identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8B1967F9F05858A2B02B034DB946FE69.text	8B1967F9F05858A2B02B034DB946FE69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chimarra (Chimarra) buenaventura Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Chimarra (Chimarra) buenaventura sp. nov.</p><p>Fig. 3</p><p>Type material.</p><p>Holotype male: Ecuador: • El Oro: Reserva Buenaventura, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.76829&amp;materialsCitation.latitude=-3.65367" title="Search Plazi for locations around (long -79.76829/lat -3.65367)">small stream near Umbrellabird Lodge</a>, 3.65367°S, 79.76829°W, el. 530 m, 11.x.2023, Ríos-Touma (UMSP 000551868) (UMSP) . Paratypes: same as holotype, 2 females (MECN); 1 female (UMSP) .</p><p>Diagnosis.</p><p>Chimarra buenaventura is a particularly distinctive species in the form of its tergum X, with very short and strongly sclerotized lateral lobes, terminating in a distinctive short, narrow projecting sclerite, and with the two sensilla of the lobe located on a recurved process near its distal margin. The phallotremal sclerite of various species in the group is often associated with a textured sclerotized “ curl, ” modified into a row of short spines in some species. In Chimarra buenaventura this structure seems to be modified into a distinct short, curved spine, buttressed basally by a sclerotized projection, appearing as a second short, branched tine. Females of many of the species have been illustrated. The female of C. buenaventura is distinguished by a pair of distinctively formed cup-shaped sclerites at the apicoventral attachment site of the vaginal apparatus, and by sinuate, paired sclerites on its ventral surface, which extend anterolaterally to the anterior cup-like sclerite, a generalized feature of the vaginal apparatus in Chimarra .</p><p>Description.</p><p>Adult. Forewing length male 5.0 mm, female 5.0– 5.25 mm (n = 3). Color pale fuscous, partially denuded (specimens in alcohol). Male. Abdominal segment IX, in lateral view, with very pronounced sinuous extension of anteroventral margin and short apodemes from anterodorsal margin; posteroventral process elongate, posteriorly projecting, subacute apically. Tergum X membranous mesally, with short, sclerotized lateral lobes; in lateral view, each more or less parallel-sided, subtruncate apically, with very narrow, acute, apical projection, projection laterally recurved, continuous with membranous dorsal extension of lateral lobe; sensilla of each lobe two, on reflexed sclerotized curl, near apex of sclerotized lateral lobe. Preanal appendage short, rounded, in lateral view, wider than long. Inferior appendage, in lateral view, with ventral margin broadly rounded; bearing minute triangular cusp on mesal margin, in ventral or caudal view; dorsal thumb-like process of inferior appendage, in lateral view, projecting posteriorly, strongly mesally curved, with apex expanded and rounded. Phallus with ventral margin of phallobase distinctly projecting; phallic spines two, asymmetrically arrayed, anterior spine elongate, straight, posterior spine shorter, slightly curved; endotheca textured with small spines and also possessing short, curved spine, with basal tine-like projection; phallotremal sclerite complex composed of elongate rod and ring structure and membranous structure with pair of associated wishbone-like sclerites apically; endotheca not expanded in specimen examined, but phallotremal sclerite complex apparently associated with short, curved endothecal spine.</p><p>Female. Ventral sclerites of sternum IX with prominent membranous lateral pouches (“ clasper receptacles ” of Blahnik 1998). Vaginal apparatus moderately elongate, largely membranous, with pair of distinctly cupped sclerites posteroventrally (attachment or anchoring site of vaginal apparatus), posterodorsal sclerites absent; at midlength, with rounded, membranous lateral projections, ventral surface with paired sclerites, narrow and sinuous, extending from membranous lateral projections to anterior cup-like sclerite, flanking lateral margin of vaginal apparatus anteriorly.</p><p>Etymology.</p><p>Named after Reserva Buenaventura of Fundación Jocotoco, located in the southwestern foothills of the Andes and contains elements of the Tumbesian dry forests of southern Ecuador and the Chocó humid forests of northwestern Ecuador. In addition to rare and endemic birds and plants, it is the type locality of this new species.</p><p>Remarks.