taxonID	type	description	language	source
7C03879DFF8EB346358CF8B1FD86E214.taxon	type_taxon	Type species. Sandalodesmus salvadorii (Silvestri, 1895), originally described as Odontopeltis salvadorii Silvestri, 1895; subsequently designated by Rojas-Buffet et al. 2022: 261.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8EB346358CF8B1FD86E214.taxon	diagnosis	Diagnosis. (Modified from Rojas-Buffet et al. 2022). Males of Sandalodesmus differ from those of Oncoleptodesmus by the presence of only two dorsal macrosetae on the gonocoxites (gcx) (Figs 6 C, 9 B) vs. several dorsal macrosetae of Oncoleptodesmus (Schubart 1958: figs 1 – 3). By the absence of spinulations on the acropodite process (ap) vs. the presence of numerous small spinulations along its lateral margin on Oncoleptodesmus (Schubart 1958: figs 1 – 3). An exception is Sandalodesmus repandus (Schubart, 1958), which exhibits spinulation in this region.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8EB346358CF8B1FD86E214.taxon	description	Modified after Rojas-Buffet et al. (2022).	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8DB34C358CF9C8FF6AE7A8.taxon	description	Figs 1 – 6	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8DB34C358CF9C8FF6AE7A8.taxon	diagnosis	Diagnosis. Males of Sandalodesmus peruibe sp. nov. can be distinguished from other Sandalodesmus species by the following combination of gonopodal characters: prefemoral process (pfp) with two acute terminations at the same height (mpfp; Figs 5 B, 6 B, D), and an ectal apical projection (epfp) with a small, rounded apex (Figs 5 A, 6 A, C). Acropodite region with apical portion subparallel to the gonocoxite (gcx) (Figs 5 A – B, D, 6), vs. the entirely perpendicular orientation observed in other species (Fig. 10). An acute process (asp) is present internally to the acropodite (Fig. 5 D) vs. absent in other species, except in S. fandango sp. nov. Presence of a conspicuous incision in the median portion of the acropodite region, in mesal view (Figs. 5 B, D, 6 D), similar to the condition observed in S. fandango sp. nov., S. avilectus Hoffman, 1982 and S. librellus Hoffman, 1982.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8DB34C358CF9C8FF6AE7A8.taxon	etymology	Etymology. The specific epithet peruibe refers to “ Peroibe ”, a folkloric figure of the Tupiniquim Indigenous people of Brazil, known as a strong hunter and lover of “ Jureia ”. The name alludes to the curved apical part of the gonopod, which resembles a flexed human arm, a symbolic reference to the strength of Peroibe. It is important to note that the curved apical part of the gonopod is a common trait to other Sandalodesmus, the curvature is not to be taken as a diagnostic feature of this species. Noun in apposition.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8DB34C358CF9C8FF6AE7A8.taxon	materials_examined	Material examined. Type material: Holotype: BRAZIL ● ♂; Estação Ecológica Juréia-Itatins (- 24.3571, - 47.0136), Peruíbe, São Paulo; iii. 1997; A. D. Brescovit leg.; IBSP 401. Paratypes: BRAZIL ● 2 ♂♂; same data as for holotype; IBSP 15767. Other material: BRAZIL ● 2 ♂♂ [fragmented specimens]; same locality as holotype; iv. 2012; J. P. P. Pena-Barbosa leg.; IBSP 3741.