identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0947107FB2315F538D7E9F781706A0C3.text	0947107FB2315F538D7E9F781706A0C3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Devario altus Sudasinghe & Pethiyagoda & Meegaskumbura 2025	<div><p>Devario altus sp. nov.</p><p>Fig. 13 A – C</p><p>Devario monticola Batuwita, de Silva &amp; Udugampala, 2017 (in part).</p><p>Type materials.</p><p>Holotype. 2025.04.01.NH, 48.7 mm SL, Bopathalawa, Menik Palama Farm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=80.70638&amp;materialsCitation.latitude=6.8155556" title="Search Plazi for locations around (long 80.70638/lat 6.8155556)">Agra Oya (Mahaweli basin)</a>, 6°48'56"N, 80°42'23"E, 1580 m asl, H. Sudasinghe, 14 June 2019 .</p><p>Paratypes. 2025.04.02.NH, 1, 44.2 mm SL (same data as holotype); DZ 5260, 13, 41.9–54.9 mm SL (same data as holotype) .</p><p>Diagnosis.</p><p>Devario altus sp. nov. is distinguished from its Sri Lankan congeners by the combination of the following characters and character states: anal-fin origin posterior to dorsal-fin origin, beneath 3 rd branched dorsal-fin ray; 1 st infraorbital usually bearing a laterally-directed process; P-stripe continuous, not bifurcated anteriorly; interstripe I + 1 long, extending uninterrupted anterior to vertical through pelvic-fin origin; caudal peduncle long, 19.2–21.4 % SL, shallow, 8.8–10.5 % SL; body shallow, its depth 21.8–26.0 % SL; dorsal-fin height 19.0–23.2 % SL; dorsal-fin base length 15.1–18.2 % SL; anal-fin base length 16.4–20.5 % SL; pelvic fin height 12.9–16.6 % SL; pectoral-fin height 20.3–23.4 % SL; head depth 17.3–19.6 % SL; scales between lateral line and dorsal-fin origin 6–7 ½; circumpeduncular scales, 13–14; branched dorsal-fin rays 8 ½ - 10 ½.</p><p>Specifically, D. altus sp. nov. differs from D. malabaricus by possessing (in 73 percent of specimens) a process on the 1 st infraorbital vs lacking (in 95 percent of D. malabaricus); having the P-stripe continuous, not bifurcated anteriorly (vs P-stripe usually bifurcated anteriorly); interstripe I + 1 long, extending uninterrupted anterior to vertical through pelvic-fin origin (vs short, not extending uninterrupted anterior to a vertical through the pelvic-fin origin in D. malabaricus); a caudal peduncle length of 19.2–21.4 % SL (vs 12.3–19.4 % SL); a body depth of 21.8–26.0 % SL (26.1–35.4 % SL); and scale rows between lateral line and dorsal-fin origin 6–7 ½ (vs 7 ½ - 9).</p><p>The new species is distinguished from D. memorialis by having the anal-fin origin beneath the 3 rd branched dorsal-fin ray, distinctly posterior to dorsal-fin origin (vs on vertical through dorsal-fin in D. memorialis); the anal-fin base length 16.4–20.5 % SL (vs 20.0–23.4 % SL); a head depth of 17.3–19.6 % SL (vs 14.1–15.8 % SL); and 13 or 14 (vs 15–18) circumpeduncular scales.</p><p>Devario altus sp. nov. differs from D. micronema by having the I + 1 interstripe long, extending uninterrupted anterior to a vertical through the pelvic-fin origin (vs short, not extending uninterrupted anterior to a vertical through the pelvic-fin origin in D. micronema); having the P-stripe continuous, uninterrupted anteriorly (vs P-stripe usually interrupted anterior to pelvic-fin origin with bars or blotches); having a longer caudal peduncle, 19.2–21.4 % SL (13.2–20.0 % SL); having a body depth of 21.8–26.0 % SL (vs 23.5–36.6 % SL); and having 6–7 ½ (7 ½ - 9 ½) scale rows between lateral line and dorsal-fin origin.