</p><p>Chimarra buenaventura is a new species in the ortiziana group, as defined by Blahnik (1998). Additional species were subsequently described by Blahnik and Holzenthal (2012) and Holzenthal et al. (2022). The group currently contains fifteen species, distributed from Mexico to the northern parts of South America, with one species extending to southeastern Brazil. This new species adds a sixteenth species. Species of the group have a characteristically formed inferior appendage, with a relatively short and mesally curved, thumb-like, dorsal projection, usually somewhat dilated and rounded apically. The various species are most readily distinguished by the form and shape of the lateral lobes of tergum X, and the position and disposition of its two sensilla.</p></div>	https://treatment.plazi.org/id/8B1967F9F05858A2B02B034DB946FE69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
AFA0645D39505941BD6382F0CA638087.text	AFA0645D39505941BD6382F0CA638087.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chimarra (Otarrha) buglas Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Chimarra (Otarrha) buglas sp. nov.</p><p>Fig. 2</p><p>Type material.</p><p>Holotype male: Ecuador: • Morona Santiago: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.43163&amp;materialsCitation.latitude=-2.99616" title="Search Plazi for locations around (long -78.43163/lat -2.99616)">tributary to Río San Antonio, between Limón Indanza &amp; San Antonio</a>, 2.99616°S, 78.43163°W, el. 1016 m, 15.ii.2023, Ríos, Holzenthal, Amigo, Huisman (UMSP 000551083) (MECN).</p><p>Diagnosis.</p><p>Character similarities between C. buglas and C. ramosa Holzenthal, Blahnik &amp; Ríos-Touma, 2022 include the general structure of the inferior appendage, with acute mesal projections both basally and at midlength; short, but distinct, projections from the posterior margin of tergum VIII; and an apically truncate posteroventral process on segment IX. The most distinctive and diagnostic feature of C. buglas is the structure of tergum X, which like C. ramosa is divided mesally, with each lateral lobe also divided into a dorsal and ventral lobe; the ventral lobe, in C. buglas is submembranous and weakly sclerotized preapically, ending in a distinctly sclerotized spine-like projection. This is perhaps the most distinctive and diagnostic feature of the species. Other diagnostic differences include the structure of the dorsal lobes of tergum VIII, which are acute apically, rather than rounded, and covered with small spines. Additionally, the ventral process of segment IX, in lateral view, is wider apically, thus more dramatically truncate, and the inferior appendage, as viewed ventrally, has an apex that is more elongate, acute, and mesally curved. The apex of the inferior appendage also lacks a preapical projection.</p><p>Description.</p><p>Adult. Forewing length male 4.5 mm. Color pale brown, denuded (specimen in alcohol). Male. Tergum VIII with pair of sclerotized, digitate projections from posterior margin, longer than tergum, apices acute, covered with small spines, denser and more elongate apically. Segment IX, in lateral view, with anterior margin nearly straight (slightly expanded in ventral half), segment longest ventrally, just above ventral process, posterior margin sinuously expanded below preanal appendage; posteroventral process elongate (length ~ 2 × width), widening apically, apex distinctly truncate. Posteromesal projection of tergum IX short, subtriangular, wide basally, concavely narrowed laterally, acute apically, ~ 1 / 3 length of lateral lobes of tergum X. Tergum X divided mesally, forming two sclerotized lateral lobes; in lateral view, lateral lobes each also divided into dorsal and ventral lobes; dorsolateral lobe distinctly shorter than ventrolateral lobe, nearly straight, uniform in width, with multiple sensilla apically and short setae dorsally at approximately midlength; ventral lobe, in lateral view, straight, narrow, submembranous preapically, apex laterally curved, strongly sclerotized, spine-like. Preanal appendage large, flattened, ear-like. Inferior appendage elongate, narrow, mesally curved, tapering apically; in ventral view, with apex strongly inturned, tapering, acute; mesal surface with two tine-like projections: basal tine short and acute, median tine elongate, narrow, sinuate, subequal in length to incurved apex of appendage. Phallus with phallobase short, tubular, distinctly ventrally flexed on ventral margin; endotheca expanded apically, with short membranous apicodorsal projection, internally with elongate spine, wide basally, very narrow and acute apically; phallotremal sclerite complex indistinct, bowed, with converging narrow sclerites.