identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
7E563D16FF80FFBFD8482570FF74FC4A.text	7E563D16FF80FFBFD8482570FF74FC4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Villaria coriacea (Merr.) Arriola & Alejandro 2020	<div><p>1. Villaria coriacea (Merr.) Arriola &amp; Alejandro, comb. nov.</p><p>Hypobathrum coriaceum Merr. (1926) 490. — Lectotype (designated here): BS(Ramos) 42615, (lecto US[photo!(US 00138112)];isolecto K!(K0003039)), Philippines, Province of Bohol, Bilar.</p><p>Shrubs to small trees, 3–6 m high; branches glabrous. Stipules ovate to narrowly ovate, 7–8.2 by 4–5 mm, keel prominent from the lower 1/3 up to the tip, outer surface glabrous, inner surface densely pilose at the base, margin glabrous; colleters numerous, basally inside, 0.2–0.4 mm long. Leaves: petioles 2–3 by 1–1.5 mm, glabrous; blades oblong to oblanceolate, 3–16 by 2.5–4.5 cm, base acute, apex acuminate, coriaceous, glabrous on both sides except midrib sparsely pubescent beneath; secondary veins 10–11 on each side. Inflorescences supra-axillary, subsessile, 2–3 mm above the node; many-flowered glomerules; peduncle very short, &lt;5 mm long; bracts ovate, 0.9–1.2 by c. 0.5 mm, apex acuminate, pubescent outside, densely pilose inside, colleters few at the base inside. Flowers subsessile, pedicel suppressed. Calyx: tube infundibuliform, 2.5–3 by c. 2.5 mm, pubescent throughout; lobes ovate, c. 1 by 1.5 mm, pubescent outside, densely pilose inside, margins densely ciliate. Corolla: tube infundibuliform, 1.5–2.5 by c. 1.5 mm, glabrous outside, densely pilose inside; lobes ovate, 2.5–3 by c. 2.5 mm, glabrous outside, densely pilose at the base inside, margins sparsely ciliate. Anthers 2–2.5 by 0.3–0.5 mm, with upper 1/3 partially exserted. Style 0.5–0.9 mm long; stigmatic branches elliptic, 2–2.3 by 0.9–1.3 mm, papillate, projected beyond the mouth of the corolla tube. Fruits green, 0.6–1 by 0.7–1.2 cm; stalk 0.5–1.5 cm long, glabrous. Seeds ovate to oblong, 1–2 mm long, yellow.</p><p>Distribution — Philippines (Aurora, Bataan, Bohol, Bulacan, Camarines Sur, Laguna, Quezon, Sorsogon, Tarlac, Zambales Prov.).</p><p>Habitat &amp; Ecology — In primary forest at 300–450 m, also in coastal areas. Flowering: April to June; fruiting: May to September.</p><p>Additional specimens studied. THE PHILIPPINES, Bataan, Arriola &amp; Banag TA025 (USTH); Bilar, Arriola T028 (USTH); Camarines Sur, PPI (Reynoso, Sagcal &amp; Fuentes) 14909 (K); Cavite, Arriola &amp; Cruz T027 (USTH); Rizal, Arriola T031 (USTH); Tarlac, Vidal y Soler 387 (K); Zambales, BS (Edaño) 38339 (K).</p><p>Note — This species resembles V. glomerata because of its oblong to oblanceolate leaf blades (leaf blades broadly lanceolate in V. glomereta) and many-flowered glomerules. Villaria coriacea is distinct by its abaxially sparsely pilose stipules, apically acuminate bracts, larger corolla tube, and shorter fruit stalk.</p></div>	https://treatment.plazi.org/id/7E563D16FF80FFBFD8482570FF74FC4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriola, A. H.;Alejandro, G. J. D.	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