</p><p>It is distinguished from Devario pathirana by a shallower body (21.8–26.0 % SL, vs 32.4–36.9 % SL in D. pathirana); a shallower caudal peduncle depth of 8.8–10.5 % SL (vs 11.3–13.1 % SL); a shorter dorsal-fin base (15.1–18.2 % SL, vs 19.0–23.8 % SL); a shorter anal-fin base (16.4–20.5 % SL, vs 20.7–22.8 % SL); a shorter pelvic fin (12.9–16.6 % SL, vs 16.1–18.8 % SL); a shorter pectoral fin (pectoral-fin height 20.3–23.4 % SL, vs 23.8–27.9 % SL); 6–7 ½ (vs 8–8 ½) scale rows between lateral line and dorsal-fin origin; and 8 ½ - 10 ½ (vs 11 ½) branched dorsal-fin rays.</p><p>Devario altus sp. nov. is genetically distinct from D. malabaricus, its closest congener, by 1.2–1.7 % in the cox 1 DNA barcoding marker and genetically distinct from the remaining species of Sri Lankan Devario by more than 3.6 %. Some populations of Devario in the central hills (‘ D. cf. malabaricus (highland phenotype) ’) possess a color pattern resembling that of D. altus sp. nov.: however, some of the individuals sequenced from these populations contain the mitochondrial haplotype for D. altus sp. nov., while most others contain the D. malabaricus mitochondrial haplotype. Sexual dimorphism is apparent in D. altus sp. nov., with males possessing (and females lacking) tubercles on the branched rays of the pectoral fin.</p><p>Description.</p><p>For general appearance, see Fig. 13 A – C; morphometric data are provided in Suppl. material 2. Head and body laterally compressed, elongate. Body depth greatest at pelvic-fin origin. Snout length subequal to eye diameter. Medial margins of dentaries straight, parallel, with an indentation (‘ danionine notch’) anteriorly. Well-developed dermal grooves present along supraorbital shelves. Small, rounded, symphysial knob present, fitting into shallow groove on inner margin of upper jaw. Lower jaw slightly longer than upper. First infraorbital with a process (n = 11) or smooth (n = 4). Tubercles on lower jaw rounded, arranged in a mediodorsal band of 2–3 rows, tapering to a single row posteriorly and towards symphysis, present in both sexes. Single row of rounded tubercles on upper margin of upper jaw. Pectoral-fin tubercles present in males. Both maxillary and rostral barbels present. Rostral barbel longer than maxillary barbel, not reaching anterior margin of orbit; maxillary barbel short, not reaching anterior margin of orbit. Lateral line complete, declining steeply for first 7 scales, then curving parallel to ventral body outline, running low on caudal peduncle, terminating on caudal-fin base, with 36 (2), 37 (6), 38 (3), 39 (1), 40 (2), or 41 (1) pored scales on body, plus 1–2 on caudal-fin base. Median predorsal scales 15 (3), 16 (10), 17 (1), or 18 (1). Lateral scale rows between origins of dorsal and pelvic fins 6 + 1 + 1 (2), ½ 6 + 1 + 1 (6), 7 + 1 + 1 (2), or ½ 7 + 1 + 1 (5). Circumpeduncular scales 13 (7) or 14 (8). Dorsal fin with 2–3 unbranched rays, 8 ½ (1), 9 ½ (10), or 10 ½ (4) branched rays, origin anterior to vertical through anal-fin origin, distal margin straight. Anal fin with 3 unbranched rays, 12 ½ (5), 13 ½ (7), 14 ½ (2), or 15 ½ (1) branched rays, distal margin slightly concave. Pectoral-fin with 1 unbranched and 9 (5), 10 (8) or 11 (2) branched rays, adpressed fin just reaching or surpassing pelvic-fin origin; axial lobe well developed. Pelvic fin with 1 unbranched and 6 (6), or 7 (9) branched rays, origin midway between anal-fin origin and pectoral-fin origin; tip of adpressed pelvic fin not reaching anal-fin origin. Pelvic ‘ axillary’ scale present. Caudal fin with 9 + 8 (15) branched rays, forked, lobes subequal, rounded distally.</p><p>Color pattern.