identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
717B87F7FF91E94DFF4FFE19FBDAFC0C.text	717B87F7FF91E94DFF4FFE19FBDAFC0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colpoptera Burmeister 1835	<div><p>Genus Colpoptera Burmeister, 1835</p><p>Type species: Colpoptera sinuata Burmeister, 1835</p><p>Diagnosis. (Abridged from Gnezdilov &amp; O’Brien 2008). Body elongated, head about as broad as prothorax, eyes prominent. Vertex transverse, anterior margin truncate, posterior margin concave. Frons elongate, with distinct median carina. Pronotum narrow, anterior margin strongly convex, posterior margin concave. Mesonotum at midline longer than vertex + pronotum combine, bearing arcuate anterior transverse carinae and median carina. Forewings elongate and narrowed to rounded apex, with narrow hypocostal plate and many transverse veins apically, hindwings large and functional. Hind tibiae with single lateral tooth distally. Male pygofer with nearly straight hind margin (elongate process absent). Aedeagus relatively simple, bilaterally symmetrical, with elongated, usually bifurcated, subapical lateral processes (“phallobase lobe” of Gnezdilov &amp; O’Brien 2008); and with pair of shorter, simple ventral processes. Gonostyli bearing broad capitulum, narrowing apically with broad lateral tooth. Anal tube elongated. Female sternite VII with rounded median process (Gnezdilov &amp; O’Brien, fig. 59). Gonoplacs nearly triangular, fused medially; gonocoxa VIII with straight hind margin (Gnezdilov &amp; O’Brien, figs. 60-62). Anal tube long and narrow, distinctly longer than gonoplacs Gnezdilov &amp; O’Brien, fig. 57).</p></div>	https://treatment.plazi.org/id/717B87F7FF91E94DFF4FFE19FBDAFC0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Barrantes Barrantes, Edwin A.;Zumbado Echavarria, Marco A.;Bartlett, Charles R.;Bahder, Brian W.	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2025): New planthoppers in the genus Colpoptera (Hemiptera: Auchenorrhyncha: Fulgoroidea: Nogodinidae) from lowland tropical rainforest and cloud forest in Costa Rica. Zootaxa 5689 (1): 114-132, DOI: 10.11646/zootaxa.5689.1.4, URL: https://doi.org/10.11646/zootaxa.5689.1.4
717B87F7FF91E948FF4FFBB8FED4F89E.text	717B87F7FF91E948FF4FFBB8FED4F89E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colpoptera acinaces Bahder & Bartlett 2025	<div><p>Colpoptera acinaces Bahder &amp; Bartlett sp. nov.</p><p>(Figures 2–6)</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.48509&amp;materialsCitation.latitude=10.203231" title="Search Plazi for locations around (long -84.48509/lat 10.203231)">Hotel Villa Blanca</a>, Alajuela Province, Costa Rica (10.203231, -84.485094) (Fig. 1) .</p><p>Diagnosis. Body dark, uniformly fuscous, face uniformly dark, forewing fuscous, less fuscous along costal margin and distal third, pale spot at RA branch apices and CuP apex. Pygofer quadrate in lateral view. Gonostyli from lateral view very broad, bearing large capitulum with helical sclerotized ridges. Aedeagus lightly sclerotized, upcurved, nearly bilaterally symmetrical, bearing pair of strongly bifurcated dorsal processes of moderate length arising subapically on lateral margins and pair of large, broad, long and hooked decussate processes on ventral margin that cross distally, nearly reaching the base. Anal segment elongated, irregularly sinuate on ventral margin, ventral margin with strong invagination approximately 2/3 from base, apex blunt.</p><p>Description. Color. Dark colored (Fig. 2), body uniformly fuscous, face uniformly dark, posterior 2/3 of vertex signficanlty paler, forewing fuscous (Fig. 4), paler along costal vein and distal third, especially near dorsoapical margin, pale spot at RA branch apices and CuP apex. (Fig. 4).</p><p>Structure. Body elongated, length male 5.5 mm, female 5.9 mm (Table 3).