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF8DB34C358CF9C8FF6AE7A8.taxon	description	Description. Male (Holotype, IBSP 401). Coloration (preserved in 70 % ethanol): head yellowish in clypeus region and reddish on vertex; antennae yellowish; body reddish orange, slight darkening on last 3 – 4 rings; paranota tips light yellow (Fig. 2 A – C); anterior legs (body rings 2 – 5) lighter than body; posterior legs (body rings 6 – 18) reddish with yellowish postfemora (Fig. 2 D – F). Telson reddish (Fig. 2 C); coloration varying among individuals. Antennae: covered with thin setae; 5 th, 6 th and 7 th antennomeres with small clusters of basiconic sensilla (br) at distal external margin (Fig. 4 F). Gnathochilarium: uniformly covered with small setae on all plates. Body rings: cuticle smooth, flat (Fig. 4 D). Paranota well projected laterally and slightly downward, forming lowered arch (Fig. 4 A); anterior margin rounded; posterior margin subtriangular (Fig. 2 A – C). Ozopore (oz) arrangement typical of Polydesmida (5, 7, 9, 10, 12, 13, 15 – 19); surrounded by peritremata, dorsally positioned in posterior region (Fig. 4 D). Sternites: body ring 4 with a pair of flat projections (Fig. 3 A); body ring 5 with one pair of subpyramidal bumps between anterior legs (Fig. 3 D), generally symmetrical, flat anterior face, longer on transverse axis (vs. longitudinal, see Bouzan et al. 2023; fig. 3 A); body ring 6 with large V-shaped posterior-ventral depression (Fig. 3 B); body ring 7 without projections (Fig. 5 C). Gonopod aperture oval with deep V-shaped anterior margin into prozonite (Fig. 5 C); body rings 8 – 19 each with two pairs of small projections posteriorly near each coxa (Fig. 3 E). Legs (Fig. 4 B – C, E): gonopores opening as longitudinal slits, on concave area at apex of subrectangular gonapophyses (Fig. 3 C). Coxae of walking legs with one long ventral seta; Leg modification patterns (ring numbers): prefemoral knob (2 <3 – 15> 16 – 18) (Fig. 4 B); tibial sole (sensu Hoffman 1982), i. e., projection on the distomesal part of the tibia (2 <3 – 12> 13 – 14> 15 – 16; absent from 17 – 18) (Fig. 4 E); presence and degree of development of lateral projections dorsal to the stigma are as follows: 2 – 7 <8 – 13 <nearly absent from 14 – 20. Telson: without modifications (Fig. 3 F); hypoproct with a posterior marginal projection and two (1 + 1) lateral marginal setae. Gonopods (Figs 5 A – B, D, 6): Gonocoxite (gcx) approximately same in size as telopodites; broad at midsection (Fig. 6 C); coxites with transverse striations at anterior margin, in ectal view; a large and robust spiniform process (gsp) present (Fig. 5 A). Cannula (c) lacking prominent projections. Prefemoral region (pfr) short, approximately one-third telopodite length (Fig. 5 B). Prefemoral process (pfp) subspatulate, subparallel to acropodite, reaching half its length (Fig. 5 B); mesal apical projection (mpfp) bearing two acute terminations at same height (Figs 5 B, 6 B); ectal apical projection (epfp) with a small and rounded apex (Figs 5 A, 6 A). Acropodite process (ap) with conspicuous dorsal striations (Fig. 6 C); short acropodite spiniform process (asp) on inner region (Figs 5 D, 6 D). Median incision visible on acropodite in mesal view, creating fold along spermatic groove (Figs. 5 B, D, 6 D). Solenomere (s) shorter than ap (Figs. 5 D, 6 B, D). Measurements (in mm): Body: total length 52.8, width of 10 th body ring 7.8. Antennomere lengths (1 − 7): 0.4; 1.3; 1.3; 1.3; 1.2; 1.3; 0.2. Podomere lengths of 6 th leg (1 − 7): 0.3; 1.2; 2.4; 1.0; 1.2; 1.3; tarsal claw 0.3. Telson length 1.3. Gonopod aperture: length 1.9, width 2.8. Gonopod: length 2.8, width 3.4. Gonocoxite: length 1.6, width 1.4. Telopodite: length 2.3, width 1.6. Female. Unknown. Species group. Sandalodesmus peruibe sp. nov. is assigned to the iguazuensis species group based on the following shared characters: prefemoral region short, not significantly extended mesally (Fig. 5 B); prefemoral process shorter than the acropodite region and non torsate (Fig. 6 D); acropodite process broad, forming a hood-like structure (Figs 5 A−B, 6); and solenomere being small and short, not extending beyond the acropodite process (Figs 5 D, 6 D).	