</p><p>For condition in life, see Fig. 13 A; in preservative, see Fig. 13 B, C. In preservative, mid-dorsal stripe narrow, extending from occiput to caudal peduncle. Cleithral spot vertically elongate, covering part of first lateral-line scale and scale above it. P-stripe broad, two scale-widths wide anteriorly, 1 scale-width wide posteriorly, continuing to end of medial caudal-fin rays; anteriorly uninterrupted above pectoral and pelvic fins, bordered dorsally and ventrally by narrow I + 1 and I, respectively. Interstripe I narrow, of uniform width, about ¼ - ⅓ width of P-stripe, slightly narrow to P- 1, slightly narrower than I + 1. Interstripe I + 1 narrower anteriorly, about ½ - ⅓ width of P-stripe, wider posteriorly, subequal to P-stripe, uninterrupted anterior to pelvic-fin origin. P + 1 originating above middle of pectoral fin, narrower than P, subequal I + 1 anteriorly, tapering to caudal peduncle, not reaching caudal-fin base. Interstripe I + 2 absent. P + 2 absent. P- 1 narrower than P, faint, with scattered melanophores, tapering to caudal peduncle, not reaching caudal-fin base; I- 1, P- 2 absent. Anterior bars indistinct or absent. Dorsal and anal fins hyaline, with light scattering of small melanophores along interradial membranes. Caudal-fin rays pigmented anteriorly; both rays and interradial membranes pigmented posteriorly. Pectoral and pelvic fins hyaline.</p><p>Habitat and distribution.</p><p>Among the localities sampled in the central hills, the only one in which the mitochondrial genotype consists solely of D. altus is Bopathalawa, at 1580 m asl, the highest elevation at which any species of Devario has been recorded in Sri Lanka. Here, the 1–2 m-wide stream traverses Manik Palama, a dairy farm. The only other fish recorded here was the exotic Poecilia reticulata Peters, 1859 .</p><p>Etymology.</p><p>The species-name altus is Latin for ‘ high’, an allusion to this species being restricted to the Sri Lankan highlands. Applied as a noun in apposition. Suggested vernacular name in English: Sri Lanka Montane Danio .</p></div>	https://treatment.plazi.org/id/0947107FB2315F538D7E9F781706A0C3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sudasinghe, Hiranya;Pethiyagoda, Rohan;Meegaskumbura, Madhava	Sudasinghe, Hiranya, Pethiyagoda, Rohan, Meegaskumbura, Madhava (2025): Species without borders: the diversification of giant danios in Sri Lanka (Teleostei, Danionidae, Devario). Evolutionary Systematics 9 (2): 229-258, DOI: 10.3897/evolsyst.9.162251
C2CCC44B8C3B5F479EAD1D5FFDE5DEE8.text	C2CCC44B8C3B5F479EAD1D5FFDE5DEE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Devario malabaricus (Jerdon 1849)	<div><p>Devario malabaricus (Jerdon, 1849)</p><p>Perilampus malabaricus Jerdon, 1849 .</p><p>Devario monticola Batuwita, de Silva &amp; Udugampala, 2017 .</p><p>Danio lineolatus Bleeker, 1863 a .</p><p>Material examined.</p><p>109 ex., see Suppl. material 1.</p><p>Diagnosis.</p><p>Devario malabaricus is distinguished from Sri Lankan congeners by the combination of the following characters and character states: anal-fin origin posterior to dorsal-fin origin, beneath 3 rd branched dorsal-fin ray (vs anal-fin origin on vertical through dorsal-fin origin in D. memorialis); 1 st infraorbital smooth, with no infraorbital process (vs present in D. micronema, D. pathirana, and most specimens of D. altus); P-stripe usually bifurcated anteriorly (vs P-stripe not bifurcated in D. altus and D. memorialis); interstripe I + 1 short, usually not extending uninterrupted anterior to vertical through pelvic-fin origin (vs long, extending uninterrupted anterior to vertical through pelvic-fin origin in D. altus); caudal peduncle length 12.3–19.4 % SL (vs 19.2–21.4 % SL in D. altus); caudal peduncle depth 9.8–12.9 % SL (vs 8.1–9.8 % SL in D. memorialis); body depth 26.1–35.4 % SL (vs 21.8–26.0 % SL in D. altus; 15.9–22.6 % SL in D. memorialis); dorsal-fin base length 16.7–23.5 % SL (vs 13.5–16.3 % SL in D. memorialis); pelvic-fin height 13.3–18.1 % SL (vs 11.6–13.3 % SL in D. memorialis); 33–39 pored lateral-line scales (vs 38–46 in D. memorialis); predorsal scales 14–18 (vs 17–19 in D. memorialis); 12–15 circumpeduncular scales (vs 15–18 in D. memorialis); 7 ½ - 9 scales between lateral line and dorsal-fin origin (vs 7 ½ in D. memorialis; 6–7 ½ in D. altus); 9 ½ - 12 ½ branched dorsal-fin rays (vs 8 ½ in D. memorialis).</p><p>Devario malabaricus is genetically distinct from D. altus, its closest congener, by 1.2–1.7 % in the cox 1 DNA barcoding marker, and from the remaining Sri Lankan species of Devario by more than 2.8 %. Sexual dimorphism is apparent in D. malabaricus, with the branched rays of the pectoral fin exhibiting tubercles in males but not in females (Sudasinghe 2024).</p><p>Habitat and distribution.</p><p>Devario malabaricus is the only non-endemic member of the genus in Sri Lanka: it occurs also in peninsular India. It is the most widespread species of Devario on the island, a generalist that utilizes lotic and lentic, shaded and unshaded, clear and turbid, and disturbed and undisturbed habitats, from lowland floodplains to hill streams and torrents.</p><p>Remarks.</p><p>Kullander et al. (2017) and Sudasinghe and Pethiyagoda (2019) showed that the neotype designation for Perilampus malabaricus Jerdon, 1849 by Batuwita et al. (2017) is inconsistent with Article 75.3. 1 of the International Code of Zoological Nomenclature and hence, invalid. Although Devario malabaricus lacks a name-bearing type, it is common in its type locality, Malabar, which is unambiguously defined in contemporaneous maps. While Indian samples of D. malabaricus were not included in the present analysis, previous work (Batuwita et al. 2017; Sudasinghe et al. 2020 a) has shown through both morphological and genetic analysis, that the Indian and Sri Lankan populations are conspecific.</p></div>	https://treatment.plazi.org/id/C2CCC44B8C3B5F479EAD1D5FFDE5DEE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sudasinghe, Hiranya;Pethiyagoda, Rohan;Meegaskumbura, Madhava	Sudasinghe, Hiranya, Pethiyagoda, Rohan, Meegaskumbura, Madhava (2025): Species without borders: the diversification of giant danios in Sri Lanka (Teleostei, Danionidae, Devario). Evolutionary Systematics 9 (2): 229-258, DOI: 10.3897/evolsyst.9.162251
FADBB3695EC4533BB7D322969544F857.text	FADBB3695EC4533BB7D322969544F857.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Devario memorialis Sudasinghe, Pethiyagoda & Meegaskumbura 2020	<div><p>Devario memorialis Sudasinghe, Pethiyagoda &amp; Meegaskumbura, 2020 a</p><p>Devario memorialis Sudasinghe, Pethiyagoda &amp; Meegaskumbura, 2020 a .</p><p>Material examined.</p><p>10 ex., see Suppl. material 1.</p><p>Diagnosis.</p><p>Devario memorialis is distinguished from its Sri Lankan congeners by the combination of the following characters and character states: anal-fin origin on vertical through dorsal-fin origin (vs anal-fin origin posterior to dorsal-fin origin, beneath 3 rd branched dorsal-fin ray in D. malabaricus, D. micronema, D. altus, and D. pathirana); 1 st infraorbital smooth, with no infraorbital process (vs infraorbital process present in D. micronema, D. pathirana, and most specimens of D. altus); anterior bars on body absent or indistinct (vs present in D. malabaricus, D. micronema, and