</p><p>Head. In dorsal view, vertex nearly rectangular, anterior margin slightly curved, approximately level with anterior margin of eyes, posterior margin concave, median carina absent, transverse carina at fastigium evident (Fig. 3A). In lateral view, generally rounded, slightly angled at fastigium, vertex slightly projected above eyes, frons curved dorsad (Fig. 3B). In frontal view, dorsal margin weakly concave at transverse carina, lateral margins expanding from dorsal margin, reaching widest point just below antennae then constricting strongly to clypeus, median carina evident, extending from fastigium to frontoclypeal suture (Fig. 3C). Eye roughly ovate (long axis diagonal), weakly emarginated above antenna. Lateral ocellus near anterioventral margin of eye. Antennae short, scape short, ring-like, pedicel ovoid, taller than wide, bearing irregularly distributed sensory plaques.</p><p>Thorax. Pronotum in dorsal view broad at midpoint, at midline about 1.5x length of vertex, widest at midpoint with anterior margin convex, posterior margin concave, tricarinate, carina at midline continuous from anterior to posterior margin, lateral carinae extending from anterior margin of pronotum, laterad, becoming obsolete ventrally (Fig. 3A). Mesonotum approximately as wide as long at midpoint, unicarinate anteriorly with pair of carinae (slightly sinuate) forked at diagonal laterally from median carina near anterior margin, extending just beyond obscure lateral carinae (Fig. 3A); in lateral view, mesonotum raised above level of pronotum (Fig. 3B). Hind tibiae with single lateral tooth approximately 3/4 length distad from femoral-tibial joint, spinulation 6-7-2.</p><p>Forewing reticulate, anterior and posterior margin subparallel, asymmetrically rounded at apex (Fig. 4); clavel apex near distal ¾ of forewing; RP arises separately from Sc+RA at basal cell.</p><p>Male terminalia. Pygofer, in lateral view (Fig. 5), quadrate, nearly uniform width along entire length, anterior margin sinuate, posterior margin slightly sinuate, dorsal and ventral margin equal length, slightly sinuate, medioventral process lacking. Gonostyli, in lateral view, very broad, generally rounded with large capitulum arising proximad, with helical sclerotized ridges, apex blunt with posterior corner rounded, anterior corner angulate (Fig. 5). Aedeagus nearly bilaterally symmetrical (Fig. 6), distally upcurved, bearing two pairs of processes; first pair (A1 &amp; A2) arising on subapical lateral margin, slender, strongly and widely bifurcated with dorsal projections (A1a &amp; A2a) curved cephalad proximally, straight distally, ventral projections nearly straight, angled cephalad, extending just beyond apex of A1a and A2a (Fig. 6), from dorsal view (Fig. 6C), processes A1b &amp; A2b converging to meet medially; second pair (A3 &amp; A4) arising on inner ventral margin just proximad of A1 and A2, curving dorsad and cephalad, stout and broadly flattened, narrowed basally, expanding immediately, lateral margins subparallel, apices strongly curved mesad (approximately 90°), decussate, nearly reaching aedeagal base. Anal segment elongated (Fig. 5), irregularly sinuate on ventral margin, dorsal margin angled ventrad approximately ¼ from base, ventral margin with strong invagination approximately 2/3 from base, apex distad of invagination comma-shaped, apex blunt (not downcurved); paraproct small.</p><p>Plant associations. Unknown, collected sweeping vegetation in vacant lot.</p><p>Distribution. Hotel Villa Blanca, Alajuela Province, Costa Rica.</p><p>Etymology. The specific epithet is derived from the Latin word ‘ acinaces ’ (scimitar or short sword), a reference to the large, scimitar-like ventral processes of the aedeagus, and is intended as masculine in gender.</p><p>Material examined. Holotype male “ Costa Rica, Alajuela Pr. / Hotel Villa Blanca / 12.VI.2018 / sweeping weeds / Coll.: B.W.Bahder // Holotype / Colpoptera acinaces ♂ ” (FLREC) . Paratypes (2 males, 6 females); same data as holotype .</p></div>	https://treatment.plazi.org/id/717B87F7FF91E948FF4FFBB8FED4F89E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Barrantes Barrantes, Edwin A.;Zumbado Echavarria, Marco A.;Bartlett, Charles R.;Bahder, Brian W.	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2025): New planthoppers in the genus Colpoptera (Hemiptera: Auchenorrhyncha: Fulgoroidea: Nogodinidae) from lowland tropical rainforest and cloud forest in Costa Rica. Zootaxa 5689 (1): 114-132, DOI: 10.11646/zootaxa.5689.1.4, URL: https://doi.org/10.11646/zootaxa.5689.1.4