</p><p>Female. Unknown.</p><p>Etymology.</p><p>Named after the Buglas Nature Reserve, an important area for avian conservation, near the type locality.</p><p>Remark.</p><p>Chimarra buglas is a new species in the Chimarra (Otarrha) patosa group, as defined by Blahnik (2002), and likely most closely related to C. ramosa . Including the new species, the group now contains eight species. This is the second species described from Ecuador.</p></div>	https://treatment.plazi.org/id/AFA0645D39505941BD6382F0CA638087	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
94A55B88750A5D0DA919D64B2F580413.text	94A55B88750A5D0DA919D64B2F580413.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chimarrhodella spinosa Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Chimarrhodella spinosa sp. nov.</p><p>Fig. 1</p><p>Type material.</p><p>Holotype male: Ecuador: • Zamora Chinchipe: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.13041&amp;materialsCitation.latitude=-4.53371" title="Search Plazi for locations around (long -79.13041/lat -4.53371)">Tributary crossing Hwy E 682, N of Valladolid</a>, Tapichalaca Reserve, 4.53371°S, 79.13041°W, el. 1898 m, 12.ii.2023, Ríos, Amigo, Huisman (UMSP 000279961) (MECN).</p><p>Diagnosis.</p><p>Chimarrhodella spinosa is very closely related to C. costaricensis Blahnik &amp; Holzenthal, 1992, resembling it particularly in the structure of tergum X, which is membranous mesally and has sensillate lateral projections, each rounded apically, with the apex reflexed on its ventral margin to form a prominent, acute, ventrally projecting, spine-like projection at approximately midlength. Also, like C. costaricensis, and unlike other species in the galeata group of Chimarrhodella, it has the apices of the inferior appendage acute, resembling species of the peruviana group in this respect. Chimarrhodella spinosa is most readily distinguished from C. costaricensis by the spines of the phallic apparatus. Like C. costaricensis, it has a pair of short, symmetrically positioned, phallic spines basally, although slightly longer in length than in C. costaricensis; it differs in having an additional pair of spinose tracts, composed of clustered, fine, needle-like spines, anterior to the paired phallic spines. These tracts of fine spines are a unique feature among known species of the genus. Chimarrhodella spinosa also differs from C. costaricensis in its somewhat larger size and longer inferior appendage, with their apicomesal margins more abruptly narrowed at approximately midlength.</p><p>Description.</p><p>Forewing length male 7.0 mm. Color pale stramineous, denuded (specimen in alcohol). Male. Segment IX, in lateral view, with anteroventral margin weakly produced, concavely excised mesally; posterolateral margins with weakly developed, setose, dorsolateral expansions at level of preanal appendage. Tergum X membranous mesally, shallowly notched at apex, laterally with pair of sclerotized, apically sensillate lobes; each lobe with large sclerotized, spine-like, anteroventrally directed projection. Preanal appendage elongate, but short in comparison to other species of Chimarrhodella; inserted dorsad of widest point of posterolateral expansion of segment IX. Inferior appendage, in lateral view, elongate, linear; in ventral view, widest basally, narrowing apically, abruptly and concavely narrowed on mesal surface in apical half, apex with short, acute, mesally directed projection. Phallus with phallobase tubular, elongate, expanded basodorsally, extended posteroventrally; endotheca elongate, with well-developed, preapical, pleated, hood-like expansion; basally with pair of subequal, elongate, phallic spines, lacking basal enlargements, and additional paired tracts of fine, needle-like spines; phallotremal sclerite complex elongate, tube-like, weakly sclerotized overall, slightly more sclerotized basally; ventral margin with wide sclerotized strip, narrowing preapically, weakly forked and more sclerotized apically.</p><p>Female. Unknown.</p><p>Etymology.</p><p>The name refers to the tract of fine spines in the endotheca, a unique feature among known species of the genus.</p></div>	https://treatment.plazi.org/id/94A55B88750A5D0DA919D64B2F580413	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
