holotype; ZFMK Ar 24780. Other material examined COLOMBIA – La Guajira • 4 ♀♀, in pure ethanol; same collection data as for holotype; ZFMK Col 290 (voucher of UH 288; one abdomen cleared and transferred to ZFMK Ar 24780). VENEZUELA – Falcón • 1 ♀; forest near Santa Cruz de La Alegría; 10.8795 ° N, 68.4949 ° W; 100 m a. s. l.; 15 Feb. 2020; B. A. Huber, O. Villarreal M. and Q. Arias C. leg.; ZFMK Ar 22013 • 4 ♀♀, in pure ethanol; same collection data as for preceding; ZFMK Ven 20 - 149 (voucher of M 092).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.8, carapace width 0.75. Distance PME – PME 170 µm; diameter PME 90 × 110 µm; distance PME – ALE 25 µm; AME absent. Leg 1: 18.7 (5.0 + 0.4 + 4.7 + 7.6 + 1.0), tibia 2: 2.8, tibia 3: 1.8, tibia 4: 2.8; tibia 1 L / d: 59; diameters of leg femora (at half length) 0.10; of leg tibiae 0.08. COLOR (in ethanol). Carapace pale whitish ochre, with median brown mark including ocular area (Fig. 2 C), clypeus not darkened; sternum whitish; legs whitish ochre, patellae and tibia-metatarsus joints brown; abdomen whitish, dorsally with few dark marks. BODY. Habitus as in Fig. 1 C. Ocular area slightly raised. Carapace without thoracic groove. Clypeus with sclerotized rim (sclerotized rim narrower than in M. uca sp. nov.; cf. Fig. 15 A). Sternum wider than long (0.54 / 0.46), unmodified. Abdomen approximately twice as long as wide, dorso-posteriorly pointed. CHELICERAE. As in Fig. 10 A – B; with pair of lateral conical processes, pair of distal frontal processes set with ~ 23 modified (globular) hairs each, and pair of slightly diverging apophyses in front of fang joints. PALPS. As in Fig. 8; coxa unmodified, apparently symmetric; trochanter with short rounded ventral process, apparently symmetric; all other segments directionally asymmetric, see below; femur with distinct process on prolateral-ventral side; procursus consisting of main branch and ventral hinged process; genital bulb simple, consisting of globular part and embolus. ASYMMETRY. Right femur rather cylindrical, with cylindrical prolateral-ventral process; left femur smaller, distally strongly widened, and with conical prolateral-ventral process. Right tibia much bigger than left tibia (maximum diameter in lateral view 0.56 – 0.61 vs 0.21 – 0.22). Right procursus consisting of strongly sclerotized main branch (distally the sclerite is partly internal, covered by weakly sclerotized semitransparent cuticle) and weakly sclerotized semitransparent ventral hinged process; left procursus much smaller, with slender main branch, with distally widened and heavily sclerotized ventral hinged process. Right genital bulb with small globular part (diameter 0.27 – 0.29) and long slender embolus; left genital bulb with larger globular part (diameter 0.36 – 0.38) and with shorter and strongly widened embolus (similar to M. uca sp. nov.; cf. Fig. 15 B – C). LEGS. Without spines, without curved hairs, without sexually dimorphic short vertical hairs; retrolateral trichobothrium of tibia 1 at 9 %; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~ 20 pseudosegments, very indistinct except distally. Variation Males Tibia 1 in three other males: 4.2, 4.2, 4.5. Color pattern on carapace consistent but pattern on abdomen variable, entirely whitish or with black and white marks dorsally. Females In general very similar to male (Fig. 1 D) but carapace pattern limited to black lines (Fig. 2 D), clypeus unmodified, sternum color variable, in some females with brown marks near leg coxae or entire sternum speckled; abdomen color variable as in males. Tibia 1 in ten females: 3.6 – 3.9 (mean 3.7). Epigynum (Fig. 11) simple, mostly weakly sclerotized, posteriorly with sclerotized scape, apparently symmetric; internal asymmetric structures visible through cuticle; posterior epigynal plate short and indistinct. Internal genitalia (Figs 10 C, 12) with sclerotized receptacle on right side, with complex system of pouches, ducts, and folds (apparently similar to M. mariguitarensis; cf. Huber 2004: figs 1 – 2); with pair of oval pore plates (possibly slightly asymmetric, but this might be an artifact of preparation). Barcodes We sequenced two specimens from two localities (geographic distance: 485 km) (Table 1; Fig. 23). The CO 1 distance was 1.7 % (Table 2). Distances to the other three species treated herein ranged from 15.5 to 20.3 %.