taxonID	type	description	language	source
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	description	(Figs 1 – 12) LSID: urn: lsid: zoobank. org: act: 0 B 5 B 6628 - AC 85 - 4 CAC-A 2 A 4 - E 8742 B 55 BA 27.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	etymology	Etymology: The species is named in honour of Amnon Freidberg.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	diagnosis	Diagnosis: This species can be easily recognized from all known species of Ulidia by the postpronotal lobe with prominent tubercle bearing one thickened, short nail-like postpronotal seta; presence of a dark spot over wing apex; entirely brown cells bc, c, sc; and combination of predominantly shining brownish yellow frons, facial carina and gena; ventral and posterobasal setae on fore femur and anteroventral setae on hind femur strongly thickened; mesonotum elongate, mostly matt rugulose, and prescutular acrostichal setae lacking.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	description	Description: Head (Figs 2 – 4) black, with mostly brown to yellowish brown frons, facial carina and gena. Head ratio h: l: w = 1: 0.8: 1.3. Frons 0.8 × as long as wide, reddish or yellowish brown medially, with black orbit, shining, with narrow silvery white microtrichose vitta turning cuneiform at border of frons and parafacial (Figs 3, 4, arrowed); medially frons conspicuously depressed, finely and sparsely yellowish setulose in anterolateral part. Vertical plates and ocellar triangle shining black; 2 pairs of black orbital setae (anterior fine and short); ocellar setae lateroclinate, 0.9 × as long as posterior orbital seta. Face reddish yellow to brown, lunule black, facial carina mostly reddish yellow 1.7 × as wide as antennal groove and 1.3 × as wide as high, subshining, finely shagreened; epistome black, smooth, without metallic sheen; subgena moderately low. Antennal groove 1.9 – 2 × as high as wide, deep, entirely black, white microtrichose except medioventral surface shining black. Gena reddish yellow to brown, 0.4 – 0.5 × as high as eye; parafacial matt brownish or reddish yellow, with very narrow white microtrichose stripe along eye; gena posteriorly separated with rather blunt genal ridge from entirely black postgena. Occiput entirely black, mostly subshining, rugulose, except vertex, occipital sutures and anterior margin of postgena shining black. Ventral half of occiput conspicuously swollen. Medial vertical seta 0.25 – 0.45 × as long as frons width, 1.1 – 1.2 × as long as lateral vertical and 1.5 – 2.2 × as long as ocellar, orbital, and postocellar setae. Antenna: scape black to brown, pedicel deeply incised, brown or sometimes apically yellow; flagellomere 1 short oval, 1.5 × as long as wide, yellow or brownish yellow, white microtrichose; arista bare, brown. Clypeus black. Palp dark brown, gray microtrichose, with 8 – 9 black lateroventral setae setose. Mouthparts black. Thorax (Figs 2, 5). Scutum and scutellum black, finely rugulose, almost matt; posterior portion of postpronotal lobe, posterior surface of notopleural triangle, postalar lobe, anterior half of katepisternum, and whole anepimeron shining black; postscutellum black, subshining; posteroventral margin of scutellum without microtrichose area; anatergite, katatergite and lateroventral parts of mediotergite subshining black. Scutum 1.5 × as long as wide, with fine, poorly visible brownish setulae in intra-alar area and regular row of 8 – 11 dorsocentral setulae and 1 dorsocentral seta (Fig. 5); acrostichal seta lacking, at most 6 – 7 tiny setulae between dorsocentral setae; postpronotal lobe with short and thickened, nail-like seta inserted into conspicuous projection; proepisternal ridge strongly expressed, bearing 2 rows of 8 – 10 short, sometimes thickened, nail-like setulae; proepisternal seta at most 1.2 × as long as proepisternal setulae; other setae moderately long, black: 2 postsutural supra-alar, 1 intra-alar and 1 postalar. Scutellum transverse, 2.8 × as wide as long, finely wrinkled, subshining black, with 2 pairs of black scutellar setae. Wing (Fig. 6). Hyaline, with dark spot at wing apex and entirely brown cells r 1, bc, c, and sc; veins brown; cell r 4 + 5 apically narrowed, apical section of M straight or slightly arcuate, ending anterior of wing apex; section of costal vein between apices