3F639C6BD6735125A476885FA08D33B2.text	3F639C6BD6735125A476885FA08D33B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sumacodella grijalvai Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Sumacodella grijalvai sp. nov.</p><p>Fig. 4</p><p>Type material.</p><p>Holotype male: Ecuador: • Imbabura: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-96.0&amp;materialsCitation.latitude=0.30966" title="Search Plazi for locations around (long -96.0/lat 0.30966)">Bosque Protector &amp; Reserva Biológica Los Cedros</a>, 0.30966°N, 78,781 96°W, el. 1400 m, 02.ix.2023, Ríos-Touma (UMSP 000551804) (MECN).</p><p>Diagnosis.</p><p>Although morphologically distinct from Sumacodella elongata, this new species possesses almost all the apomorphic characters considered to be potentially diagnostic for the genus when it was established. Character similarities include a very elongate segment IX, with the posteroventral margin greatly produced, a dorsal margin that has almost disappeared, and a posterior margin from which both elongate, narrow preanal appendage and an extremely elongate and narrow, parallel-sided, tergum X emerge. The species are also similar in that the sensilla of tergum X are confined to a relatively short, digitate apicomesal projection, bordered laterally by narrow lateral projections. Both species also have inferior appendage with an extremely elongate anteromesal apodeme, and a very elongate and narrowly tubular phallus, lacking a basodorsal expansion. Sumacodella grijalvai differs from S. elongata in having segment IX even more elongate, with its anterolateral apodemes broader, and in having a tergum X that is also more elongate, almost excessively so, extending almost to the apex of the apical segment of the inferior appendage, with the lateral projections near its apex very narrow and laterally hooked apically, rather than simple and digitate. It lacks the short, apically setose projections found at the juncture of the mesal and lateral lobes of tergum X in S. elongata . The tubular phallus has an elongate, narrow, posteriorly directed, spine-like projection at approximately midlength, not present in S. elongata, and has only an indistinct tract of very small spines. Also, the apical segment of the inferior appendage is more broadly rounded apically, especially in lateral view.</p><p>Description.</p><p>Adult. Forewing length male 5.5 mm. Color pale brown, partially denuded (in alcohol). Male. Segment VIII short, sternum and tergum subequal in length, sternum densely covered with short, fine setae, tergum with setae confined to posterior region of segment, posterior margin with membranous extension, tightly surrounding segment IX. Segment IX, in lateral view, very elongate, strongly tapering, anterolateral margins modified dorsally to form pair of large, tapering apodemes, rounded apically, ventral margin strongly produced posteriorly, subtruncate in dorsal and ventral views, posterior margin very obliquely narrowed dorsally, with lateral margin converging from ca. mid-height to anterior margin; in dorsal view, with posterior margin forming V-shaped convergence at anterior margin. Tergum X extremely elongate, narrow, nearly parallel-sided, extending to (or almost to) apex of apical segment of inferior appendage; in lateral view, upturned in apical extremity forming digitate mesal lobe, densely covered with sensilla, bordered by pair of very narrow, sclerotized lateral lobes, shorter than mesal lobe, acute, laterally hooked apically. Preanal appendage elongate, extending nearly half length of tergum X, very narrow for most of length, widened preapically. Inferior appendage bi-segmented, segments subequal in length, nearly uniform in width; apical segment rounded, with dense pad of short, stiff apical spines, extended anteriorly on ventromesal surface; with extremely elongate anteroventral apodeme. Phallus very elongate, narrow, tubular, without basodorsal projection; ventrally with elongate, narrow, spine-like projection just past midlength, extending parallel to phallobase to its apex; phallus internally with only small tract of very minute spines, scarcely evident. Phallotremal sclerite very indistinct, weakly sclerotized, small, ring-like.</p><p>Female. Unknown.</p><p>Etymology.</p><p>We dedicate this species to Dr. Agustín Grijalva, a former Constitutional Judge of Ecuador, who successfully applied the constitutional “ Rights of Nature ” provision in a ruling that saved Bosque Protector Los Cedros from mining concessions. This Constitutional Court mandate protected this species and all those inhabiting Los Cedros.</p></div>	https://treatment.plazi.org/id/3F639C6BD6735125A476885FA08D33B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
