taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
70C57444733F5CC1B42AE716455B51E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
70C57444733F5CC1B42AE716455B51E1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
36C2048CD31E566B8399BCEE29EF6580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
36C2048CD31E566B8399BCEE29EF6580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
36C2048CD31E566B8399BCEE29EF6580.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
5DE22A3C14D3591A82BED8AA0EF56DF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
5DE22A3C14D3591A82BED8AA0EF56DF9.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9FB30488ACD2566D8EEA52D9D9235FD3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9FB30488ACD2566D8EEA52D9D9235FD3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F221AB6043A652C798A27235C7018788.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F221AB6043A652C798A27235C7018788.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F221AB6043A652C798A27235C7018788.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
2C57081BD4BB510CB7F73E8EF00E8186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
2C57081BD4BB510CB7F73E8EF00E8186.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
17653170638E5DE4836C36C4ED07CCE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
17653170638E5DE4836C36C4ED07CCE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
17653170638E5DE4836C36C4ED07CCE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
39EF56621CCE5BF492AC15B1ED539DE5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F8F38414BA1E5CE09AC770954CBDEA85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F8F38414BA1E5CE09AC770954CBDEA85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F8F38414BA1E5CE09AC770954CBDEA85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
F8F38414BA1E5CE09AC770954CBDEA85.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
B980BDC8A9B25FFA811ABAA0487F9070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
B980BDC8A9B25FFA811ABAA0487F9070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
45B02AE51170537C9292453D8D500106.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
45B02AE51170537C9292453D8D500106.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
45B02AE51170537C9292453D8D500106.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
45B02AE51170537C9292453D8D500106.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
45B02AE51170537C9292453D8D500106.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
C74F133673595E7087AA0B784EFC7317.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
C74F133673595E7087AA0B784EFC7317.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
C74F133673595E7087AA0B784EFC7317.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
C74F133673595E7087AA0B784EFC7317.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
428FCF1C77E756299B727F57400D4E78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
428FCF1C77E756299B727F57400D4E78.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
970319F3E10F50F9813F8F087AB27BF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
970319F3E10F50F9813F8F087AB27BF6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6F6ECC4648225AFE9201C8CFB0D24189.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6F6ECC4648225AFE9201C8CFB0D24189.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3344D494A1405E52808E20046747AE15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3344D494A1405E52808E20046747AE15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3344D494A1405E52808E20046747AE15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3344D494A1405E52808E20046747AE15.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
E240DE1214085EE69BC1A3043200C861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
E240DE1214085EE69BC1A3043200C861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
E240DE1214085EE69BC1A3043200C861.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269565	https://doi.org/10.3897/zookeys.1229.119155.figure1	Figure 1. a analysis using IQTree (UFbootstrap values as percentage of 1000) and MrBayes 3.2. 7 (/ posterior probabilities) of DNA barcodes of Topiris, including all unique haplotypes sequenced (except one 133 bp sequence of Topiris albidella), showing their relationship to the genus Athrypsiastis and two exemplars of Metathrinca, rooted by Xylorycta (type species: X. luteotactella). The reconstructed DNA barcode Topiris candidella (NHMUK 010219690, highlighted in blue) from Sarawak falls in a clade with specimens of other members of the BIN cluster BOLD: ADR 9781 from which it differs by 1-3 bp. Topiris is recovered as monophyletic. HT = holotype, PT = paratype, NT = neotype; bp = no of nucleotides sequenced, if not 658 b morphological dataset (65 characters) as analysed for 24 taxa in MrBayes 3.2. 