D. pathirana); preanal length 58.5–63.8 % SL (vs 64.9–69.8 % SL in D. pathirana); caudal peduncle depth 8.1–9.8 % SL (vs 9.8–12.9 % SL in D. malabaricus, 11.3–13.1 % SL in D. pathirana); body depth 15.9–22.6 % SL (vs 26.1–35.4 % SL in D. malabaricus, 23.5–36.6 % SL in D. micronema, 21.8–26.0 % SL in D. altus, 32.4–36.9 % SL in D. pathirana); dorsal-fin height 15.7–20.0 % SL (vs 19.0–23.2 % SL in D. altus, 20.6–24.3 % SL in D. pathirana); dorsal-fin base length 13.5–16.3 % SL (vs 16.7–23.5 % SL in D. malabaricus, 16.6–23.3 % SL in D. micronema, 19.0–23.8 % SL in D. pathirana); anal-fin base length 20.0–23.4 % SL (vs 16.4–20.5 % SL in D. altus); pelvic-fin height 11.6–13.3 % SL (vs 13.3–18.1 % SL in D. malabaricus, 13.8–18.5 % SL in D. micronema, 12.9–16.6 % SL in D. altus, 16.1–18.8 % SL in D. pathirana); pectoral-fin height 19.1–23.6 % SL (vs 23.8–27.9 % SL in D. pathirana); head depth 14.1–15.8 % SL (vs 16.0–23.0 % SL in D. micronema, 17.3–19.6 % SL in D. altus, 17.6–21.7 % SL in D. pathirana); interorbital width 37.2–40.7 % HL (vs 40.7–45.3 % HL in D. pathirana); rostral barbel 4.0–8.9 % HL (vs 8.3–14.2 % HL in D. altus, 8.6–17.7 % HL in D. pathirana); 38–46 lateral-line scales (vs 33–39 in D. malabaricus); predorsal scales 17–19 (vs 14–18 in D. malabaricus, 13–17 in D. micronema, 14–15 in D. pathirana); 15–18 circumpeduncular scales (vs 12–15 in D. malabaricus, D. micronema, 13–14 in D. altus, D. pathirana); 7 ½ scales between lateral line and dorsal-fin origin (vs 7 ½ – 9 in D. malabaricus, 7 ½ – 9 ½ in D. micronema, 6–7 ½ in D. altus, 8–8 ½ in D. pathirana); 8 ½ branched dorsal-fin rays (vs 9 ½ – 12 ½ in D. malabaricus, D. micronema, 8 ½ – 10 ½ in D. altus, 11 ½ in D. pathirana); 14 ½ – 15 ½ branched anal-fin rays (vs 13 ½ – 14 ½ in D. pathirana).</p><p>Devario memorialis is genetically distinct from the remaining Sri Lankan species of Devario by an uncorrected p-distance of more than 3.9 % in the cox 1 DNA barcoding marker.</p><p>Habitat and distribution.</p><p>Devario memorialis has been recorded only from a 3 - kilometer section of a headwater tributary of the Ma Oya (river) near Aranayake, at an elevation of 238–266 m above sea level. This species typically inhabits the fast-flowing sections of the stream.</p><p>Remarks.</p><p>At some of the sampling sites at Aranayake, D. malabaricus occurs in syntopy with D. memorialis but at a lower frequency than the latter. All the specimens dissected to establish sex were females. Pectoral-fin tuberculation is absent in all the specimens in our series: it is possible that pectoral-fin tuberculation is absent in both sexes of D. memorialis .</p></div>	https://treatment.plazi.org/id/FADBB3695EC4533BB7D322969544F857	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sudasinghe, Hiranya;Pethiyagoda, Rohan;Meegaskumbura, Madhava	Sudasinghe, Hiranya, Pethiyagoda, Rohan, Meegaskumbura, Madhava (2025): Species without borders: the diversification of giant danios in Sri Lanka (Teleostei, Danionidae, Devario). Evolutionary Systematics 9 (2): 229-258, DOI: 10.3897/evolsyst.9.162251