6402FC81EE1659FE9B412C722C32BA25.text	6402FC81EE1659FE9B412C722C32BA25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia insolita Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Wormaldia insolita sp. nov.</p><p>Fig. 6</p><p>Type material.</p><p>Holotype male: Ecuador: • Pichincha: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.86681&amp;materialsCitation.latitude=0.03249" title="Search Plazi for locations around (long -78.86681/lat 0.03249)">small forest stream, Milpe Bird Sanctuary</a>, 0.03249°N, 78.86681°W, el. 1125 m, 20.xi.2023, Ríos, Holzenthal, Frandsen, Amigo (UMSP 000507032) (UMSP) . Paratype: Ecuador: • Pichincha: small trickle, Milpe Garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.87086&amp;materialsCitation.latitude=0.03739" title="Search Plazi for locations around (long -78.87086/lat 0.03739)">Mindo Cloud Forest Foundation</a>, 0.03739°N, 78.87086°W, el. 1114 m, 21.xi.2023, Ríos, Holzenthal, Frandsen, Amigo, 1 male (UMSP 000551499) (MECN) .</p><p>Diagnosis.</p><p>Wormaldia insolita, because of the very distinctive and unusual development of its inferior appendage, is readily identified and unlikely to be confused with any other described species. Particularly diagnostic is the apical segment of the inferior appendage, which has its apex strongly deflexed and narrowed, and has a longitudinal tract of very dense and minute, spine-like setae on its mesal surface, not quite extending to the apex. The basal segment of the inferior appendage is very bulbously enlarged. Like W. milpe sp. nov. and related species, it has a distinctive mesal notch on the posterior margin of tergum VIII. However, we are uncertain about the closest affinity of this new species.</p><p>Description.</p><p>Adult. Forewing length male 4.5 mm (n = 2). Color fuscous. Male. Sternum VII without distinct posteromesal projection or process, very slightly convexly produced mesally. Tergum VIII with posterior margin modified to form broad mesal notch, bordered laterally by short rounded sclerotized projections, margin of notch also sclerotized; sternum VIII unmodified, only weakly notched mesally. Segment IX, in lateral view, with anterolateral margin broadly and convexly rounded, posterior margin nearly straight, narrowing dorsally. Tergum X, in lateral view, with apex rounded, sensillate; preapically, on dorsal margin, with concavely rounded excavation; tergum, as viewed dorsally, subtriangular, narrow apically, widened basally, with small, subtriangular lateral projections at approximately midlength. Preanal appendage elongate, digitate, somewhat irregular in shape, extending to preapical excavation of tergum X, apparently fused to posterior margin of segment IX, lateral to base of tergum X. Inferior appendage, in lateral view, with basal segment short and very wide, length subequal to width, wide at apex; apical segment, in lateral view, subequal in length to basal segment, basally as wide as apex of basal segment, extending short distance and then abruptly ventrally flexed from dorsal margin, narrowed, acute apically, with apex projecting downward; posterior and apical margins of apical segment with elongate setae, pad of very dense, minute, spine-like setae on mesal surface, though not exposed in lateral view, distinctly visible along posterior margin, visible in dorsal view, but not directly visible, though apparent, in ventral view; in ventral view, both segments of inferior appendage much narrower and more uniform in width, basal segment wider than apical segment, apical segment subacute apically. Phallus with phallobase expanded basodorsally, very short ventrally, sclerotization extending laterally into endotheca; endotheca moderately elongate, with two elongate, narrow spines, one very strongly curved, with enlarged base, other very narrow and sinuous, with base narrow and nearly as elongate as spine. Phallotremal sclerite indistinct, apparently widened or flared apically.</p><p>Female. Unknown.</p><p>Etymology.</p><p>From the Latin insolita meaning unusual, uncommon or strange, and attributed to the unusual development of its inferior appendage, which are unique for this species in the genus.</p></div>	https://treatment.plazi.org/id/6402FC81EE1659FE9B412C722C32BA25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