7 under a gamma model, rooted by the type species of Xylorycta (X. luteotactella). The specimen numbers and sexes of the exemplars used for describing the morphological characters are set out in this figure. Where possible, the holotype (HT) [or neotype (NT) was used for scoring. PT = paratype, NT = neotype, m = male, f = female. The male of Topiris candidella (highlighted in blue) is represented by a modern specimen (NHMUK 019223170), since the holotype is incomplete. T. candidella is morphologically cryptic with T. cinderella except for small differences in the male genitalia.	Figure 1. a analysis using IQTree (UFbootstrap values as percentage of 1000) and MrBayes 3.2. 7 (/ posterior probabilities) of DNA barcodes of Topiris, including all unique haplotypes sequenced (except one 133 bp sequence of Topiris albidella), showing their relationship to the genus Athrypsiastis and two exemplars of Metathrinca, rooted by Xylorycta (type species: X. luteotactella). The reconstructed DNA barcode Topiris candidella (NHMUK 010219690, highlighted in blue) from Sarawak falls in a clade with specimens of other members of the BIN cluster BOLD: ADR 9781 from which it differs by 1-3 bp. Topiris is recovered as monophyletic. HT = holotype, PT = paratype, NT = neotype; bp = no of nucleotides sequenced, if not 658 b morphological dataset (65 characters) as analysed for 24 taxa in MrBayes 3.2. 7 under a gamma model, rooted by the type species of Xylorycta (X. luteotactella). The specimen numbers and sexes of the exemplars used for describing the morphological characters are set out in this figure. Where possible, the holotype (HT) [or neotype (NT) was used for scoring. PT = paratype, NT = neotype, m = male, f = female. The male of Topiris candidella (highlighted in blue) is represented by a modern specimen (NHMUK 019223170), since the holotype is incomplete. T. candidella is morphologically cryptic with T. cinderella except for small differences in the male genitalia.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
3F7F5D9B58505E4CBB1AA38CEDF12C30.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269567	https://doi.org/10.3897/zookeys.1229.119155.figure3	Figure 3. ‘ Framework’ tree showing position of Topiris, Metathrinca and Athrypsiastis falling within ‘ core Xyloryctidae ’ (i. e. with the clade including the type species of Xylorycta, X. luteotactella), rather than with the genera Eumenodora plus Izatha (part of Xyloryctidae sensu lato, rooted by Martyringa (Lecithoceridae )). The combined matrix with the COI data was analysed in the context of multi-gene data (for six terminals that include labels ‘ MM’ and ‘ NKU’; all of COI plus between four-seven nuclear genes) using Phyml 3.0 (nodes are shown with ABayes support values in red). The total number of genes is shown (‘ g’) for taxa with multiple loci, where not only COI - 5 P. COI sequences below 200 bp or near-identical are excluded. Support values shown are ABayes values > 0.94 from analysis in Phyml 3.0 (online) under a GTR + G model.	Figure 3. ‘ Framework’ tree showing position of Topiris, Metathrinca and Athrypsiastis falling within ‘ core Xyloryctidae ’ (i. e. with the clade including the type species of Xylorycta, X. luteotactella), rather than with the genera Eumenodora plus Izatha (part of Xyloryctidae sensu lato, rooted by Martyringa (Lecithoceridae )). The combined matrix with the COI data was analysed in the context of multi-gene data (for six terminals that include labels ‘ MM’ and ‘ NKU’; all of COI plus between four-seven nuclear genes) using Phyml 3.0 (nodes are shown with ABayes support values in red). The total number of genes is shown (‘ g’) for taxa with multiple loci, where not only COI - 5 P. COI sequences below 200 bp or near-identical are excluded. Support values shown are ABayes values > 0.94 from analysis in Phyml 3.0 (online) under a GTR + G model.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9D59D67AEC045D739692DE6CE2DE8CDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9D59D67AEC045D739692DE6CE2DE8CDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269572	https://doi.org/10.3897/zookeys.1229.119155.figures48-56	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	Figures 48–56. Male genitalia of Topiris and Athrypsiastis spp. 48 T. sampitella (Lvovsky, 2014), holotype, Gen. prep. No 63 49 A, B Athrypsiastis phaeoleuca Meyrick, 1910, holotype, slide no. NHMUK 010316398 50 A, B A. cheesmanae sp. nov., holotype, slide no. NHMUK 010316428 51 A, B A. edelweissella sp. nov., holotype, slide no. NHMUK 010316429 52 A. penumbrella sp. nov. A holotype, slide no. NHMUK 010316430 B paratype, slide no. NHMUK 010316843 53 A, B A. symmetra Meyrick, slide no. NHMUK 010316443 54 A. halmaherella (Lvovsky), holotype, Gen. prep. No 62 55 A. chionodes Diakonoff, 1954, holotype 56 A. delicata Diakonoff, 1954, holotype. Scale bar: 1 mm (49–53 B).