of R 4 + 5 and M very narrow, ≤ 0.25 × as long as vein DM – Cu. Postero-apical extension of cell cup shorter than transverse shoulder of vein Cu 2. Calypters with long white fringe. WL = 3.9 – 4.2 mm. Haltere with base and stem black, knob creamy white. Legs black; most setae and setulae black, except fore basitarsus ventrally with brush of dark yellow setulae; fore femur conspicuously thickened, posterodorsally, postero- and anteroventrally with 4 – 5 short thickened setae inserted into tubercles; mid femur slender, with row of 15 – 17 long and thin setae posteroventrally and 2 rows of slightly shorter setae on anterior surface; mid tibia apically with 1 long and 2 shorter setae; hind femur slightly thickened apically, with 2 strong preapical setae dorsally and 2 rows of 3 – 5 thickened setae, some inserted into tubercles, ventrally. Abdomen entirely black, dorsally matt, finely shagreened, laterally subshining, only tergite 1 with inconspicuous and sparse microtrichia laterally; syntergite 1 + 2 anteriorly narrow, widened posteriorly, with short black setae laterodorsally, tergites 3 – 4 almost bare; tergite 5 of male and female conspicuously shagreened, with sparse and relatively long setae posteriorly; female abdominal tergite 2 without dimple-like structures; female tergite 6 transverse, entirely developed, but hidden underneath of tergite 5, with 2 rows of setulae. Sternites wide in both sexes, separated by rather narrow, black membrane from tergites; sternite 1 wider than long; sternite 2 moderately wide, 1.3 × as long as wide, with desclerotized “ window ” in anterior ⅓; sternites 3 – 4 of male and 3 – 5 of female subquadrate, with few sparse setae (4 – 6 longest at posterior margins); tergite and sternite 6 of female conspicuously narrower than preceding tergites, as wide as and half as long as oviscape, both with 2 rows of setae, setae of posterior row moderately long. Sternites 4 – 6 of female without anteromedial apodemes. Male postabdomen brown to black; pregenital sternites moderately long; sternite 8 with numerous subequal setulae. Epandrium as in Figs 7, 8, cerci deeply separated, with long, anteroventrally directed apices (Fig 7: cyan arrows); lateral surstylus simple, with one mesally directed lobe; medial surstylus entirely flat, forming no lobes, bearing 20 – 25 uniformly thin, moderately long setulae, and no prensisetae. Phallus 1.0 mm long, with stipe almost as long as preglans (without glans); membranous inflatable valve as long as preglans, basally with caecum 2 – 3 × as long as stipe width; preglans without spines; glans with one wide, moderately sclerotized lobe (Figs 9, 10, cyan arrows) and small, strongly sclerotized inner structure. Hypandrium (Fig. 8) asymmetric, with strongly developed lateral sclerite (pregonite) at right side and pair of button-like sensory plates at both sides of basiphallus; phallapodeme dorsoventrally compressed. Ejaculatory apodeme fan-like. Female terminalia: oviscape black, 0.95 × as long as tergite 5, aculeus 0.8 – 0.85 mm long, 7 – 7.5 × as long as wide, with long and thin setae on tergite and sternite 8 and cercal unit (Fig. 11); 3 subspherical spermathecae with smooth surface and inverted necks (Fig. 12).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	materials_examined	Holotype: ♂ Afghanistan: 10 km S of Bamyan [34.7160 ° N 67.8400 ° E], 2800 m, 22 – 30. v. 2010, Yu. Skrylnik (SIZK). Paratypes: 5 ♂ 2 ♀, same data as holotype; 2 ♀, same locality, 12. vi. 2016, Yu. Skrylnik (SIZK, SMNHTAU).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44146FFA1FE2DFCF8FE98E9B9.taxon	biology_ecology	Biology: Unknown.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	description	(Figs 13 – 30)	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	etymology	Etymology: The species is named in honor of Miroslav Barták, a Czech dipterist, the first collector of this species.