7E563D16FF83FFBFDB07272EFC3BFE44.text	7E563D16FF83FFBFDB07272EFC3BFE44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Villaria multibracteata (Elmer) Arriola & Alejandro 2020	<div><p>2. Villaria multibracteata (Elmer) Arriola &amp; Alejandro, comb. nov.</p><p>Hypobathrum multibracteatum Elmer (1934) 3250. — Lectotype (designated here): Elmer 15674 (lecto NY131934), Philippines, Province of Sorsogon, Irosin.</p><p>Shrubs to small trees, 2–2.5 m tall; branches glabrous. Stipules triangular, 5.5–6 by 1.5–2 mm, keel prominent from the lower 1/3 to the tip, outer surface glabrous, inner surface densely pilose at the base, margin sparsely ciliate, colleters few, 0.3–0.5 mm long. Leaves: petioles 2–4 by c. 2 mm, glabrous; blades lanceolate to ovate, 2.5–5.5 by 2.5–4 cm, recurved towards the tip, base obtuse, apex acuminate, submembranous, glabrous on both sides except for the sparsely pubescent midrib underneath; secondary veins 8–11 on each side. Inflorescences supra-axillary, 2–2.5 mm above the node, few-flowered dichasial cymes; peduncles 1–3.5 cm long, glabrous; pedicels 4–8 mm long, glabrous; bracts triangular, 0.6–1.5 by c. 0.4 mm, apex acute, glabrous outside, densely pilose inside, margin sparsely ciliate; colleters few at the base inside. Calyx: tube infundibuliform, 2–3 by 1.5–2 mm, glabrous throughout; lobes triangular, 0.3–0.5 by c. 0.2 mm, glabrous on both sides, margins sparsely ciliate. Corolla: tube infundibuliform, 2.5–3 by 0.8–1.5 mm, glabrous inside and outside; lobes ovate, c. 1.5 by 0.5–1 mm, glabrous outside, sparsely pilose at the base inside, margins glabrous. Anthers 0.9–1.2 by c. 0.5 mm, with upper 1/3 partially exserted. Style 1–1.5 mm long; stigmatic branches narrowly elliptic to elliptic, c. 2.5 mm long, papillate inside, projected beyond the mouth of the corolla tube. Fruits green, 1–2.3 by 1–1.5 cm; stalk 1–2.5 cm long, glabrous. Seeds ovate to oblong, 3–5 mm long, brown.</p><p>Distribution — Philippines (Aurora, Bataan, Bulacan, Laguna, Quezon, Samar, Sorsogon, Zambales Prov.).</p><p>Habitat &amp; Ecology — Along forest trails and deep ravines in primary forest. Flowering: January to March; fruiting: March to May.</p><p>Additional specimens studied. THE PHILIPPINES, Bataan, Arriola T023 (USTH), Arriola &amp; Banag TA023 (USTH), Vidal y Soler 3076 (K), Whitford 1061 (K), Williams 1061 (K); Irosin, Elmer 15925 (P); Mariveles, Loher 1477 (K); Quezon, BS (Ramos &amp; Edaño) 28586 (K); Samar, BS (Ramos) 17503 (K, P); Sorsogon, Arriola &amp; Buera T012 (USTH), Elmer 15511 (P).</p><p>Note — Villaria multibracteata resembles V. purpurea in its lanceolate leaf blades and cymose inflorescences. However, V. multibracteata differs by its narrower leaf blades with fewer nerves and the presence of multiple triangular bracts. This feature of the bracts distinguishes the species from all other Villaria species.</p></div>	https://treatment.plazi.org/id/7E563D16FF83FFBFDB07272EFC3BFE44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriola, A. H.;Alejandro, G. J. D.	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