717B87F7FF95E944FF4FFF3FFED4FBB6.text	717B87F7FF95E944FF4FFF3FFED4FBB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colpoptera alacutus Bahder & Bartlett 2025	<div><p>Colpoptera alacutus Bahder &amp; Bartlett sp. nov.</p><p>(Figures 7–11)</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.00596&amp;materialsCitation.latitude=10.431269" title="Search Plazi for locations around (long -84.00596/lat 10.431269)">La Selva Biological Station</a>, Heredia Province, Costa Rica (10.431269, -84.005961) (Fig. 1) .</p><p>Diagnosis. Body pale, uniformly testaceous, face uniformly pale, forewing slightly fuscous in distal 1/3, pale along costal margin at midpoint. Apex of forewing pointed. Pygofer quadrate in lateral view. Gonostyli from lateral view very broad, bearing large capitulum with helical sclerotized ridges. Aedeagus lightly sclerotized, upcurved, nearly bilaterally symmetrical, bearing pair of strongly bifurcated dorsal processes of moderate length arising subapically on lateral margins and pair of short, narrow, slightly bulbous processes on ventral margin. Anal segment elongated, slightly sinuate on ventral margin, apex slightly hooked ventrally.</p><p>Description. Color. Pale colored (Fig. 7), body uniformly testaceous, antennae with pedicel dorsally black at base, forewing pale, paler along costal vein at midpoint, lightly fuscous in distal 1/3 (Fig. 9).</p><p>Structure. Body elongated, length male (with wings) 7.5 mm (Table 3).</p><p>Head. In dorsal view, vertex nearly rectangular, anterior margin slightly curved, approximately level with anterior margin of eyes, posterior margin concave, median carina absent, transverse carina at fastigium evident (Fig. 8A). In lateral view, generally rounded, slightly angled at fastigium, vertex slightly projected above eyes, frons curved dorsad (Fig. 8B). In frontal view, dorsal margin weakly concave at transverse carina, lateral margins expanding from dorsal margin, reaching widest point at antennae then constricting strongly to clypeus, median carina evident, extending from fastigium to frontoclypeal suture (Fig. 8C). Eye roughly ovate (long axis diagonal), weakly emarginated above antenna. Lateral ocellus near anterioventral margin of eye. Antennae short, scape short, ring-like, pedicel ovoid, taller than wide, bearing irregularly distributed sensory plaques.</p><p>Thorax. Pronotum in dorsal view broad at midpoint, at midline about 1.5x length of vertex, widest at midpoint with anterior margin convex, posterior margin concave, tricarinate, carina at midline continuous from anterior to posterior margin, weakly developed, lateral carinae extending from anterior margin of pronotum, laterad, becoming obsolete ventrally (Fig. 8A). Mesonotum slightly wider than long at midpoint, unicarinate anteriorly with pair of carinae (slightly sinuate) forked at diagonal laterally from median carina near anterior margin, extending just beyond obscure lateral carinae (Fig. 8A); in lateral view, mesonotum raised above level of pronotum (Fig. 8B). Hind tibiae with single lateral tooth approximately 3/4 length distad from femoral-tibial joint, spinulation 5-7-2.</p><p>Forewing reticulate, anterior and posterior margin subparallel, asymmetrically rounded at apex (Fig. 9); claval apex near distal ¾ of forewing; RP arises separately from Sc+RA at basal cell. Apex pointed, posterior margin in distal 1/3 straight (Fig. 9)</p><p>Male terminalia. Pygofer, in lateral view (Fig. 10), quadrate, nearly uniform width along entire length, anterior margin sinuate, posterior margin slightly sinuate, dorsal and ventral margin equal length, slightly sinuate, medioventral process lacking. Gonostyli, in lateral view, very broad, generally rounded with large capitulum arising