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	diagnosis	Diagnosis: This species can be easily recognized from all known species of Ulidia by the combination of entirely hyaline cells bc, c, and cup; brownish pterostigma, dark spot at wing apex, creamy haltere, shining occiput, matt, finely shagreened mesonotum, and prescutellar acrostichal seta lacking. In the key to the known Palaearctic species (Morgulis & Freidberg 2014) it runs to the couplet 11. together with U. hirsuta Morgulis & Freidberg, 2014, U. apicalis (Meigen, 1826), and U. sp. nr. apicalis, differing from them by entirely black tarsi, and lack of the prescutellar acrostichal seta. It is similar to U. apicalis from Western Mediterranean region in having moderately large dark apical spot in cells r 1, r 2 + 3 and r 4 + 5, clearly differing from it by the almost round eye (length: height ratio = 1: 1.1 – 1.23) and mostly matt, finely shagreened mesonotum (in U. apicalis, eye vertical oval, length: height ratio = 1: 1.4 – 1.55, mesonotum polished black without sculpture). Similarly to U. hirsuta, it possesses moderately large dark apical spot in cells r 1, r 2 + 3 and r 4 + 5, almost round eye and head in profile, shagreened mesonotum, differing from it by moderately long (setulae at most ⅔ × as long as orbital seta) setulose orbits, lack of prescutellar acrostichal setae, and male genitalia with surstylus covered by fine setulae (in U. hirsuta, vertical and orbital plates of frons long setulose, prescutellar acrostichal setae present, and male genitalia with surstylus covered by thickened prensisetae, as shown by Morgulis & Freidberg 2014: figs 23, 57). This species also somewhat reminds some specimens of variable U. ruficeps in wing pattern and head colouration, as well as entirely black tarsi, readily differing from that species by mesonotum matt, densely shagreened (glossy in U. ruficeps) and palpus short and setulose throughout its length (long and wide, setulose only at base in U. ruficeps). See also diagnosis of U. facialis below.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	description	Description: Head (Figs 14 – 16) black, with frons and facial yellowish brown to dark brown. Head ratio h: l: w = 1: 1 – 1.05: 1.45 – 1.5. Frons 0.6 – 0.7 × as long as wide, yellowish brown to dark brown medially, with orbits widely black, shining, laterally shallowly wrinkled, with small white microtrichose triangular spot at anterolateral margin, slightly convex, with numerous black setulae, 0.5 – 0.65 × as long as orbital setae), inserted into shallow pits. Vertical plates and ocellar triangle shining black; 2 pairs of black orbital setae, posterior orbital seta 0.7 – 0.8 × as long as medial and lateral vertical seta, anterior orbital seta short, hardly distinguishable from surrounding inclinate and lateroclinate setulae; ocellar setae lateroclinate, 0.2 – 0.3 × as long as posterior orbital seta. Face and lunule black, facial carina 1.6 × as wide as antennal groove and as wide as high, subshining, finely shagreened; epistome black, finely shagreened; subgena moderately low. Antennal groove 1.5 – 1.6 × as high as wide, deep, entirely black, white microtrichose except ventral surface shining brown to black. Eye 1.23 × as high as long. Gena reddish yellow to brown, half as high as eye; parafacial matt brownish or reddish yellow to black, with very narrow white microtrichose stripe along eye; gena posteriorly separated with rather blunt genal ridge from entirely black postgena. Occiput entirely black, mostly subshining, rugulose, except smooth vertex, occipital sutures and anterior margin of postgena shining black. Ventral half of occiput conspicuously swollen. Medial vertical seta 0.25 – 0.45 × as long as frons width, 1.1 – 1.2 × as long as lateral vertical and 1.5 – 2.2 × as long as ocellar, orbital, and postocellar setae. Antenna: scape black to brown, pedicel deeply incised, brown or sometimes apically yellow; flagellomere 1 short oval, 1.5 × as long as wide, brownish yellow to black, white microtrichose; arista bare, brown. Clypeus black. Palp dark brown, gray microtrichose, with 8 – 9 black lateroventral setae. Mouthparts black. Thorax (Fig. 17). Scutum and scutellum black, finely rugulose, almost matt; posterior portion of postpronotal lobe, entire notopleural triangle, postalar lobe, anterior half of anepisternum and katepisternum, and whole anepimeron shining black; postscutellum black, subshining; posteroventral margin of scutellum without microtrichose area; anatergite, katatergite and lateroventral parts of mediotergite subshining black, finely rugulose. Scutum wide, 1.1 – 1.2 × as long as wide, with irregular, moderately short setulae in intra-alar area and regular row of 12 – 16 dorsocentral setulae and 1 dorsocentral seta (Fig. 