taxonID	type	description	language	source
7E563D16FF80FFBFD8482570FF74FC4A.taxon	description	Hypobathrum coriaceum Merr. (1926) 490. — Lectotype (designated here): BS (Ramos) 42615, (lecto US [photo! (US 00138112)]; isolecto K! (K 0003039 )), Philippines, Province of Bohol, Bilar. Shrubs to small trees, 3 – 6 m high; branches glabrous. Stipules ovate to narrowly ovate, 7 – 8.2 by 4 – 5 mm, keel prominent from the lower 1 / 3 up to the tip, outer surface glabrous, inner surface densely pilose at the base, margin glabrous; colleters numerous, basally inside, 0.2 – 0.4 mm long. Leaves: petioles 2 – 3 by 1 – 1.5 mm, glabrous; blades oblong to oblanceolate, 3 – 16 by 2.5 – 4.5 cm, base acute, apex acuminate, coriaceous, glabrous on both sides except midrib sparsely pubescent beneath; secondary veins 10 – 11 on each side. Inflorescences supra-axillary, subsessile, 2 – 3 mm above the node; many-flowered glomerules; peduncle very short, <5 mm long; bracts ovate, 0.9 – 1.2 by c. 0.5 mm, apex acuminate, pubescent outside, densely pilose inside, colleters few at the base inside. Flowers subsessile, pedicel suppressed. Calyx: tube infundibuliform, 2.5 – 3 by c. 2.5 mm, pubescent throughout; lobes ovate, c. 1 by 1.5 mm, pubescent outside, densely pilose inside, margins densely ciliate. Corolla: tube infundibuliform, 1.5 – 2.5 by c. 1.5 mm, glabrous outside, densely pilose inside; lobes ovate, 2.5 – 3 by c. 2.5 mm, glabrous outside, densely pilose at the base inside, margins sparsely ciliate. Anthers 2 – 2.5 by 0.3 – 0.5 mm, with upper 1 / 3 partially exserted. Style 0.5 – 0.9 mm long; stigmatic branches elliptic, 2 – 2.3 by 0.9 – 1.3 mm, papillate, projected beyond the mouth of the corolla tube. Fruits green, 0.6 – 1 by 0.7 – 1.2 cm; stalk 0.5 – 1.5 cm long, glabrous. Seeds ovate to oblong, 1 – 2 mm long, yellow. Distribution — Philippines (Aurora, Bataan, Bohol, Bulacan, Camarines Sur, Laguna, Quezon, Sorsogon, Tarlac, Zambales Prov.). Habitat & Ecology — In primary forest at 300 – 450 m, also in coastal areas. Flowering: April to June; fruiting: May to September. Additional specimens studied. THE PHILIPPINES, Bataan, Arriola & Banag TA 025 (USTH); Bilar, Arriola T 028 (USTH); Camarines Sur, PPI (Reynoso, Sagcal & Fuentes) 14909 (K); Cavite, Arriola & Cruz T 027 (USTH); Rizal, Arriola T 031 (USTH); Tarlac, Vidal y Soler 387 (K); Zambales, BS (Edaño) 38339 (K). Note — This species resembles V. glomerata because of its oblong to oblanceolate leaf blades (leaf blades broadly lanceolate in V. glomereta) and many-flowered glomerules. Villaria coriacea is distinct by its abaxially sparsely pilose stipules, apically acuminate bracts, larger corolla tube, and shorter fruit stalk.	en	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
7E563D16FF83FFBFDB07272EFC3BFE44.taxon	distribution	Distribution — Philippines (Aurora, Bataan, Bulacan, Laguna, Quezon, Samar, Sorsogon, Zambales Prov.). Habitat & Ecology — Along forest trails and deep ravines in primary forest. Flowering: January to March; fruiting: March to May. Additional specimens studied. THE PHILIPPINES, Bataan, Arriola T 023 (USTH), Arriola & Banag TA 023 (USTH), Vidal y Soler 3076 (K), Whitford 1061 (K), Williams 1061 (K); Irosin, Elmer 15925 (P); Mariveles, Loher 1477 (K); Quezon, BS (Ramos & Edaño) 28586 (K); Samar, BS (Ramos) 17503 (K, P); Sorsogon, Arriola & Buera T 012 (USTH), Elmer 15511 (P). Note — Villaria multibracteata resembles V. purpurea in its lanceolate leaf blades and cymose inflorescences. However, V. multibracteata differs by its narrower leaf blades with fewer nerves and the presence of multiple triangular bracts. This feature of the bracts distinguishes the species from all other Villaria species.	en	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
7E563D16FF83FFBFD848213CFB5AF7C5.taxon	distribution	Distribution — Philippines (Bataan, Quezon, Aurora, Sorsogon, Surigao, Visayas, Zambales Prov.). Habitat & Ecology — In primary forest to deep wooded ravines, also near coastal areas. Flowering: April to June; fruiting: May to September. Additional specimens studied. THE PHILIPPINES, Alabat, Quezon, BS (Ramos & Edaño) 48148 (P); Alas-asin, Bataan, Arriola & Banag TA 002 (USTH); Angat, Bulacan, BS (Ramos & Edaño) 34067 (P); Baler, Aurora, Alejandro et al. 10 ­ 110 (USTH); Bataan, FB (Pascual) 28779 (P); Bulusan, Elmer 15925 (L); Ilocos, BS (Ramos) 33007 (K); Irosin, Elmer 16598 (K); Lamao, Bataan, Borden 2928 (P), Borden 21726 (K); Maria Aurora, Arriola & Alejandro 11 ­ 048 (USTH); Montalban, Rizal, Loher 6428 (K); Quezon, Merrill 34067 (P); Rizal, BS (Ramos) 41871 (P); Surigao, Banag & Tandang SU 024 (PNH); Visayas, Langenberger 2074 (L). Notes — 1. The lanceolate to narrowly ovate leaf shape of V. purpurea resembles that of V. acutifolia and V. glomerata. However, V. purpurea differs from all other known Villaria species by having the longest fruit stalks of all. 2. Elmer (1906) indicated Elmer 7959 to be the type. No duplicate with this number was found. In the Biodiversity Heritage Library, the number in the scan of Elmer (1906) has been changed by hand to 7616 (a number, among others, also mentioned by Merrill 1923), of which a duplicate is present in E. However, it is not clear that Elmer made a typing error. Therefore, we select a neotype here.	en	Arriola, A. H., Alejandro, G. J. D. (2020): Three new combinations in Villaria (Octotropideae, Rubiaceae), an endemic genus from the Philippines. Blumea 65 (1): 86-89, DOI: 10.3767/blumea.2020.65.01.11, URL: https://doi.org/10.3767/blumea.2020.65.01.11