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21638FFA8FDA5E9BDFBEFFD96.taxon	distribution	Distribution Known from northern Colombia (La Guajira) and northwestern Venezuela (Falcón) (Fig. 24). Natural history At the type locality, the spiders were collected from palm and Heliconia L. leaves in a disturbed forest remnant along a river. The locality was shared with Mesabolivar eberhardi Huber, 2000 and an undescribed species of Chibchea Huber, 2000. In Falcón, the five females were collected close to the forest margin of a well-preserved forest. They shared the locality with two further representatives of Metagonia, M. latigo Huber, 2020 and M. guttata Huber, 2020. The latter species seemed to share the same large dicot plant leaves with M. wayuu sp. nov. (Huber & Villarreal 2020).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	description	Figs 1 E – F, 2 E – F, 13 – 16	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	diagnosis	Diagnosis Easily distinguished from most known congeners (except M. mariguitarensis and M. wayuu sp. nov.) by strongly asymmetric male palps (Fig. 13), by male chelicerae with pair of strong lateral protrusions (Fig. 15 D), and by female external and internal genitalia (Figs 15 F, 16; epigynum with posterior semicircular process, or ‘ scape’; internal genitalia with complex system of pouches, ducts, and folds). Distinguished from both M. mariguitarensis and M. wayuu by different color pattern on carapace (both in males and females; Fig. 2), by larger size and longer legs (e. g., male tibia 1> 6.5 vs <5.0; female tibia 1> 5.5 vs <4.0), and by female genitalia (epigynal scape with asymmetric groove; Fig. 16 B). Further distinguished from M. mariguitarensis by male chelicerae with pointed rather than rounded distal apophyses (Fig. 15 D), by main branch of left procursus without distinctive ventral indentation (compare Fig. 4 C with Fig. 14 C), by absence of hair-like process on right procursus (compare Fig. 4 D with Fig. 14 D), and by stronger palpal asymmetry, i. e., absolutely and relatively bigger right palp (e. g., right / left tibia diameter> 2.6 vs <2.5; see also Fig. 22).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	etymology	Etymology The species name alludes to fiddler crabs (Uca and other genera), where the males have a major claw significantly larger than the minor claw.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	materials_examined	Type material Holotype COLOMBIA – Quindío • ♂; Armenia, Universidad del Quindío; 4.5537 ° N, 75.6608 ° W; 1500 m a. s. l.; 8 Sep. 2022; B. A. Huber and G. A. Rodríguez leg.; MUSENUV Ar 3533. Paratypes COLOMBIA – Quindío • 1 ♂, 2 ♀♀; same collection data as for holotype; MUSENUV Ar 3534 • 7 ♂♂, 9 ♀♀; same collection data as for holotype; ZFMK Ar 24781, 24782. Other material examined COLOMBIA – Quindío • 3 ♀♀, 1 juv., in pure ethanol; same collection data as for holotype; ZFMK Col 238 (voucher of UH 542) • 1 ♀; 3 km N of Circasia, Reserva Bosque del Silencio; 4.6411 ° N, 75.6379 ° W; 1850 m a. s. l.; 7 Sep. 2022; B. A. Huber and G. A. Rodríguez leg.; ZFMK Ar 24783 • 2 ♀♀, in pure ethanol; same collection data as for preceding; ZFMK Col 237 (voucher of UH 254). – Risaralda • 1 ♀; La Celia, Vereda San Eugenio; 4.970 ° N, 75.006 ° W; 1600 m a. s. l.; 25 Feb. 2010; N. Betancour leg.; MUSENUV 2214 • 1 ♀; same collection data as for preceding; MUSENUV 2220 • 1 ♂, 1 ♀; same collection data as for preceding; MUSENUV 2226.