7E563D16FF83FFBFD848213CFB5AF7C5.text	7E563D16FF83FFBFD848213CFB5AF7C5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Villaria purpurea (Elmer 1906) Arriola & Alejandro 2020	<div><p>3. Villaria purpurea (Elmer) Arriola &amp; Alejandro, comb. nov.</p><p>Tricalysia purpurea Elmer (1906) 70. — Hypobathrum purpureum (Elmer) Merr. (1923) 534. — Type: Elmer 7659 (lost in PNH, no duplicates found), Philippines, Province of Tayabas, Mount Banahaw. Neotype (designated here): FB (Borden) 2928 (neo K!; isoneo K!, L), Philippines, Province of Bataan, Mount Mariveles. See note 2.</p><p>Shrubs to small trees, up to 3 m tall; branches glabrous. Stipules triangular, 5–7 by 3–4 mm, keel prominent from the base to the tip, outer surface glabrous, inner surface pilose at the base, margin sparsely ciliate, colleters numerous, 0.8–1 mm long. Leaves: petioles 7–8 by 1.5–2 mm, glabrous; blades lanceolate to narrowly ovate, 7–12 by 2.5–4 cm, base acute, apex acuminate, subcoriaceous, glabrous on both sides; secondary veins 8–12 on each side. Inflorescences supra-axillary, 3.5–5 mm above the node, few- to many-flowered dichasial cymes; peduncle 3–4.5 by c. 0.9 mm, glabrous; pedicels 1–1.3 cm long, glabrous; bracts triangular, 0.3–0.8 by c. 0.3 mm, apex acuminate and sharply pointed, glabrous outside, densely pilose inside, margins sparsely ciliate; colleters numerous at the base inside. Calyx: tube infundibuliform, 2.5–5 by c. 3 mm, glabrous throughout; lobes ovate, 0.9–1.3 by 0.5–1 mm, glabrous on both sides, margin sparsely ciliate. Corolla: tube infundibuliform, 1.5–2 mm long, glabrous on both sides; lobes ovate, 3–3.5 by 2–2.5 mm, glabrous outside, sparsely pilose on the base inside, margin sparsely ciliate. Anthers 2.5–3 by c. 1 mm, base truncate, apex obtuse. Style 1–2.3 mm long; stigmatic branches elliptic, 4–4.5 by c. 1 mm, papillate inside. Fruits green, purple when mature, 2.4–4 by c. 2 cm; stalk 3.5–4.6 cm long, glabrous. Seeds angular-ovoid, 6 – 8 mm long, greenish to yellow.</p><p>Distribution — Philippines (Bataan, Quezon, Aurora, Sorsogon, Surigao, Visayas, Zambales Prov.).</p><p>Habitat &amp; Ecology — In primary forest to deep wooded ravines, also near coastal areas. Flowering: April to June; fruiting: May to September.</p><p>Additional specimens studied. THE PHILIPPINES, Alabat, Quezon, BS (Ramos &amp; Edaño) 48148 (P); Alas-asin, Bataan, Arriola &amp; Banag TA002 (USTH); Angat, Bulacan, BS (Ramos &amp; Edaño) 34067 (P); Baler, Aurora, Alejandro et al. 10­110 (USTH); Bataan, FB (Pascual) 28779 (P); Bulusan, Elmer 15925 (L); Ilocos, BS (Ramos) 33007 (K); Irosin, Elmer 16598 (K); Lamao, Bataan, Borden 2928 (P), Borden 21726 (K); Maria Aurora, Arriola &amp; Alejandro 11­048 (USTH); Montalban, Rizal, Loher 6428 (K); Quezon, Merrill 34067 (P); Rizal, BS (Ramos) 41871 (P); Surigao, Banag &amp; Tandang SU024 (PNH); Visayas, Langenberger 2074 (L).</p><p>Notes — 1. The lanceolate to narrowly ovate leaf shape of V. purpurea resembles that of V. acutifolia and V. glomerata . However, V. purpurea differs from all other known Villaria species by having the longest fruit stalks of all.</p><p>2. Elmer (1906) indicated Elmer 7959 to be the type. No duplicate with this number was found. In the Biodiversity Heritage Library, the number in the scan of Elmer (1906) has been changed by hand to 7616 (a number, among others, also mentioned by Merrill 1923), of which a duplicate is present in E. However, it is not clear that Elmer made a typing error. Therefore, we select a neotype here.</p></div>	https://treatment.plazi.org/id/7E563D16FF83FFBFD848213CFB5AF7C5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Arriola, A. H.;Alejandro, G. J. D.	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