17); acrostichal seta lacking, at most 4 – 6 setulae between dorsocentral rows; postpronotal lobe with 1 thin, unmodified seta and 2 – 3 setulae; proepisternal ridge poorly expressed, bearing numerous fine setulae; proepisternal seta short, not distinguished among setulae; other setae moderately long, black: 1 postsutural supra-alar, 1 intra-alar and 1 postalar. Scutellum moderately long, 1.7 × as wide as long, finely wrinkled, with 2 pairs of black scutellar setae. Wing (Fig. 18) hyaline, with dark spot at wing apex reaching from apex of cell r 1 to middle of cell r 4 + 5, cells bc and base of cell c yellowish, cell sc and veins brown; cell r 4 + 5 moderately narrowed at apex: section of costal vein between apices of R 4 + 5 and M 0.35 – 0.45 × as long as crossvein DM – Cu. Postero-apical extension of cell cup 2 × as long as transverse shoulder of vein Cu 2. Calypters with long white fringe. WL = 3.8 – 4.5 mm. Haltere with base brown, knob creamy. Legs entirely black; most setae and setulae black, except fore and hind basitarsi ventrally with brush of dark yellow setulae; fore femur moderately thickened, with fine, unmodified setae; mid femur narrow, with row of 8 – 10 moderately long and thin setae posteroventrally and shorter setulae on anterior surface; mid tibia apically with 1 long and 2 shorter apical setae; hind femur slightly thickened apically, with 2 strong preapical setae dorsally. Abdomen entirely black, dorsally matt, finely shagreened, laterally subshining, with inconspicuous and sparse microtrichia only at sides of tergite 1; syntergite 1 + 2 anteriorly narrow, widened posteriorly, with short black setae laterodorsally, tergites 3 – 4 almost devoid of setae; tergite 5 of male and female conspicuously shagreened, with sparse and relatively long setae posteriorly; female abdominal tergite 2 without dimple-like structures; female tergite 6 transverse, entirely developed, but hidden underneath tergite 5, with 2 rows of setulae. Sternites wide in both sexes, separated by rather narrow, black membrane from tergites; sternite 1 wider than long; sternite 2 1.3 × as long as wide, with desclerotized ʻwindowʼ in anterior ⅓; sternites 3 – 4 of male and 3 – 5 of female subquadrate, with few sparse setae (4 – 6 longest at posterior margins); tergite and sternite 6 of female conspicuously narrower than preceding tergites, as wide as and half as long as oviscape, both with 2 rows of setae, setae of posterior row moderately long. Sternites 4 – 6 of female without anteromedial apodemes. Male postabdomen (Figs 20 – 27) brown to black; pregenital sternites moderately long; sternite 8 with numerous subequal setulae. Epandrium as in Figs 20 – 22; cerci deeply separated, apically approximated and rounded (Fig. 21); lateral surstylus simple, with one mesally directed lobe (Fig. 21); medial surstylus entirely flat, forming no lobes, bearing 20 – 25 uniformly thin, moderately long setulae, and no prensisetae (Fig. 20). Phallus 1.0 mm long, with stipe almost half as long as preglans (without glans); membranous inflatable valve as long as preglans, basally with caecum 2 – 3 × as long as stipe width; preglans without spines; glans with wide transverse sclerite basally, one wide, moderately sclerotized, apically rounded claw-like lobe and two complex, strongly sclerotized inner structures bearing radiate sculpture (Figs 24 – 27). Hypandrium (Figs 22, 23) asymmetric, with strongly developed lateral sclerite (pregonite) at right side and pair of button-like sensory plates at both sides of basiphallus; phallapodeme dorsoventrally flattened, suboval. Ejaculatory apodeme fan-like. Female terminalia: oviscape black, 0.95 × as long as tergite 5, aculeus 0.85 mm long, 7 – 7.5 × as long as wide, with long and thin setae on tergite and sternite 8 and cercal unit; 3 spherical spermathecae with smooth surface and inverted necks (Figs 29 – 30).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	biology_ecology	Biology: Unknown.