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF87B349358CFB4DFD21E2A4.taxon	description	Figs 1, 7 – 10	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF87B349358CFB4DFD21E2A4.taxon	diagnosis	Diagnosis. Males of Sandalodesmus jureia sp. nov. differ from other Sandalodesmus by the following combination of gonopodal characters: prefemoral process (pfp) trifid (Figs 9 A, 10 A, C) vs. bifid or unbranched in the other species. Ectal projection (epfp) twists toward the acropodite and ends in a sharp, dorsally pointing tip (Figs 9 A, 10 A, C), while the mesal projection (mpfp) twists away from the acropodite and has a rounded apex (Figs 9 B, 10 B, D). Acropodite process (ap), in ectal view, with a small laminar process (alp) with three main spines (Figs 9 A, C, 10 A, C) vs. absent. Solenomere (s) exceeds the length of the acropodite process, with its tip directed mesally (Figs 9 A, B, 10), as in the stramineus species group.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF87B349358CFB4DFD21E2A4.taxon	etymology	Etymology. The specific epithet jureia refers to a folkloric character from the indigenous Tupiniquim people of Brazil, representing a woman of remarkable beauty and the lover of “ Peroibe ”. Noun in apposition.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF87B349358CFB4DFD21E2A4.taxon	materials_examined	Material examined. Type material: Holotype: BRAZIL ● ♂; Estação Ecológica Juréia-Itatins (- 24.3571, - 47.0136), Peruíbe, São Paulo, Brazil; xii. 1998; A. D. Brescovit leg.; IBSP 1300. Paratype: BRAZIL ● ♂; Cachoeira do Guilherme, Estação Ecológica Juréia-Itatins (- 24.3571, - 47.0136), Peruíbe, São Paulo, Brazil; i. 1994; A. Eterovic leg.; IBSP 1127.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF87B349358CFB4DFD21E2A4.taxon	description	Description. Male (Holotype, IBSP 1300). Coloration (preserved in 70 % ethanol): head and antennae reddish yellow; body entirely reddish brown, some body rings slightly darkened (Fig. 7 A – C). Paranota tip slightly yellowish (Fig. 7 B). Legs and sternites yellowish (Fig. 7 E – F). Telson reddish, lighter towards tip (Fig. 7 C). Antennae: covered in thin setae; 5 th, 6 th and 7 th antennomeres with small clusters of basiconic sensilla distally on external margin. Gnathochilarium: small setae uniformly distributed on all plates (Fig. 8 C). Body rings: cuticle smooth, flat. Paranota well projected laterally, slightly downward (Fig. 7 A – C); anterior margin rounded, posterior margin subtriangular (Fig. 7 A – C). Ozopore (oz) arrangement typical for Polydesmida (5, 7, 9, 10, 12, 13, 15 – 19); surrounded by peritremata (Fig. 7 B), located dorsally in posterior region. Sternites: body ring 4 with a pair of flat projections; body ring 5 with one pair of small bumps (Fig. 8 B), between the anterior legs; generally symmetrical, reduced; body ring 6 and 7 without depressions or projections. Gonopod aperture oval, unmodified (Fig. 8 E); body rings 8 – 19 with two pairs of small projections, near each coxae. Legs (Fig. 8 A, D): gonopores opening as longitudinal slits, on concave area at apex of subrectangular gonapophyses. Coxae of walking legs with one long ventral seta. Leg modifications greatly reduced (vs. other species described herein), except for tibial sole on legs of rings 2 – 6. Leg modification patterns (ring numbers): prefemoral knob (2 – 10> 11 – 14; absent on 15 – 18); tibial sole (2 – 6> 7> 8 – 9> 10 – 11; absent on 12 – 18); presence and degree of development of lateral projections dorsal to the stigma are as follows: 2 – 7 <8 – 13 <nearly absent from 14 – 20. Telson: without modifications; hypoproct with a posterior marginal projection and two (1 + 1) lateral marginal setae (Fig. 8 F). Gonopods (Figs 9, 10): Gonocoxite (gcx) approximately same size as telopodite; relatively uniform circumference along entire length (Fig. 9 A). Anterior margin with transverse striations, in ectal view; spiniform process small (gsp) (Fig. 10 C – D). Cannula (c) without prominent projections. Prefemoral region (pfr) short, approximately one-third length of telopodite (Fig. 10 D). Prefemoral process (pfp) trifid, subspatulate, subparallel to acropodite; reaching mid-length of acropodite (Fig. 10 D). Ectal projection (epfp) twisted toward acropodite, apex sharp, dorsally directed (Figs 9 A, 10 A, C); mesal projection (mpfp) twisted away from acropodite, apex rounded (Figs 9 B, 10 B, D). Acropodite process (ap), in ectal view, with a small laminar process (alp) with three main spines and a fourth secondary spine branching from median one (Figs 9 A – B, 10 A, C); ventral margin with conspicuous longitudinal striations (Fig. 9 A, D). Acropodite region without incisions near spermatic groove. Solenomere (s) slightly longer than acropodite process; tip directed upward and toward the mesal side (Fig. 9 A – B). Measurements (in mm): Body: total length 42.2, width of 10 th body ring 6.2. Antennomere lengths (1 − 7): 0.7; 1.3; 1.1; 1.2; 1.3; 1.3; 0.2. Podomere lengths of 6 th leg (1 − 7): 0.8; 1.2; 2.2; 0.8; 0.9; 1.4; tarsal claw 0.3. Telson length 1.1. Gonopod aperture: length 0.8, width 1.7. Gonopod: length 1.5, width 1.7. Gonocoxite: length 1.1, width 0.6. Telopodite: length 1.4, width 0.5. Female. Unknown. Species group. Sandalodesmus jureia sp. nov. is assigned to the stramineus species group based on the following shared characters: solenomere longer than acropodite process and with a posterior extension (= subterminal lobe; Hoffman, 1967) on mesal side (Figs 9 A−B, 10).	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF82B34B358CFE41FF48E4C5.taxon	description	Figs 1, 11 – 12	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF82B34B358CFE41FF48E4C5.taxon	diagnosis	Diagnosis. Males of S. fandango sp. nov. differ from other Sandalodesmus by the following combination of gonopodal characters: prefemoral process (pfp) with a serrated medial apical margin (Fig. 12 D) vs. smooth in the other species, and with a small acute projection on the ectal apical margin. Mesal projection of prefemoral process (mpfp) folds posteriorly, positioning the frontal face of the pfp dorsally (Figs 12 B, D). Acropodite spiniform process (asp) present (Figs 12 B, D) vs. absent, except in S. peruibe sp. nov. Presence of an incision in the median-basal portion of the acropodite region, in mesal view (Fig. 12 B, D), similar to the condition observed in S. peruibe, S. avilectus and S. librellus.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF82B34B358CFE41FF48E4C5.taxon	etymology	Etymology. The specific epithet “ fandango ” refers to the “ Fandango Caiçara ”, a traditional cultural dance of the coastal inhabitants of the southern Atlantic Forest in São Paulo state, where the species was collected. Noun in apposition.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF82B34B358CFE41FF48E4C5.taxon	materials_examined	Material examined. Type material: Holotype: BRAZIL ● ♂; Estação Ecológica Juréia-Itatins (- 24.3571, - 47.0136), Peruíbe, São Paulo, Brazil; iii. 1997; A. D. Brescovit leg.; IBSP 320.	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