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9D59D67AEC045D739692DE6CE2DE8CDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269565	https://doi.org/10.3897/zookeys.1229.119155.figure1	Figure 1. a analysis using IQTree (UFbootstrap values as percentage of 1000) and MrBayes 3.2. 7 (/ posterior probabilities) of DNA barcodes of Topiris, including all unique haplotypes sequenced (except one 133 bp sequence of Topiris albidella), showing their relationship to the genus Athrypsiastis and two exemplars of Metathrinca, rooted by Xylorycta (type species: X. luteotactella). The reconstructed DNA barcode Topiris candidella (NHMUK 010219690, highlighted in blue) from Sarawak falls in a clade with specimens of other members of the BIN cluster BOLD: ADR 9781 from which it differs by 1-3 bp. Topiris is recovered as monophyletic. HT = holotype, PT = paratype, NT = neotype; bp = no of nucleotides sequenced, if not 658 b morphological dataset (65 characters) as analysed for 24 taxa in MrBayes 3.2. 7 under a gamma model, rooted by the type species of Xylorycta (X. luteotactella). The specimen numbers and sexes of the exemplars used for describing the morphological characters are set out in this figure. Where possible, the holotype (HT) [or neotype (NT) was used for scoring. PT = paratype, NT = neotype, m = male, f = female. The male of Topiris candidella (highlighted in blue) is represented by a modern specimen (NHMUK 019223170), since the holotype is incomplete. T. candidella is morphologically cryptic with T. cinderella except for small differences in the male genitalia.	Figure 1. a analysis using IQTree (UFbootstrap values as percentage of 1000) and MrBayes 3.2. 7 (/ posterior probabilities) of DNA barcodes of Topiris, including all unique haplotypes sequenced (except one 133 bp sequence of Topiris albidella), showing their relationship to the genus Athrypsiastis and two exemplars of Metathrinca, rooted by Xylorycta (type species: X. luteotactella). The reconstructed DNA barcode Topiris candidella (NHMUK 010219690, highlighted in blue) from Sarawak falls in a clade with specimens of other members of the BIN cluster BOLD: ADR 9781 from which it differs by 1-3 bp. Topiris is recovered as monophyletic. HT = holotype, PT = paratype, NT = neotype; bp = no of nucleotides sequenced, if not 658 b morphological dataset (65 characters) as analysed for 24 taxa in MrBayes 3.2. 7 under a gamma model, rooted by the type species of Xylorycta (X. luteotactella). The specimen numbers and sexes of the exemplars used for describing the morphological characters are set out in this figure. Where possible, the holotype (HT) [or neotype (NT) was used for scoring. PT = paratype, NT = neotype, m = male, f = female. The male of Topiris candidella (highlighted in blue) is represented by a modern specimen (NHMUK 019223170), since the holotype is incomplete. T. candidella is morphologically cryptic with T. cinderella except for small differences in the male genitalia.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9D59D67AEC045D739692DE6CE2DE8CDA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269566	https://doi.org/10.3897/zookeys.1229.119155.figure2	Figure 2. Combined morphological and molecular tree run in MrBayes 3.2. 7 under a split-partition gamma and inverse gamma model (with posterior supports where > 0.94) and with the morphological part of the matrix mapped onto this tree using WinClada v. 1.00. 08. This tree shows supporting characters (filled circles show forward nonhomoplasious changes, which may be either synapomorphies or autapomorphies; unfilled circles show forward changes with homoplasy or reversal) for Topiris and other ingroup species. The support values in red are posterior probabilities of at least 0.94 for a two partition run in MrBayes 3.2. 7 for the combined 723 character matrix. HT = holotype, NT = neotype, PT = paratype, m = male, f = female. Character numbers appear above the circles. State numbers appear below the circles.	Figure 2. Combined morphological and molecular tree run in MrBayes 3.2. 7 under a split-partition gamma and inverse gamma model (with posterior supports where > 0.94) and with the morphological part of the matrix mapped onto this tree using WinClada v. 1.00. 08. This tree shows supporting characters (filled circles show forward nonhomoplasious changes, which may be either synapomorphies or autapomorphies; unfilled circles show forward changes with homoplasy or reversal) for Topiris and other ingroup species. The support values in red are posterior probabilities of at least 0.94 for a two partition run in MrBayes 3.2. 7 for the combined 723 character matrix. HT = holotype, NT = neotype, PT = paratype, m = male, f = female. Character numbers appear above the circles. State numbers appear below the circles.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