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 3.2, carapace width 0.9. Distance PME – PME 250 µm; diameter PME 120 × 130 µm; distance PME – ALE 30 µm; AME absent. Leg 1: 31.5 (7.8 + 0.5 + 7.7 + 14.0 + 1.5), tibia 2: 4.9, tibia 3: 2.8, tibia 4: 4.4; tibia 1 L / d: 86; diameters of leg femora (at half length) 0.10 – 0.11; of leg tibiae 0.09. COLOR (in ethanol). Carapace anteriorly pale ochre-yellow, posterior half dark brown (Fig. 2 E), ocular area with brown pattern, clypeus brown; sternum whitish; legs ochre-yellow, patellae and tibia-metatarsus joints dark brown; abdomen pale ochre-gray, dorsally and laterally with dark and whitish marks. BODY. Habitus as in Fig. 1 E. Ocular area slightly raised. Carapace without thoracic groove. Clypeus with strongly sclerotized rim (Fig. 15 A). Sternum wider than long (0.68 / 0.56), unmodified. Abdomen approximately twice as long as wide, dorso-posteriorly pointed. CHELICERAE. As in Fig. 15 D – E; with pair of lateral conical processes, pair of distal frontal processes set with ~ 12 – 14 modified (globular) hairs each, and pair of diverging apophyses near fang joints. PALPS. As in Fig. 13; coxa unmodified, apparently symmetric; all other segments directionally asymmetric, see below; trochanter with short ventral apophysis; femur with distinct process on prolateral-ventral side; procursus consisting of main branch and ventral hinged process; genital bulb simple, consisting of globular part and embolus. ASYMMETRY. Right trochanter slightly larger than left trochanter. Right femur rather cylindrical, with cylindrical prolateral-ventral process; left femur smaller, distally strongly widened, and with conical prolateral-ventral process. Right tibia much bigger than left tibia (maximum diameter in lateral view: 0.70 – 0.77 vs 0.25 – 0.27). Right procursus consisting of strongly sclerotized main branch (distally the sclerite is mostly internal, covered by weakly sclerotized and semitransparent cuticle) and weakly sclerotized semitransparent ventral hinged process; left procursus much smaller, with slender main branch, with distally widened and heavily sclerotized ventral hinged process. Right genital bulb with small globular part (diameter 0.30 – 0.34) and long slender embolus; left genital bulb with larger globular part (diameter 0.42 – 0.46) and with shorter and strongly widened embolus (Fig. 15 B – C). LEGS. Without spines, without curved hairs, without sexually dimorphic short vertical hairs; retrolateral trichobothrium of tibia 1 at 8 %; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~ 25 pseudosegments, very indistinct. Variation Males Tibia 1 in ten males: 6.7 – 7.8 (mean 7.4). Females In general very similar to male (Fig. 1 F) but carapace posteriorly with two Y-marks (Fig. 2 F), ocular area and clypeus not darkened, clypeus unmodified, sternum darker (either dark brown with small light spots or light brown and medially ochre). Tibia 1 in 18 females: 5.7 – 6.6 (mean 6.2). Epigynum (Fig. 16 A – C) simple, mostly weakly sclerotized, posteriorly with sclerotized and slightly asymmetric scape (groove on left side; all females same-sided); internal asymmetric structures visible through cuticle; posterior epigynal plate short and indistinct. Internal genitalia (Figs 15 F, 16 D – E) with sclerotized receptacle on right side, with complex system of pouches, ducts, and folds (apparently similar to M. mariguitarensis; cf. Huber 2004: figs 1 – 2); with pair of elongated pore plates (possibly slightly asymmetric, but this might be an artifact of preparation). One cleared female from Vereda San Eugenio apparently identical to female from type locality. Barcodes We sequenced two specimens from two localities (geographic distance: 10 km) (Table 1; Fig. 23). The CO 1 distance was 0.2 % (Table 2). Distances to the other three species treated herein ranged from 18.8 to 20.9 %.