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414AFFA4FE2EFB73FC67EF00.taxon	materials_examined	Holotype: ♂ Uzbekistan: Aman-Kutan [pass], alpine meadow, 39.17 ° N 66.54 ° E, 1600 m, 23. v. 1989, M. Barták (SIZK). Paratypes: Qazaqstan: 5 ♂ 6 ♀, Karatau Mts, Aktobe R. 15 km N Atabaj, 750 – 820 m, 8. v. 1994 (2 ♂ [1 dissected], 1 ♂ ♀ in copula); 9. v. 1994 (2 ♂), 10. v. 1994 (5 ♀), Korneyev; 1 ♂ (dissected), Jambyl Region, S slopes of Karatau Mts NE of Biblikol, 1100 m, 19. v. 1988, Antropov; 1 ♀ Jambyl Region, S slopes of Karatau Mts NE of Biblikol, 700 – 1300 m, 19. v. 1998, Kireychuk; 20 ♂ (1 dissected) 11 ♀ 1 ♂ ♀ in copula, Karatau Mts, N Achisai 35 km NNE Kentau 1200 – 1700 m, 12. v. 1994, Korneyev (SIZK, SMNHTAU, ZMHB). Kyrghyzstan: 1 ♂ (dissected), Kara-Archa vill., SW Kyrghyz Alatau Mts., 35 km ESE Jambyl [= Taraz], 3. v. 1994, Korneyev; 2 ♂ 1 ♀, 10 km N Tash Kumyr 1500 m, 21. v. 1994 (1 ♂), 22. v. 1994 (1 ♂ 1 ♀), Korneyev; 1 ♂ 1 ♀, Chichkan R. 1000 m, 12 km N Toktogul, 23. v. 1994, Korneyev; 1 ♀, Chon- Aryk prope Bishkek, 1200 m, 26. v. 1994, Korneyev; 1 ♂ 1 ♀, same, 42 ° 43.2 ' N 74 ° 34.4 ' E, 1060 – 1100 m, 19. vi. 1999, Korneyev & Kameneva; 3 ♀, Talas Region, 20 km ESE of Pokrovka (= 45 km ESE of Taraz), 20 – 22. v. 1988, Kireychuk; 1 ♂, Talas ridge, Chatkal flood plain, 76 km of Jangy-Bazar, 42 ° 04.0 ' N 71 ° 35.8 ' E, 2210 m, 29. vi. 1998 Korneyev & Kameneva; 12 ♂ 9 ♀ Chatkal River valley, 15.3 km of Jangy-Bazar, 41 ° 41.1 ' N 70 ° 39.7 ' E, 1400 – 1700 m, 1. vii. 1998 (7 ♂ 6 ♀), 2. vii. 1998 (5 ♂ 3 ♀), Korneyev & Kameneva (SIZK). Uzbekistan: 24 ♂ 23 ♀, same data as holotype (MBC, SIZK).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	description	(Figs 31 – 36) LSID: urn: lsid: zoobank. org: act: 280 CFF 6 F- 05 A 3 - 434 D-BF 13 - B 72 BAD 21 F 019.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	etymology	Etymology: The species is named in honor of Yuriy Skrylnik, Ukrainian entomologist, collector of the type series.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	diagnosis	Diagnosis: This species can be easily recognized from all known species of Ulidia by the combination of frons, occiput, mesonotum and abdominal tergites 1 – 3 sparsely gray pollinose without shining dots at bases of setae and setulae, entirely hyaline wing, face and gena yellow, frons depressed, palp long, extending beyond oral margins. It is similar to U. ruficeps, widespread from Greece and Israel to Iran and Kyrghyzstan, in having elongate head with strongly depressed frons, widely yellow face, and long palp, readily differing from that species by the uniformly microtrichose frons and mesonotum (in U. ruficeps, frons subshining and mesonotum glossy).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	description	Description: Head (Figs 32 – 35) black, with mostly yellow to yellowish brown frons, facial carina and gena. Head ratio h: l: w = 1: 1.2: 1.6. Frons 0.9 – 1.0 × as long as wide, black in posterior half and yellow to orange in anterior half, entirely and uniformly covered with sparse white microtrichia, including vertical plate, orbits and ocellar triangle (Fig. 34), frons slightly depressed medially, moderately densely and short setulose. Vertical plate with 2 orbital setae and 2 – 3 rows of moderately long erect setulae 0.5 – 0.75 × as long as setae); ocellar setae lateroclinate, 1.0 – 1.2 × as long as posterior orbital seta. Face yellow, lunule conspicuously projected anteriorly, with 6 – 8 setulae; facial carina yellow 0.9 – 1.0 × as wide as antennal groove, subshining, finely shagreened; epistome yellow, smooth, shining; subgena high. Antennal groove 1.5 – 1.6 × as high as wide, deep, entirely yellow, almost entirely white microtrichose. Parafacial yellow, white microtrichose in posterior 0.3 – 0.5 of width along eye. Gena yellow, matt, white microtrichose in posterior part, black setulose, 0.55 × as high as eye (Fig. 32). Occiput black, entirely white microtrichose. Ventral half of occiput conspicuously swollen. Medial vertical seta 0.39 – 0.45 × as long as frons width, 1.0 – 1.1 × as long as lateral vertical and 1.3 – 1.5 × as long as ocellar, orbital, and postocellar setae. Antenna: scape black, pedicel deeply incised, black, white microtrichose; flagellomere 1 round, as long as wide, black, silvery white microtrichose; arista bare, entirely black. Clypeus yellow. Palp black in apical 0.6 of length, brownish yellow at base, with long basoventral setae and a few short marginal apical setae, silvery white microtrichose. Mouthparts black, short, labellum 0.8 × as long as eye; prementum shining black. Thorax (Figs 31, 33) entirely black, covered with silvery white sparse white microtrichia not hiding colour of