D892D67698445960B07BEE9F9E624773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
D892D67698445960B07BEE9F9E624773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
D892D67698445960B07BEE9F9E624773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
D892D67698445960B07BEE9F9E624773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
D892D67698445960B07BEE9F9E624773.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6E6EFC4D656153848A7E90266FC4F0D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6E6EFC4D656153848A7E90266FC4F0D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269570	https://doi.org/10.3897/zookeys.1229.119155.figures32-39	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	Figures 32–39. Male genitalia of Topiris spp. 32 A, B T. candidella Walker, 1863, Brunei, slide no. NHMUK 010316445 33 A, B T. candidella, Sarawak, slide no. NHMUK 014331342 34 T. cinderella sp. nov. A holotype, slide no. NHMUK 010316400 B paratype, slide no. NHMUK 010316448 35 A, B T. ochrotincta sp. nov., holotype, slide no. NHMUK 010316401 36 A, B T. schneeweissella sp. nov., holotype, slide no. NHMUK 010316358 37 A, B T. sericella sp. nov., holotype, slide no. NHMUK 010316873 38 A, B T. salva (Meyrick, 1932), comb. nov., slide no. NHMUK 010316399 39 A, B T. salva (Meyrick, 1932), comb. nov., paraneotype, slide no. NHMUK 010316441. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6E6EFC4D656153848A7E90266FC4F0D8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269566	https://doi.org/10.3897/zookeys.1229.119155.figure2	Figure 2. Combined morphological and molecular tree run in MrBayes 3.2. 7 under a split-partition gamma and inverse gamma model (with posterior supports where > 0.94) and with the morphological part of the matrix mapped onto this tree using WinClada v. 1.00. 08. This tree shows supporting characters (filled circles show forward nonhomoplasious changes, which may be either synapomorphies or autapomorphies; unfilled circles show forward changes with homoplasy or reversal) for Topiris and other ingroup species. The support values in red are posterior probabilities of at least 0.94 for a two partition run in MrBayes 3.2. 7 for the combined 723 character matrix. HT = holotype, NT = neotype, PT = paratype, m = male, f = female. Character numbers appear above the circles. State numbers appear below the circles.	Figure 2. Combined morphological and molecular tree run in MrBayes 3.2. 7 under a split-partition gamma and inverse gamma model (with posterior supports where > 0.94) and with the morphological part of the matrix mapped onto this tree using WinClada v. 1.00. 08. This tree shows supporting characters (filled circles show forward nonhomoplasious changes, which may be either synapomorphies or autapomorphies; unfilled circles show forward changes with homoplasy or reversal) for Topiris and other ingroup species. The support values in red are posterior probabilities of at least 0.94 for a two partition run in MrBayes 3.2. 7 for the combined 723 character matrix. HT = holotype, NT = neotype, PT = paratype, m = male, f = female. Character numbers appear above the circles. State numbers appear below the circles.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6B445B7B55565B3AA17526B9F00DCA3F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269568	https://doi.org/10.3897/zookeys.1229.119155.figures4-21	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	Figures 4–21. Dorsal images of Topiris spp. 4 Topiris candidella Walker, 1863, holotype ♂ 5 T. candidella, Sarawak, leg. A. R. Wallace, ♂ 6 T. candidella, Brunei, ♂ 7 T. cinderella sp. nov., paratype ♂ 8 T. ochrotincta sp. nov., holotype ♂ 9 T. schneeweissella sp. nov., holotype ♂ 10 T. sericella sp. nov., holotype ♂ 11 T. salva (Meyrick, 1932), comb. nov., neotype ♂ 12 T. salva (Meyrick, 1932), comb. nov., paraneotype ♂ 13 T. salva, ♂ 14 T. albogrisella sp. nov., holotype ♂ 15 T. lacteella sp. nov., paratype ♂ 16 T. madonna sp. nov., holotype ♂ 17 T. meyricki sp. nov., holotype ♂ 18 T. sampitella (Lvovsky, 2014), comb. nov., holotype ♂ 19 T. thunbergella sp. nov., holotype ♂ 20 T. albidella sp. nov., holotype ♂ 21 T. digiticosta sp. nov., holotype ♂. Scale bar: 10 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