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E2162EFFAFFD97EAC7FD5AF8FB.taxon	distribution	Distribution Known from three neighboring localities in Colombia (Fig. 24). Natural history In Armenia, most specimens were collected from palm leaves in the entrance area of the small secondary forest within the university campus; deeper in the forest they seemed to be very rare or absent. They shared the locality with a second (undescribed, symmetric) species of leaf-dwelling Metagonia. In Reserva Bosque del Silencio, the three females were collected on a single small palm tree; no further specimens were found despite focused search.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	description	Figs 1 G – H, 17 – 21 M 025 Metagonia Car 260 – Huber et al. 2022: 678 (molecular data).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	diagnosis	Diagnosis Easily distinguished from known congeners by male palpal asymmetry largely restricted to procursus (Figs 17 – 18), i. e., barely involving palp size as in other species with asymmetric males described above. Also by rounded median process on clypeus (Fig. 19 A), by male chelicerae with small, modified (globular) hairs arranged in two lateral bands (Fig. 19 A), by epigynum without scape (Fig. 20 B), and by asymmetric structures in female internal genitalia (Figs 19 B, 20 D – G; sclerotized receptacle on right side and arched membranous structure on left side).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	etymology	Etymology The species name honors the Emberá, an indigenous people in the Chocó Department of western Colombia and in Panama.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	materials_examined	Type material Holotype COLOMBIA – Risaralda • ♂; near Santa Cecilia; 5.3458 ° N, 76.1094 ° W; 450 m a. s. l.; 9 Sep. 2022; B. A. Huber and G. A. Rodríguez leg.; on leaves at forest edge; MUSENUV Ar 3535. Paratype COLOMBIA – Risaralda • 1 ♂; same collection data as for holotype; ZFMK Ar 24784. Other material examined COLOMBIA – Chocó • 1 ♂, 1 ♀, in pure ethanol; Jardin Botánico del Pacífico, “ trail to Mirador after stream crossings ”; 6.266 ° N, 77.375 ° W; 16 Jan. 2014; CarBio team leg.; ZFMK Car 260 (voucher of M 025) • 1 juv., assigned tentatively, in pure ethanol; Jardin Botánico del Pacífico, Ceiba Loop Trail; 6.266 ° N, 77.375 ° W; 13 Jan. 2014; CarBio team leg.; ZFMK Car 257. – Risaralda • 1 ♂, in pure ethanol; same collection data as for holotype; ZFMK Col 247 (voucher of UH 260). PANAMA – Colón • 1 ♂; San Lorenzo Protected Area; 9.267 ° N, 79.967 – 79.983 ° W; ~ 150 m a. s. l.; 13 May 2004; J. Schmidl and J. Bail leg.; fogging; ZMUC • 1 ♂; same locality as for preceding; 15 May 2004; J. Schmidl and J. Bail leg.; ZMUC • 1 ♀; same locality as for preceding; 17 Oct. 2003; J. Schmidl and A. Floren leg.; ZMUC • 1 ♂; same locality as for preceding; 17 Oct. 2003; J. Schmidl and J. Bail leg.; ZMUC • 1 ♂; same locality as for preceding; 12 May 2003; J. Schmidl and J. Bail leg.; ZMUC • 1 ♀; same locality as for preceding; 27 May 2004; J. Schmidl and J. Bail leg.; ZMUC • 1 ♂; same locality as for preceding; 8 Oct. 2004; J. Bail leg.; ZMUC • 2 ♂♂; same locality as for preceding; 15 May 2004; J. Schmidl and J. Bail leg.; ZMUC.