7C03879DFF82B34B358CFE41FF48E4C5.taxon	description	Description. Male (Holotype, IBSP 320). Coloration (preserved in 70 % ethanol): clypeus reddish; labrum region yellowish; vertex reddish; head, antennae, and body rings uniformly brownish red (Fig. 11 A – C). Paranota tip yellowish (Fig. 11 B). Sternites reddish (Fig. 11 E – F). Legs reddish orange, lighter than sternites (Fig. 11 D – E). Telson reddish, lighter at tip (Fig. 11 C). Antennae: thin setae; 5 th, 6 th and 7 th antennomeres with small clusters of basiconic sensilla on distal external margin. Gnathochilarium: all plates with uniform coverage of small setae. Body rings: cuticle smooth, flat. Paranota well projected laterally and slightly downward, forming lowered arch; anterior margin rounded; posterior margin subtriangular. Ozopore (oz) arrangement typical for Polydesmida (5, 7, 9, 10, 12, 13, 15 – 19); surrounded by peritremata; located dorsally in posterior region (Fig. 11 B). Sternites: body ring 4 with a pair of flat projections; body ring 5 with one pair of subpyramidal bumps; symmetrical; anterior face flat and transversely elongated; between anterior legs; body ring 6 with V-shaped posterior-ventral depression; reduced; body ring 7 without projections. Gonopod aperture extending into prozonite; anterior margin oval; body rings 8 – 19 with two pairs of small projections near each coxae (Fig. 11 E). Legs: gonopores opening as longitudinal slits, on concave area at apex of subrectangular gonapophyses. Coxae of walking legs with one long ventral seta; Leg modification patterns (ring numbers): prefemoral knob (2 <3 – 15> 16 – 18); tibial sole (2 <3 – 12> 13 – 14> 15 – 16; absent on 17 – 18); presence and degree of development of lateral projections dorsal to the stigma are as follows: 2 – 7 <8 – 13 <nearly absent from 14 – 20. Telson: without modifications (Fig. 11 F); hypoproct with a posterior marginal projection and two (1 + 1) lateral marginal setae. Gonopods (Fig. 12): Gonocoxite (gcx) approximately as long as telopodite; broad at mid-length, narrowing toward ectal margin (Fig. 12 A); Anterior margin with transverse striations, in ectal view (Fig. 12 C); spiniform process small (gsp) (Fig. 12 C). Cannula (c) without prominent projections. Prefemoral region (pfr) short, approximately one-third telopodite length (Fig. 12 D). Prefemoral process (pfp) with serrated medial apical margin (Fig. 12 D); small acute projection on ectal apical margin; mesal prefemoral process (mpfp) folded posteriorly, positioning frontal face of pfp dorsally (Fig. 12 B, D). Acropodite spiniform process (asp) present (Fig. 12 B, D). Acropodite process (ap) with longitudinal striations on ventral margin (Fig. 12 D); without apical curvature, perpendicular to gonocoxite (gcx). Median-basal incision present in mesal view; creates fold along spermatic groove path (Fig. 12 B, D). Solenomere (s) shorter than acropodite. Measurements (in mm): Body: total length 49.9, width of 10 th body ring 7.0. Antennomere lengths (1 − 7): 0.5; 1.5; 1.2; 1.2; 1.2; 1.0; 0.2. Podomere lengths of 6 th leg (1 − 7): 0.8; 1.0; 2.0; 1.0; 1.0; 1.0; 1.0; tarsal claw 0.2. Telson length 1.5. Gonopod aperture: length 1.5, width 2.6. Gonopod: length 2.6, width 2.6. Gonocoxite: length 2.0, width 1.2. Telopodite: length 2.0, width 1.0. Female. Unknown. Species group. Sandalodesmus fandango sp. nov. is assigned to the iguazuensis species group based on the following shared characters: prefemoral region short, not significantly extended mesally (Fig. 12 D); prefemoral process shorter than the acropodite region and non torsate (Fig. 12 B); acropodite process broad, forming a hood-like structure (Fig. 12); and solenomere being small and short, not extending beyond the acropodite process (Fig. 12 D).	en	Figueiredo, Thiago D., Iniesta, Luiz F. M., Brescovit, Antonio D., Bouzan, Rodrigo S. (2025): Three new species of Sandalodesmus Silvestri, 1902 from the last intact stretches of Atlantic Forest in Brazil (Polydesmida: Chelodesmidae: Sandalodesmini). Zootaxa 5723 (4): 514-532, DOI: 10.11646/zootaxa.5723.4.3, URL: https://doi.org/10.11646/zootaxa.5723.4.3