proximad, with helical sclerotized ridges, apex blunt with posterior corner rounded, anterior corner angulate (Fig. 10). Aedeagus nearly bilaterally symmetrical (Fig. 11), distally upcurved, bearing two pairs of processes; first pair (A1 &amp; A2) arising on subapical lateral margin, slender, strongly and widely bifurcated with dorsal projections (A1a &amp; A2a) curved ventrad and angled cephalad, ventral projections (A1b &amp; A2b) curved ventrad, angled cephalad, extending just beyond apex of A1a and A2a (Fig. 11), from dorsal view (Fig. 11C), processes A1b &amp; A2b converging to meet medially; second pair (A3 &amp;A4) arising on inner ventral margin just proximad of A1 and A2, curving dorsad and cephalad, short, narrow basally, expanding slightly at midpoint, constricting to apex. Anal segment elongated, slightly sinuate on ventral margin, dorsal margin angled ventrad approximately ¼ from base, apex pointed, hooked ventrad (Fig. 10).</p><p>Plant associations. Unknown, collected sweeping trailside vegetation.</p><p>Distribution. La Selva Biological Station, Heredia Province, Costa Rica.</p><p>Etymology. The specific epithet is derived from the Latin words ‘ ala ’ for wing and ‘ acutus ’ for pointed, referencing the unique shape of the wing.</p><p>Material examined. Holotype male “ Costa Rica, Heredia Pr. / La Selva Biological Station / 12.VI.2018 / sweeping vegetation / Coll.: B.W.Bahder // Holotype / Colpoptera alacutus ♂ ” (FLREC) . Paratypes (2 males); same data as holotype .</p></div>	https://treatment.plazi.org/id/717B87F7FF95E944FF4FFF3FFED4FBB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Barrantes Barrantes, Edwin A.;Zumbado Echavarria, Marco A.;Bartlett, Charles R.;Bahder, Brian W.	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2025): New planthoppers in the genus Colpoptera (Hemiptera: Auchenorrhyncha: Fulgoroidea: Nogodinidae) from lowland tropical rainforest and cloud forest in Costa Rica. Zootaxa 5689 (1): 114-132, DOI: 10.11646/zootaxa.5689.1.4, URL: https://doi.org/10.11646/zootaxa.5689.1.4
717B87F7FF99E95EFF4FFF3FFB63F824.text	717B87F7FF99E95EFF4FFF3FFB63F824.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colpoptera falx Bahder & Bartlett 2025	<div><p>Colpoptera falx Bahder &amp; Bartlett sp. nov.</p><p>(Figures 12–16)</p><p>Type locality. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-84.00596&amp;materialsCitation.latitude=10.431269" title="Search Plazi for locations around (long -84.00596/lat 10.431269)">La Selva Biological Station</a>, Heredia Province, Costa Rica (10.431269, -84.005961) (Fig. 1) .</p><p>Diagnosis. Dark species, body generally uniformly fuscous, slight less fuscous around eyes and along anterior margin of mesnotum, forewing fuscous, less fuscous along costal vein and distal third, face without markings. Aedeagus lightly sclerotized, nearly symmetrical, pair of strongly bifurcated dorsal processes of moderate length arising subapically on lateral margins and pair of large, broad and hooked processes on ventral margin that cross distally.</p><p>Description. Dark species, body generally uniformly fuscous, slight less fuscous around eyes and along anterior margin of mesonotum, forewing fuscous, less fuscous along costal vein and distal third, face without markings (Fig. 12).</p><p>Structure. Body elongated, length male (with wings) 5.2–5.3 mm, female 5.4–5.6 mm (Table 4).</p><p>Head. In dorsal view, vertex nearly rectangular, anterior margin slightly curved, approximately level with anterior margin of eyes, posterior margin concave, median carina absent, transverse carina at fastigium evident (Fig. 13A). In lateral view, generally rounded, slightly angled at fastigium, vertex slightly projected above eyes, frons curved dorsad (Fig. 13B). In frontal view, transverse carina at fastigium evident, lateral margins expanding from dorsal margin, reaching widest point just below antennae then constricting strongly (Fig. 13C).