AB19308D316358CFA5E35067DCEC1741.text	AB19308D316358CFA5E35067DCEC1741.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia milpe Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Wormaldia milpe sp. nov.</p><p>Fig. 7</p><p>Type material.</p><p>Holotype male: Ecuador: Pichincha: small trickle, Milpe Garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.87086&amp;materialsCitation.latitude=0.03739" title="Search Plazi for locations around (long -78.87086/lat 0.03739)">Mindo Cloud Forest Foundation</a>, 0.03739°N, 78.87086°W, el. 1114 m, 21.xi.2023, Ríos, Holzenthal, Frandsen, Amigo (UMSP 000551500) (MECN).</p><p>Diagnosis.</p><p>Wormaldia milpe most closely resembles W. gallardoi Muñoz-Quesada &amp; Holzenthal, 2015, and several related species, including W. andrea, Muñoz-Quesada &amp; Holzenthal, 2015; W. prolixa Flint, 1991; and W. francovilla Muñoz-Quesada &amp; Holzenthal, 2015 . These species have the apical segment of the inferior appendage approximately subequal in length to the basal segment, only moderately narrower in width, with the apex slightly enlarged in lateral view, and with the apicomesal spine-like setae somewhat longitudinally arranged. The species also have a similar posteromesal notch on tergum VIII, though variably developed, and a tergum X with a distinctly enlarged apex and modified lateral margins. Distributions of the species range from Costa Rica through Colombia. Wormaldia milpe has the posteromesal notch of tergum VIII relatively wide and shallow, with short rounded lateral projections and the mesal notch bordered posteriorly with short angular projections. Tergum X has an apex that is sensillate and very sharply upturned; basally it has a distinctly rounded and flattened projection, with its posterior margin slightly projecting; the lateral margins of tergum X have a uniformly rounded lateral projection in the basal half, with several scattered sensilla. The phallic spines are very unequal in size, one very short and curved, the other much longer and slightly curved; both spines have their bases enlarged, that of the longer spine forming an elongate sclerotized curl. The above characters, considered collectively, adequately diagnosis this new species.</p><p>Description.</p><p>Adult. Forewing length male 3.5 mm. Color pale stramineous, denuded (specimen in alcohol). Male. Sternum VII without distinct posteromesal projection or process, very slightly convexly produced mesally. Tergum VIII with posterior margin modified mesally to form broad and shallow notch, bordered laterally by short, rounded projections, margin of notch with acute sclerotized projections; sternum VIII unmodified, only shallowly indentate mesally. Segment IX, in lateral view, with anterolateral margin convexly rounded, very slightly angular in middle; posterior margin nearly straight (very slightly concave); dorsal margin abruptly narrowed from posterior margin, forming cleft extending almost to anterior margin, where continuous with tergum X and preanal appendage. Tergum X, in lateral view, with apex sensillate, distinctly upturned and enlarged, with dorsally flattened mesobasal protuberance, with posterior margin distinctly projecting; mesobasal protuberance, in dorsal view, rounded, with narrowed base; tergum X, in dorsal view, with apex rounded, narrowing to form continuous narrow mesal sclerotization, lateral margins more membranous, widened anterior to sensillate mesal lobe, subparallel, in basal half forming uniformly convex, but weakly projecting, lateral processes with several marginal sensilla. Preanal appendage apparently fused to base of tergum X basolaterally, digitate, slightly irregular in shape, distinctly shorter in length than tergum X. Inferior appendage with basal segment wide, slightly elongate, enlarged at middle, narrowing apically; apical segment, in lateral view, slightly shorter than basal segment, narrower basally than apex of basal segment, slightly widened and rounded apically; in ventral view, with apex rounded, with tract of short, spine-like setae, extending longitudinally from apex. Phallus semi-membranous, phallobase with bulbous basodorsal expansion, strongly narrowed apically, internally with very short, curved spine with enlarged base, and second much longer, curved, apically acute spine, with elongate, curled basal sclerotization; phallotremal sclerite indistinct, apparently short and tubular.</p><p>Female. Unknown.</p><p>Etymology.</p><p>Named after the type locality, the town of Milpe and the nearby preserved cloud forest, managed by the laudable work of the Mindo Cloud Forest Foundation.</p></div>	https://treatment.plazi.org/id/AB19308D316358CFA5E35067DCEC1741	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