cuticle; antepronotum, posterior portion of postpronotal lobe, anterior half of anepisternum anterior half of katepisternum, and meron partly shining black; posteroventral margin of scutellum, anatergite, and mediotergite almost entirely microtrichose. Scutum narrow, 1.1 – 1.2 × as long as wide, one intra-alar row of 3 – 4 setulae and regular row of 9 – 11 dorsocentral setulae and 1 dorsocentral seta (Fig. 33); acrostichal seta lacking; at most 4 setulae between dorsocentral setae; postpronotal lobe with 1 seta and 2 – 3 setulae; proepisternal ridge low, with 8 – 10 black setulae; other setae moderately long, black, 1 postsutural supra-alar, 1 intra-alar and 1 postalar seta present. Scutellum transverse, twice as wide as long, with 2 pairs of black scutellar setae. Wing hyaline, with yellow cell bc, gray cell sc, and brown veins; cell r 4 + 5 apically slightly narrowed, vein M reaching costa at wing apex; section of costal vein between apices of R 4 + 5 and M wide, ≤ 0.50 – 0.55 × as long as crossvein DM – Cu. Postero-apical extension of cell cup 1.2 × as long as transverse section of vein Cu 2. Calypters with long white fringe. WL = 2.9 – 3.0 mm. Haltere with base and stem brown, knob brownish yellow. Legs entirely black, mostly white microtrichose; most setae and setulae fine, black; fore basitarsus and 2 basal hind tarsomeres with brush of brownish yellow setulae on ventral surface; mid tibia with 1 apicoventral seta; hind femur slender, with 1 – 2 preapical dorsal setae. Abdomen entirely black, dorsally matt, finely shagreened, sparsely white microtrichose; tergite 2 laterally long setose; syntergite 1 + 2 wide, tergites 3 – 4 setose mosly laterally; female abdominal tergite 2 without dimple-like structures; female tergite 6 hidden underneath of tergite 5. Sternites wide, separated by rather narrow, black membrane from tergites; oviscape shining black, black setulose, 1.2 × as wide as long and 1.1 – 1.3 × as long as tergites 4 and 5. Female terminalia not dissected. Male. Unknown.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	biology_ecology	Biology: Unknown.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B4414FFFBAFE2CFDAFFC54E9EA.taxon	materials_examined	Holotype: ♀ Afghanistan: 10 km S of Bamyan, 34 ° 67.40 ' N 68 ° 80.08 ' E, 2800 – 2900 m, 12. vi. 2016, Yu. Skrylnik (SIZK). Paratype: 1 ♀ Afghanistan: same locality as holotype, 22 – 30. v. 2010, I. Pljusch (SIZK).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	description	(Figs 37 – 54)	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	diagnosis	Diagnosis: This species is similar to U. bartaki n. sp. in having acrostichal seta lacking, wing with darkened pterostigma, frons widely reddish or yellowish brown, finely dotted, mesonotum finely rugulose, haltere knob yellow, and all tarsi entirely black, differing from it by smaller size, surstylus angulate, with 3 – 5 thick prensisetae, cerci short oval, non-produced, and phallus with stipe as long as or longer than preglans, and glans poorly sclerotized, bearing no long projected lobes (in U. bartaki n. sp. size bigger, surstylus apically rounded, with thin setae only, cerci long oval, produced ventrally, phallus with stipe conspicuously shorter than preglans, and glans with at least one moderately sclerotized lobe and largely sclerotized inner structures). Most specimens from Egypt, Israel, Iraq and Iran also differ from U. bartaki n. sp. in having hyaline wing apex without dark spot, whereas the specimens from Greece have apical dark spot. In addition, specimens of U. facialis can be recognized from cell r 4 + 5 strongly narrowed at apex (distance between apices of veins R 4 + 5 and M ≤ 0.33 × as long as crossvein DM – Cu (in other Ulidia, cell r 4 + 5 only slightly narrowed to the apex, distance between apices of veins R 4 + 5 and M ≥ 0.33 ×, usually 0.35 – 0.55 × as long as crossvein DM – Cu).