</p><p>Thorax. Pronotum in dorsal view widest at midpoint with anterior margin moderately convex, posterior margin moderately concave, tricarinate, carina at midline continuous from anterior to posterior margin, lateral carinae extending from anterior margin of pronotum, laterad, becoming obsolete ventrally (Fig. 13A). Mesonotum approximately as wide as long at midpoint, intiatially unicarinate at anterior margin, two additional carinae arising at mid carina near anterior margin, extending diagonally to just beyond lateral carinae, slightly sinuate (Fig. 13A). In lateral view, greatly raised, angled where diagonal carinae arise at mid carina (Fig. 13B). Hind tibiae with single lateral spine approximately 3/4 distance from femoral-tibial joint, spinulation 6-7-2.</p><p>Forewing reticulate, broadest basally, constricting at claval apex, expanding slightly, apex broadly rounded (Fig. 14).</p><p>Male terminalia. Pyfoger in lateral view nearly uniform width along entire length, anterior margin sinuate, posterior margin slightly sinuate, dorsal and ventral margin equal length, slightly sinuate, medioventral process lacking (Fig. 15). Gonostyli in lateral view generally rounded with large dorsal process arising basally, with helical sclerotized ridges, apex blunt with posterior corner rounded, anterior corner hooked (Fig. 15). Aedeagus nearly symmetrical with two pairs of processes (Fig. 16); first pair (A1 &amp; A2) arising on subapical lateral margin, strongly bifurcated with dorsal projections (A1a &amp; A2a) curved cephalad basally, straight distally, ventral projections nearly straight, angled cephalad, extending just beyond apex of A1a and A2a (Fig. 16), second pair (A3 &amp; A4) arising on inner ventral margin just basad of A1 and A2, curving dorsad and cephalad, broadly flattened, narrowed basally, expanding immediately, lateral margins subparallel, apex strongly curved mesad (approximately 90°), reaching aedeagal base. Anal segment elongate, irregularly sinuate on dorsal and ventral margins, dorsal margin angled ventrad approximately ¼ from base, ventral margin with strong invagination approximately 2/3 from base, apex distad of invagination comma-shaped.</p><p>Plant associations. Unknown, collected sweeping trailside vegetation.</p><p>Distribution. La Selva Biological Station, Heredia Province, Costa Rica.</p><p>Etymology. The specific epithet ‘ falx ’ is a reference to the large, scythe-like ventral process of the aedeagus.</p><p>Material examined. Holotype male “ Costa Rica, Heredia Pr. / La Selva Biological Station / 12.VI.2018 / sweeping weeds / Coll.: B.W.Bahder // Holotype / Colpoptera falx ♂ ” (FLREC) . Paratypes (3 males, 4 females); same data as holotype .</p><p>Sequence Data. For the COI gene, 18S gene, and for the 28S D8 and D9-D10 expansion region, sequence data was successfully obtained and accessioned (Table 2). There was strong to moderate bootstrap support (100, 89, 99 respectively for 18S, D8 and D9–D10) for the monophyly of Colpoptera relative to out-groups with all new taxa resolving within Colpoptera (Fig. 17). Based on COI, C. acinaces sp. nov. and C. falx sp. nov. resolve adjacent to eachother with strong bootstrap support (100). Additionally, C. alacutus sp. nov. resolves adjacent to C. japortla with moderate bootstrap support (60) for COI. These relationships were also reflected in the consensus tree based on concatenated data for all markers with strong bootstrap support for both the relationship of C. acinaces sp. nov. to C. falx sp. nov. (99) and C. alacutus sp. nov. to C. japortla (93) (Fig. 18). Finally, the consensus tree based on concatenated data for all loci analyzed for available taxa show strong bootstrap support (100) for the genus Colpoptera based on the taxa analyzed (Fig. 18).