15DF0E2861425A7696CD5B97CF92D35F.text	15DF0E2861425A7696CD5B97CF92D35F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wormaldia nilssoni Holzenthal & Blanhik & Ríos-Touma 2025	<div><p>Wormaldia nilssoni sp. nov.</p><p>Fig. 5</p><p>Type material.</p><p>Holotype male: Ecuador: • Napo: Wildsumaco Lodge, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.59824&amp;materialsCitation.latitude=-0.67631" title="Search Plazi for locations around (long -77.59824/lat -0.67631)">small stream # 2, Benavides Trail</a>, 0.67631°S, 77.59824°W, el. 1479 m, 13.xi.2023, Ríos, Holzenthal, Frandsen, Amigo (UMSP 000551141) (MECN).</p><p>Diagnosis.</p><p>Wormaldia nilssoni most closely resembles W. machadorum Muñoz-Quesada &amp; Holzenthal, 2015, described from Costa Rica and known also from Panama (Armitage et al. 2015). Similarities include the form of the inferior appendage, with a bulbous basal segment and apically narrowed apical segment, with a longitudinal arrangement of short spine-like setae on its mesal surface; and the narrow and lightly sclerotized posteromesal processes of tergum VIII. As compared to W. machadorum, the projections from tergum VIII are more V-shaped, closely situated basally and diverging apically. The anterior margin of segment IX is not as distinctly angular as it is in W. machadorum, or in other species mentioned as being close to that species in its original description. Other diagnostic features of W. nilssoni include the form of tergum X, with its apex slightly upturned and lateral margins broadly rounded, apparently with several scattered sensilla, and with a distinct short basomesal protuberance; and its phallic spines, one very short and curved, and the other lacking any noticeable projection (possibly as an aberration).</p><p>Description.</p><p>Adult. Forewing length male 4.5 mm. Color pale stramineous, denuded (specimen in alcohol). Male. Sternum VII without distinct posteromesal projection or process, very slightly convexly produced mesally. Tergum VIII with pair of posteromesal processes; processes weakly sclerotized, elongate, narrow, digitate, closely opposed basally, diverging apically, forming V-shaped configuration; sternum VIII unmodified, nearly straight. Segment IX, in lateral view, with anterolateral margin convexly rounded, very slightly angular in middle; posterior margin nearly straight (very slightly concave); dorsal margin abruptly narrowed from posterior margin, forming cleft extending almost to anterior margin, where continuous with tergum X and preanal appendage. Tergum X with apex sensillate, slightly enlarged and upturned, with small, subquadrate mesobasal protuberance; in dorsal view, with apex forming rounded projection, somewhat continuous anteromesally, lateral margins forming uniformly convex, but rather weakly projecting, lateral processes with several marginal sensilla. Preanal appendage apparently fused to base of tergum X basolaterally, simple in form, elongate, digitate, narrowed basally, shorter in length than tergum X. Inferior appendage with basal segment relatively elongate, bulbously enlarged at middle, narrowing apically; apical segment, in lateral view, distinctly shorter than basal segment, narrow, nearly uniform in width, narrowing apically; in ventral view, somewhat flattened on mesal surface, with patch of short, spine-like setae, extending longitudinally from apex. Phallus semimembranous, phallobase with bulbous basodorsal expansion, strongly narrowed apically, internally with very short, curved spine with enlarged base, and second basal small, sclerotized structure, lacking accompanying spine (possibly as aberration); phallotremal sclerite indistinct, apparently tubular, inflated at middle.</p><p>Female. Unknown.</p><p>Etymology.</p><p>This species is dedicated to Jonas Nilsson, founder of Wildsumaco Lodge and its surrounding preserved forest, for his efforts to protect this amazing habitat and its near pristine forest.</p></div>	https://treatment.plazi.org/id/15DF0E2861425A7696CD5B97CF92D35F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Holzenthal, Ralph W.;Blanhik, Roger J.;Ríos-Touma, Blanca	Holzenthal, Ralph W., Blanhik, Roger J., Ríos-Touma, Blanca (2025): New Philopotamidae (Insecta, Trichoptera) from Ecuador: seven new species and updated country checklist. ZooKeys 1263: 123-145, DOI: 10.3897/zookeys.1263.147996