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	description	Redescription: Head (Figs 14 – 16) black, with frons, face, parafacial and gena brownish yellow to dark brown. Head ratio h: l: w = 1: 1.05 – 1.1: 1.35 – 1.4. Frons (from lunule to anterior ocellus) as long as wide, reddish brown to dark brown medially, with orbits widely black, shining, laterally shallowly wrinkled, with small, usually inconspicuous white microtrichose triangular spot at anterolateral margin, slightly convex, with numerous short (0.2 – 0.3 × as long as orbital setae) black setulae inserted into shallow pits. Vertical plates and ocellar triangle shining black; 2 pairs of black orbital setae, of them anterior seta inclinate and posterior seta lateroclinate, 0.6 – 0.7 × as long as medial and lateral vertical seta; ocellar setae lateroclinate, as long as posterior orbital seta. Face and lunule reddish brown to dark brown, facial carina 1.1 × as wide as antennal groove and 1.2 × as wide as high, shining, finely shagreened; epistome brownish yellow to dark brown, finely shagreened; subgena moderately low, slightly lower than width of antennal groove, brown to dark brown, shagreened or rugulose. Antennal groove 1.8 – 1.9 × as high as wide, deep, brownish yellow, white microtrichose except ventral surface shining yellow to brown. Eye 1.1 – 1.15 × as high as long. Gena brownish yellow to brown (sometimes to black in posterior quarter), half as high as eye; parafacial entirely shining, brownish yellow to brown. Occiput entirely black, mostly subshining, rugulose; ventral half of occiput conspicuously swollen. Medial vertical seta 0.65 – 0.75 × as long as frons width, 1.1 × as long as lateral vertical and 1.6 – 1.8 × as long as ocellar and orbital setae; postocellar seta 0.3 – 0.4 × as long as medial vertical seta or lacking. Antenna: scape brown, pedicel shallowly incised, brown to black; flagellomere 1 short oval, 1.1 × as long as wide, brownish to black, white microtrichose; arista bare, brown to black. Clypeus black, at most as long as eye. Palp dark brown, gray microtrichose, with 5 – 6 black lateroventral setae basally and 3 – 4 short setae at apex. Mouthparts black, with labellum short and prementum shining black, elongate, 1.0 – 1.1 × as long as labellum. Thorax (Figs 37, 38, 41) entirely black, scutum and scutellum sparsely rugulose, subshining to matt; other parts subshining, sparsely rugulose or shagreened; posteroventral margin of scutellum without microtrichose area; postscutellum and lateroventral parts of mediotergite subshining black, finely rugulose or inconspicuously microtrichose. Scutum wide, 1.1 – 1.2 × as long as wide, with irregular, moderately short setulae in intra-alar area and regular row of 10 – 12 dorsocentral setulae and 1 dorsocentral seta (Fig. 37); acrostichal seta lacking, at most 4 – 5 setulae between dorsocentral rows; postpronotal lobe with 1 thin, unmodified seta and 2 – 3 setulae; proepisternal ridge poorly expressed, with 1 – 3 fine setulae; proepisternal seta tiny, inconspicuous; other setae moderately long, black: 2 postsutural supra-alar, 1 intra-alar and 1 postalar. Scutellum moderately long, twice as wide as long, sparsely shagreened, with 2 pairs of black scutellar setae. Wing (Figs 43 – 46) hyaline with pale brownish tinge, with cells bc, sc, pterostigma and veins gray to brown; wing apex usually hyaline, in specimens from Greece, pale brown spot at wing apex reaching from apex of cell r 1 to middle of cell r 4 + 5 present; cell r 4 + 5 strongly narrowed to apex: section of costal vein between apices of R 4 + 5 and M 0.25 – 0.3 × as long as vein DM – Cu. Postero-apical extension of cell cup as long as transverse section of vein Cu 2. Calypters with long white fringe. WL = 2.3 – 2.9 mm. Haltere with base brown, knob creamy white. Legs entirely black; all setae and setulae black, except fore tibia, fore and hind basitarsi ventrally with brush of dark yellow setulae; femora slender, with fine, unmodified setae; mid tibia apically with 1 long seta; hind femur slightly thickened apically, with 1 – 2 strong preapical setae dorsally. Abdomen entirely black subshining, finely shagreened, subshining, with inconspicuous and sparse microtrichia only at sides of tergite 1, basal half of tergite 2, and base of tergite 5; syntergite 1 + 2 anteriorly narrow, widened posteriorly, with short black setae laterodorsally, tergites 3 – 4 laterally short and sparsely setose; tergite 5 of male and female sparsely shagreened, with sparse and relatively long setae posteriorly and laterally; female abdominal tergite 2 without dimple-like structures; female tergite 6 transverse, entirely developed, but hidden underneath of tergite 5, with 2 rows of setulae. Sternites wide in both sexes, separated by rather narrow, black