D2B5A728E3825FE6B3E9747659499FD6.text	D2B5A728E3825FE6B3E9747659499FD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Devario micronema (Bleeker 1863)	<div><p>Devario micronema (Bleeker, 1863 a)</p><p>Danio micronema Bleeker, 1863 a .</p><p>Devario annnataliae Batuwita, de Silva &amp; Udugampala, 2017 .</p><p>Devario udenii Batuwita, de Silva &amp; Udugampala, 2017 .</p><p>Material examined.</p><p>137 ex., see Suppl. material 1.</p><p>Diagnosis.</p><p>Devario micronema is distinguished from Sri Lankan congeners by the combination of the following characters and character states: anal-fin origin posterior to origin of dorsal fin, beneath 3 rd branched dorsal-fin ray (vs anal-fin origin on vertical through dorsal-fin origin in D. memorialis); 1 st infraorbital with an infraorbital process (vs infraorbital process absent in D. malabaricus and D. memorialis); interstripe I + 1 short, usually not extending uninterrupted anterior to vertical through pelvic-fin origin (vs long, extending uninterrupted anterior to pelvic-fin origin in D. altus); caudal peduncle length 13.2–20.0 % SL (vs 19.2–21.4 % SL in D. altus); body depth 23.5–36.6 % SL (vs 15.9–22.6 % SL in D. memorialis); dorsal-fin base length 16.6–23.3 % SL (vs 13.5–16.3 % SL in D. memorialis); pelvic-fin height 13.8–18.5 % SL (vs 11.6–13.3 % SL in D. memorialis); head depth 16.0–23.0 % SL (vs 14.1–15.8 % SL in D. memorialis); predorsal scales 13–17 (vs 17–19 in D. memorialis); circumpeduncular scales 12–15 (vs 15–18 in D. memorialis); scales between lateral line and dorsal-fin origin 7 ½ - 9 ½ (vs 7 ½ in D. memorialis); branched dorsal-fin rays 9 ½ - 12 ½ (vs 8 ½ in D. memorialis).</p><p>Devario micronema is genetically distinct from D. pathirana, its closest congener by 1.2–1.6 % in the cox 1 DNA barcoding marker, and from the remaining Sri Lankan species of Devario by over 2.8 %. Among these, D. micronema shows, across its range, the greatest phenotypic diversity in terms of color-pattern and habitus (Fig. 14) as well as greatest genetic diversity among the Sri Lankan species of Devario . Several of our specimens are suggestive of mitonuclear discordance and / or phenotypes that are intermediate between D. malabaricus and D. micronema . These specimens are considered in this study as potential hybrids. Sexual dimorphism in D. micronema is apparent, with males exhibiting tubercles on the branched rays of the pectoral fin, a character absent in females (Sudasinghe 2024).</p><p>Habitat and distribution.</p><p>Among the endemic species of Devario, D. micronema is the most widely distributed giant danio in Sri Lanka. It is recorded from clear, shaded, undisturbed, low-elevation rainforest streams in the perhumid southwestern wet zone, from sea level to about 500 m asl in Kelani, Kalu, Bentara, and Gin River basins.</p><p>Remarks.</p><p>The Sri Lankan fishes described by Bleeker (1863 a) have not been reported on since their original description: they were presumed lost. The collection was, however, located among uncatalogued material in RMNH by Martien J P. Van Oijen and Maurice Kottelat ca 2015 (Sudasinghe et al. 2025). One of us (HS) examined the holotype of D. micronema, a 60.0 mm SL specimen (Fig. 13 D). This is entirely consistent with our conception of the species based on the material examined here.</p></div>	https://treatment.plazi.org/id/D2B5A728E3825FE6B3E9747659499FD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sudasinghe, Hiranya;Pethiyagoda, Rohan;Meegaskumbura, Madhava	Sudasinghe, Hiranya, Pethiyagoda, Rohan, Meegaskumbura, Madhava (2025): Species without borders: the diversification of giant danios in Sri Lanka (Teleostei, Danionidae, Devario). Evolutionary Systematics 9 (2): 229-258, DOI: 10.3897/evolsyst.9.162251