6B445B7B55565B3AA17526B9F00DCA3F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9BC8E1F9998D585BA2C368B7598E336C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269569	https://doi.org/10.3897/zookeys.1229.119155.figures22-31	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	Figures 22–31. Dorsal images of Topiris, Athrypsiastis and Paralecta spp. 22 Topiris RMNH. INS. 20000 23 Athrypsiastis phaeoleuca Meyrick, 1910, holotype ♂ 24 A. cheesmanae sp. nov., paratype ♂ 25 A. edelweissella sp. nov., holotype ♂ 26 A. halmaherella (Lvovsky, 2014), comb. nov., holotype ♂ 27 A. penumbrella sp. nov., holotype ♂ 28 A. symmetra Meyrick, 1915, ♂ 29 Paralecta rosiflora (Meyrick, 1930), comb. nov., holotype 30 Topiris salva (Meyrick, 1932) ♀ 31 T. salva ♀. Scale bars: 10 mm (22–29).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9BC8E1F9998D585BA2C368B7598E336C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269573	https://doi.org/10.3897/zookeys.1229.119155.figures57-65	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	Figures 57–65. Female genitalia of Topiris and Linoclostis spp., wing preparations 57 Topiris candidella Walker, 1863, slide no. NHMUK 014331340 58 T. schneeweissella sp. nov., paratype, slide no. NHMUK 010316451 59 T. salva Meyrick, 1932, comb. nov., slide no. NHMUK 014331352 60 A T. albidella sp. nov., paratype, slide no. NHMUK 010316865 60 B corpus bursae imaged during preparation 61 Linoclostis gonatias Meyrick, 1908, type, slide no. JFGC 7678 62 T. candidella, Brunei, slide no. NHMUK 014331889 63 Athrypsiastis cheesmanae sp. nov., paratype, slide no. NHMUK 014331890 64 T. salva, forewing venation slide no. NHMUK 010316844 65 L. gonatias, type, forewing venation, slide no. JFGC 7678. Scale bars: 2 mm (57–61).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9BC8E1F9998D585BA2C368B7598E336C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269574	https://doi.org/10.3897/zookeys.1229.119155.figures66-77	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	Figures 66–77. Topiris and Athrypsiastis, pre-genital abdomens and selected genitalia characters (enlarged) 66 Topiris candidella Walker, 1863, Brunei, slide no. NHMUK 014331344 67 Athrypsiastis phaeoleuca Meyrick, 1910, slide no. NHMUK 010316398 68 T. meyricki sp. nov., holotype, aedeagus, slide no. NHMUK 010316359 69 T. cinderella sp. nov., aedeagus, slide no. NHMUK 010316448 70 T. albidella sp. nov., aedeagus, slide no. NHMUK 014331359 71 A. phaeoleuca Meyrick, 1910 holotype, aedeagus, slide no. NHMUK 010316398 72 T. salva (Meyrick, 1932), antrum, slide no. NHMUK 014331352 73 T. salva (Meyrick, 1932), corpus bursae, slide no. NHMUK 014331352 74 T. albidella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316354 75 T. schneeweissella sp. nov., holotype, uncus and gnathos, slide no. NHMUK 010316358 76 Athrypsiastis phaeoleuca Meyrick, 1910, holotype, uncus and gnathos, slide no. NHMUK 010316398 77 T. cinderella sp. nov., holotype, vinculum, slide no. NHMUK 010316400. Scale bars: 2 mm (66, 67).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9BC8E1F9998D585BA2C368B7598E336C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269575	https://doi.org/10.3897/zookeys.1229.119155.figures78-87	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	Figures 78–87. Selected genitalia characters and labels 78 T. ochrotincta sp. nov., holotype, process of ventral costal membrane, slide no. NHMUK 010316401 79 T. candidella Walker, 1863, sacculus with strong mesad shoulder, slide no. NHMUK 010316360 80 T. candidella Walker, 1863, valva, slide no. NHMUK 014331342 81 T. cinderella sp. nov., holotype, costal ventral membrane of valva distally raised, slide no. NHMUK 010316400 82 T. candidella Walker, 1863, costal ventral membrane of valva distally flat, slide no. NHMUK 010316360 83 T. schneeweissella sp. nov., holotype, valva showing setae, slide no. NHMUK 010316358 84 T. candidella Walker, 1863, processes on S 8, slide no. NHMUK 014331340 85 A T. candidella Walker, 1863 type labels 85 B Wallace’s label (enlarged) 86 T. candidella Walker, 1863 labels of Wallace specimen, specimen no. NHMUK 013700125 87 hindwings at some time wrongly affixed to type of T. candidella. Scale bar: 0.2 mm (83).	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
9BC8E1F9998D585BA2C368B7598E336C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1269571	https://doi.org/10.3897/zookeys.1229.119155.figures40-47	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	Figures 40–47. Male genitalia of Topiris spp. 40 T. albogrisella sp. nov. A paratype, slide no. NHMUK 010316437 B holotype, slide no. NHMUK 010316356 41 A, B T. lacteella sp. nov., holotype, slide no. NHMUK 010316362 42 A, B T. madonna sp. nov., holotype, slide no. NHMUK 010316357 43 A, B T. meyricki sp. nov., holotype, slide no. NHMUK 010316359 44 A, B T. thunbergella sp. nov., holotype, slide no. NHMUK 010316361 45 T. albidella sp. nov., A holotype, slide no. NHMUK 010316354 B paratype, slide no. NHMUK 014331359 46 A, B T. digiticosta sp. nov., holotype, slide no. NHMUK 010316355 47 T. albidella sp. nov., paratype, lateral, slide no. NHMUK 014331360. Scale bar: 1 mm.	2025-03-04	Sterling, Mark J.;Price, Ben W.;Lees, David C.		Zenodo	biologists	Sterling, Mark J.;Price, Ben W.;Lees, David C.			