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	description	Description Male (holotype) MEASUREMENTS. Total body length 2.1, carapace width 0.7. Distance PME – PME 70 µm; diameter PME 90 × 110 µm; distance PME – ALE 30 µm; AME absent. Leg 1: 22.9 (5.7 + 0.3 + 5.4 + 10.3 + 1.2), tibia 2: 3.5, tibia 3: 2.1, tibia 4: 3.3; tibia 1 L / d: 77; diameters of leg femora (at half length) 0.09; of leg tibiae 0.07. COLOR (in ethanol). Carapace pale ochre-yellow with distinct ochre-brown pattern on posterior half, ocular area and clypeus without dark marks (Fig. 1 G – H); sternum mostly whitish, only posteriorly with small ochre mark; legs ochre-yellow, patellae and tibia-metatarsus joints dark brown; abdomen pale ochre-gray, dorsally and laterally with dark marks. BODY. Habitus as in Fig. 1 G – H. Ocular area slightly raised. Carapace without thoracic groove. Clypeus with short median process at rim (length ~ 30 µm). Sternum wider than long (0.56 / 0.44), unmodified. Abdomen approximately twice as long as wide, dorso-posteriorly pointed. CHELICERAE. As in Fig. 19 A; with series of ~ 16 – 19 modified (globular) hairs on each side and pair of distal apophyses near fang joints, without proximal lateral processes. PALPS. As in Fig. 17; coxa unmodified; trochanter with short ventral apophysis; femur short, distally strongly widened, with short conical process on prolateral side, apparently symmetric; femur-patella hinges and tibia-tarsus hinges slightly shifted toward prolateral side; tibia short, slightly asymmetric (see below), with two trichobothria in relatively distal position; procursi strongly asymmetric, see below; genital bulb simple, with small conical process on prolateral side and long weakly sclerotized embolus ending with pointed process, slightly asymmetric (see below). ASYMMETRY. Proximal palpal segments (coxa, trochanter, femur) apparently symmetric. Right tibia slightly thicker than left tibia (maximum diameter in lateral view ~ 220 vs 210 µm). Procursi directionally asymmetric (Fig. 18): left procursus with strong and distally sclerotized ventral hinged process, with slender prolateral sclerotized process, prominent dorsal hump, without distal sclerite; right procursus with slender and weakly sclerotized ventral hinged process, without prolateral sclerotized process, with smaller dorsal hump, with long distal sclerite directed towards ventral and prolateral. Genital bulbs weakly asymmetric: right embolus slightly longer than left embolus (~ 340 vs 310 µm). LEGS. Without spines, without curved hairs, without sexually dimorphic short vertical hairs; retrolateral trichobothrium of tibia 1 at 7 %; prolateral trichobothrium absent on tibia 1; tarsus 1 with ~ 20 pseudosegments, very indistinct. Variation Males Tibia 1 in eight males: 4.8 – 5.9 (mean 5.5). Sternum in some males with small dark marks near bases of leg coxae. Females In general, very similar to male (including color pattern on carapace) but clypeus unmodified. Tibia 1 in two females: 4.5, 5.3 (missing in third female). Epigynum (Fig. 20 A – C) very simple, weakly sclerotized, without scape; internal asymmetric structures visible through cuticle; posterior epigynal plate short and indistinct. Internal genitalia (Figs 19 B, 20 D – G) asymmetric, with sclerotized receptacle on right side connected with arched membranous structure on left side; the three examined females were same-sided; with pair of elongated pore plates (possibly slightly asymmetric, but this might be an artifact of preparation). One cleared female from Panama very similar but pore plate slightly longer (Fig. 21). Barcodes We sequenced two specimens from two localities (geographic distance: 173 km) (Table 1; Fig. 23). The CO 1 distance was 0.6 % (Table 2). Distances to the other three species treated herein ranged from 16.7 to 20.9 %.	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
5F0C87E21629FFA5FDBEEF1CFBCAFCD8.taxon	distribution	Distribution Known from two localities in Colombia and one locality in Panama (Fig. 24).	en	Huber, Bernhard A., Meng, Guanliang (2025): New species of Metagonia Simon with directionally asymmetric male and female genitalia (Araneae, Pholcidae). European Journal of Taxonomy 1026: 199-235, DOI: 10.5852/ejt.2025.1026.3117, URL: https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/3117/13893