4ED80C87D23A5943B0FA3154A640AA30.text	4ED80C87D23A5943B0FA3154A640AA30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Devario pathirana (Kottelat & Pethiyagoda 1990)	<div><p>Devario pathirana (Kottelat &amp; Pethiyagoda, 1990)</p><p>Danio pathirana Kottelat &amp; Pethiyagoda, 1990 .</p><p>Material examined.</p><p>8 ex., see Suppl. material 1.</p><p>Diagnosis.</p><p>Devario pathirana is distinguished from Sri Lankan congeners by the combination of the following characters and character states: it is the only completely barred Devario in the island, with 7–11 bars on the side of the body (vs fewer than 4 bars in D. malabaricus, D. micronema, D. memorialis, D. altus); P-stripe short, ending above the caudal peduncle (P-stripe long, extending uninterrupted above the caudal peduncle in D. malabaricus, D. micronema, D. memorialis, D. altus); anal-fin origin posterior to origin of dorsal fin, beneath 3 rd branched dorsal-fin ray (vs anal-fin origin on vertical through dorsal-fin origin in D. memorialis); 1 st infraorbital with an infraorbital process (vs infraorbital process absent in D. malabaricus, D. memorialis); preanal length 64.9–69.8 % SL (vs 58.5–63.8 % SL in D. memorialis); caudal peduncle length 15.7–19.6 % SL (vs 19.2–21.4 % SL in D. altus); caudal peduncle depth 11.3–13.1 % SL (vs 8.1–9.8 % SL in D. memorialis, 8.8–10.5 % SL in D. altus); body depth 32.4–36.9 % SL (15.9–22.6 % SL in D. memorialis; 21.8–26.0 % SL in D. altus); dorsal-fin height 20.6–24.3 % SL (vs 15.7–20.0 % SL in D. memorialis); dorsal-fin base length 19.0–23.8 % SL (vs 13.5–16.3 % SL in D. memorialis, 15.1–18.2 % SL in D. altus); anal-fin base length 20.7–22.8 % SL (vs 16.4–20.5 % SL in D. altus); pelvic-fin height 16.1–18.8 % SL (vs 11.6–13.3 % SL in D. memorialis; 12.9–16.6 % SL in D. altus); pectoral-fin height 23.8–27.9 % SL (vs 19.1–23.6 % SL in D. memorialis, 20.3–23.4 % SL in D. altus); head depth 17.6–21.7 % SL (vs 14.1–15.8 % SL in D. memorialis); interorbital width 40.7–45.3 % HL (vs 37.2–40.7 % HL in D. memorialis); rostral barbel 8.6–17.7 % HL (vs 4.0–8.9 % HL in D. memorialis); 14–15 predorsal scales (vs 17–19 in D. memorialis); 13–14 circumpeduncular scales (vs 15–18 in D. memorialis); 8–8 ½ scales between lateral line and dorsal-fin origin (vs 7 ½ in D. memorialis; 6–7 ½ in D. altus); 11 ½ branched dorsal-fin rays (vs 8 ½ in D. memorialis); 13 ½ – 14 ½ branched anal-fin rays (vs 14 ½ – 15 ½ in D. memorialis).</p><p>Devario pathirana is genetically distinct from D. micronema, its closest congener by 1.2–1.6 % for the cox 1 DNA barcoding marker and genetically distinct from rest of the Sri Lankan species of Devario by over 3.6 %.</p><p>Habitat and distribution.</p><p>Devario pathirana is confined to the streams associated with the headwaters of the Nilwala River basin in the southwestern wet zone of Sri Lanka. The population of Devario in Suriyakanda, at the type locality of Rasboroides rohani in the Walawe basin, comprises of potential hybrids between D. pathirana and D. malabaricus . This is likely a result of an anthropogenic translocation (Sudasinghe et al. 2018).</p><p>Remarks.</p><p>At some of the sampling sites in the headwaters of the Nilwala basin, both D. malabaricus and D. micronema were recorded along with D. pathirana (T. Ranasinghe, S. Wijayasekara pers. comm.). This suggests that there may be zones of contact between these species. Sexual dimorphism is apparent in D. pathirana, with males possessing tubercles on the branched rays of the pectoral fin, which are absent in females.</p></div>	https://treatment.plazi.org/id/4ED80C87D23A5943B0FA3154A640AA30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Sudasinghe, Hiranya;Pethiyagoda, Rohan;Meegaskumbura, Madhava	Sudasinghe, Hiranya, Pethiyagoda, Rohan, Meegaskumbura, Madhava (2025): Species without borders: the diversification of giant danios in Sri Lanka (Teleostei, Danionidae, Devario). Evolutionary Systematics 9 (2): 229-258, DOI: 10.3897/evolsyst.9.162251