membrane from tergites; sternite 1 wider than long; sternite 2 moderately wide, 1.3 × as long as wide, with desclerotized ‘ window’ in anterior ⅓; sternites 3 – 4 of male and 3 – 5 of female subquadrate, with a few sparse setae (4 – 6 longest at posterior margins); tergite and sternite 6 of female conspicuously narrower than preceding tergites, as wide as and half as long as oviscape, both with 2 rows of setae, setae of posterior row moderately long. Sternites 4 – 6 of female without anteromedial apodemes. Male postabdomen (Figs 48 – 54) brown to black; pregenital sternites moderately long; sternite 8 with numerous subequal setulae. Epandrium as in Figs 49, 51, 53; cerci very short, not produced (Fig. 51, cyan arrows); lateral surstylus simple, with two mesally directed, angulate lobes (Fig. 51, red arrows); medial surstylus, bearing 11 – 55 thin, moderately long setulae, and 3 – 4 prensisetae (Fig. 51, black arrow). Phallus with stipe almost as long as or slightly longer than preglans (without glans); membranous inflatable valve as long as preglans; preglans without spines; glans poorly sclerotized, with 3 – 4 short acute projections (Figs 48, 52, 54). Female terminalia: oviscape black, 0.9 × as long as tergite 5, aculeus 0.75 mm long, 8 – 8.5 × as long as wide, with long and thin setae on tergite and sternite 8 and subtriangular cercal unit (see also Morgulis & Freidberg 2014: fig. 96); 3 globular spermathecae with smooth surface and inverted necks.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	biology_ecology	Biology: Unknown.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	materials_examined	Material examined: Syntypes of Ulidia facialis 2 ♀: Egypt: Mariout, 5 – 10. iii. 1929 (EFC). Holotype of Ulidia salonikiensis ♂: Greece: “ Saloniki / 26 301 ”, “ Miari / - smeca I ”, “ Holotypus ” [red label] “ Ulidia / salonikiensis / n. sp. / det. Dr. W. Hennig 1939 ” (ZMHB). Paratypes: 2 ♀: Greece: “ Saloniki / 26 299 ” and “ Saloniki / 26 299 ” [“ Saloniki Turkei an Garry de N. Hough ” according to the Museum Catalogue], “ Paratypus ” (ZMHB). Holotype ♂ and paratypes 1 ♂ 2 ♀ of Ulidia omani: Israel: Be'er Sheva' [Beersheba], 24. iii. 1964 [64], P. Oman (USNM). Holotype ♂ of T. persica: Iran: Belochistan, Iranshahr, 800 m, 1 – 10. iii. 1954, Richter & Schäuffele. Paratypes: 2 ♂, same data as holotype; 11 – 18. iii. 1954, 2 ♂ 3 ♀, SE Iranshahr, Barnpurufer, Richter & Schäuffele, 13. iii. 1954 (SMNS). Other material examined: Egypt: 6 ♂, Mariout, 5 – 10. iii. 1929, Efflatoun (EFC). Israel: 3 ♂ 2 ♀, Ein Avda, 29. v. 1980, Mathis & Freidberg; 1 ♂, Yeruham, 25. iii. 1987, Freidberg; 2 ♂ 1 ♀, Bor Mashash, 25. iii. 1987, Kaplan; 4 ♀, Dimona, 12. iv. 1990, Freidberg; 4 ♂ 3 ♀, Nahal Seher, 6. iv. 1992, Freidberg; 4 ♂ 1 ♀, Mizpe Ramon observatory, 17. iii. 1995, Freidberg (SIZK). Iraq: 1 ♂ 2 ♀, Haira, pools in steppe, 6. iv. 1988, Olejnicek (MBC, SIZK).	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
6C0A87B44151FFBDFE09FAB3FD62E9FE.taxon	discussion	Remarks: Hendel (1931) had used two alternative spellings: “ fascialis ” (in the title) and “ facialis ” (in the body of description); later Hennig (1940) cited both names together and selected the spelling “ facialis ” as correct; according to the Principle of the First Reviser (ICZN 1999: Art. 24), the spelling Ulidia facialis Hendel, 1931 was fixed as the valid name for this species. The statement of Morgulis and Freidberg (2014: 230) that “ the spelling “ facialis ” … is an unjustified emendation ” is therefore incorrect. Probable conspecificity of specimens identified as U. omani and U. facialis was noted by Morgulis and Freidberg (2014), but the synonymy was pending, despite no characters to distinguish them were found. Detailed study by M. S. El-Hawagry of genitalia in the topotypic specimens collected together with the syntype females, but formally not included by Hendel (1931) into the type series, has shown that U. omani and U. facialis have the same structure of the male surstyli, cerci and phallus, which are very constant and species-specific.	en	Kameneva, Elena P., I., Valery A. Korneyev I. (2019): New species and synonymies in the genus Ulidia (Diptera: Ulidiidae) from the Middle East. Israel Journal of Entomology 49 (2): 243-266, DOI: 10.5281/zenodo.3559701, URL: http://dx.doi.org/10.5281/zenodo.3559700