</p><p>Remarks. The general habitus of C. acinaces sp. nov., C. alacutus sp. nov. and C. falx sp. nov. appears similar to many described species of Colpoptera . Additionally, both the form of the terminalia and geography appear to exclude the other described genera in the Colpopterini (most genera are Antillean). Of the novel taxa, C. alacutus sp. nov. possesses an aedeagus that is similar to many taxa within the Colpoptera (dorsal pair of processes that are deeply bifid and small ventral pair of processes that are all generally symmetrical). However, the wing shape of C. alacutus sp. nov. is more pointed than many of the described taxa. The only species with somewhat pointed apical margins of the forewings are C. acutata and C. caldwelli, both of which have similar structure of the genitalia. However, C. caldwelli is a very dark/fuscous species whereas C. alacutus sp. nov. is very pale. Despite color, C. caldwelli has an aedeagal structure more similar to C. alacutus sp. nov. than does C. acutata . Additional differences that preclude the novel taxon from C. caldwelli is the shape of the anal segment, where it is long and slender in C. alacutus sp. nov. and more robust with a truncated apex in C. caldwelli (where it is tapered in C. alacutus sp. nov.). Finally, the shape of the anal segment differs between C. alacutus sp. nov. and C. acutata as well as the color patterns described, where C. alacutus sp. nov. is lighter in color and lacks the white markings on the frons observed in C. acutata . Both C. acinaces sp. nov. and C. falx sp. nov. also superficially resemble many species of Colpoptera based on body shape and general color pattern (resembling C. marginalis and C. sinuata most noticeably based on original descriptions, however being darker than these species), the aedeagus of these novel taxa is significantly different from the structure observed in any other taxa. In both C. acinaces sp. nov. and C. falx sp. nov. the ventral processes are very broad, flattened and curved mesad, beak-like when paired and individually resembling swords. The two species differ primarily from each other in the size of these ventral processes with C. acinances sp. nov. having ventral processes only reaching the midpoint of the aedeagal shaft whereas the processes in C. falx sp. nov. extend to the base of the aedeagus. Potentially related to this difference is the size of the dorsal processes, where C. acinaces sp. nov. has larger dorsal processes compared to C. falx sp. nov. which inversely has large ventral processes. There are also subtle differences in the form of the anal segment (primarily the apex) that also separate these species. The similarity of the ventral processes in these two species appear to be reflected in the phylogeny generated based on the loci analyzed. These taxa resolve adjacent to each other with strong support (and lower than normal variance COI compared to other taxa) and the similar form (differing mainly in size) of structures indicate these taxa are very closely related.</p><p>While females were unavailable for C. alacutus sp. nov., the terminal sternites for females of C. acinances sp. nov. and C. falx sp. nov. (Fig. 19) were observed. The overall general form among the two species is very similar, with the only noticeable difference being that the apex of the tergite of C. acinaces sp. nov. is rounded whereas it is slightly truncated in C. falx sp. nov. Because these novel taxa are very similar in appearance, male morphology and genetically, it is not unexpected that this feature in females does not differ significantly. In Caldwell (1945), this character appears useful in distinguishing species, and for more distantly related species appears so. However, based on the observations of this structures here, this feature may not be reliable for closely related taxa.</p></div>	https://treatment.plazi.org/id/717B87F7FF99E95EFF4FFF3FFB63F824	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Barrantes Barrantes, Edwin A.;Zumbado Echavarria, Marco A.;Bartlett, Charles R.;Bahder, Brian W.	Barrantes Barrantes, Edwin A., Zumbado Echavarria, Marco A., Bartlett, Charles R., Bahder, Brian W. (2025): New planthoppers in the genus Colpoptera (Hemiptera: Auchenorrhyncha: Fulgoroidea: Nogodinidae) from lowland tropical rainforest and cloud forest in Costa Rica. Zootaxa 5689 (1): 114-132, DOI: 10.11646/zootaxa.5689.1.4, URL: https://doi.org/10.11646/zootaxa.5689.1.4
