taxonID	type	description	language	source
7C3D87FBB617902C71E4FC76C3A1FDA2.taxon	description	SUBFAMILY: CHIRONOMINAE NEWMAN, 1834 TRIBE: TANYTARSINI ZAVŘEL, 1917 SUBTRIBE: TANYTARSINA ZAVŘEL, 1917	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB617902C722BFF5DC5DFFB6B.taxon	description	(FIGS 1 A – F, 11 A, B) Material examined Adult male, inventory no. BSIP Tad- 610 (left: antenna, wing, foreleg, tarsi of mid- and hindlegs and tarsus of right foreleg damaged; Fig. 1 A, B) preserved in an 8.5 mm × 4.5 mm × 3.5 mm piece of amber (early Eocene, ~ 54 Mya, Tadkeshwar mine, Gujarat state, India; IR spectrum no. 14258 IAA; Fig. 11 A). Holotype and paratype, inventory no. BSIP Tad- 607 a and b, syninclusions (see Zakrzewska et al., 2018); IR spectrum no. 14259 IAA (Fig. 11 B).	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB611902A7004FD0CC3FAFA34.taxon	materials_examined	Type species: Tanytarsus signatus (van der Wulp, 1858). TANYTARSUS FORFEX GIŁKA & ZAKRZEWSKA SP.	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61090277187F952C35BFC57.taxon	description	(FIGS 4 A – F, 5 A – F, 11 D) LSID: urn: lsid: zoobank. org: act: D 8802738 - 369 F- 4 D 19 - 823 F- 379 C 31 FFA 80 B Type material Holotype, inventory no. BSIP Tad- 521: adult male (tarsi of left foreleg and of both mid- and hindlegs missing; Fig. 4 A, B) in a 7 mm × 5 mm × 2.5 mm piece of amber preserved in a cubicoid plastic mass (early Eocene, ~ 54 Mya, Tadkeshwar mine, Gujarat state, India; IR spectrum no. 14257 IAA, Fig. 11 D). Etymology From Latin ramus, branch or twig, refering to the frail, simply branched median volsellae. Noun in apposition. Diagnosis Wing vein Sc extending well beyond RM; FCu placed well distal to RM; distance between ends of R 1 – R 2 + 3 and R 2 + 3 – R 4 + 5 unequal, VR C ~ 5. Tibial combs and spurs stout, combs fan shaped, only one comb with spur on mid-leg, each comb with spur on hindleg. Anal tergite with three spinulae in a row, placed in pit surrounded by crest-like ridges. Anal point slender, long, reaching apex of inferior volsella. Superior volsella pipe shaped. Stem of median volsella twig shaped, branched into two simple, slender lamellae. Inferior volsella parallel sided. Description Adult male (N = 1). Total length ~ 2.1 mm; wing length ~ 840 μm. Head (Fig. 4 B): Eyes bare, slightly produced in dorsomedian position. Frontal tubercles not observed. Antenna with 13 flagellomeres, AR unmeasurable, plume fully developed. Length of palpomeres 2 – 5 (in micrometres): ~ 25, 105, 105 and 169. At least 12 strong clypeals. Thorax: Dc at least nine, Pa at least one, other thoracic setae unobservable. Wing (Fig. 4 C): Anal lobe relatively well developed, broadest at two-thirds of length, width 265 μm, length / width ratio 3.17. Sc extending well beyond RM. RM slightly oblique relative to M and R. R 1 and R 2 + 3 arcuate in distal section. FCu placed well distal to RM; VR Cu 1.35. Veins ending as follows (in order from base to tip): An, Sc, Cu 1, R 1, R + 2 3, M + 3 4, R + 4 5, M + 1 2; distance between ends of R 1 – R 2 + 3 and R 2 + 3 – R 4 + 5 unequal; VR C ~ 5. Wing membrane covered with macrotrichia in distal half. Legs: Tibia of foreleg with stout spur ~ 25 μm long. Tibial combs of mid leg fan shaped, teeth ≤ 25 μm long, only one comb bearing slender spur ~ 55 μm long (Fig. 4 D, E); combs of hindleg broadly separated, fan shaped; one comb broad, with teeth ≤ 25 μm long, bearing spur ~ 50 μm long; second comb slender, teeth ≤ 30 μm long, bearing spur ~ 60 μm long (Fig. 4 F). For the leg segment lengths and leg ratios, see Table 3. Hypopygium (Fig. 5 A – F): Gonostylus slightly longer than gonocoxite, ~ 85 μm long, tapering towards pointed apex. Anal tergite semicircular, bearing three spinulae arranged in a row, placed in rounded, longitudinal pit, its crest-like ridges tapering towards base of anal point; several setae on each side. Anal point long, reaching apex of inferior volsella, slender, parallel sided with blunt tip; severalstrongposterolateralsetaeoneachside (Fig. 5 A, B). Superior volsella pipe shaped, with oval basal part, distinctly narrowed at mid-length, distal part elongate with apex blunt; several short setae at median margin of basal part (Fig. 5 B, D). Digitus not observed. Stem of median volsella ~ 20 μm long, posteromedially directed, twig shaped, branched into two simple, slender lamellae (Fig. 5 C, E, F). Inferior volsella reaching half the length of the gonostylus, nearly parallel sided, with distal half armed, bearing numerous stout setae. Remarks Tanytarsus ramus differs from all known species of the genus by a set of distinct characters as follows: the VR C ratio is nearly five (Fig. 4 C), the anal tergite has a couple of spinulae placed in a row in a large pit, and the anal point is slender and extraordinarily long (Fig. 5 A, B); the pipe shaped superior volsella should be considered a key feature (Fig. 5 B, D), whereas the digitus, typical of Tanytarsus, is not observed (probably absent). The most peculiar structure diagnostic of T. ramus is the hypopygial median volsella (Fig. 5 E, F) of a shape perceived as a plesiomorphy (see Discussion: Evolution of the diagnostic structures).	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61C90277276FB11C492FC13.taxon	materials_examined	Type species: Stempellina bausei (Kieffer, 1911). STEMPELLINA STEBNERI GIŁKA & ZAKRZEWSKA SP.	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61C90277075FAE0C421FD74.taxon	description	(FIG. 6 A – F) Material examined Ta n y t a r s u s s p. 1, o n e a d u l t m a l e i n a n 8.5 mm × 6 mm × 3.5 mm piece of amber (inventory no. BSIP Tad- 533; Fig. 6 A, B); Tanytarsus sp. 2, one adult male in a 6.5 mm × 5 mm × 4 mm piece of amber (inventory no. BSIP Tad- 883 a; Fig. 6 C, D), syninclusion: Coleoptera (inventory no. BSIP Tad- 883 b); Tanytarsus sp. 3, one adult male in a 6 mm × 3 mm × 1 mm piece of amber preserved in a cubicoid plastic mass (inventory no. BSIP Tad- 884; Fig. 6 E, F); Early Eocene, ~ 54 Mya, Tadkeshwar mine, Gujarat state, India. Remarks The above three specimens are ascribed to the genus Tanytarsus on the basis of the head, wing and hypopygium characters (cf. Stebner et al., 2017). However, most of the diagnostic structures are weakly observable, deformed or insufficiently preserved to define the individuals definitively to species level. Nevertheless, these males differ distinctly from each other and from those designated as new species described in this paper, thus they most probably belong to another species. Therefore, with presumably five species, Tanytarsus is the most species-rich genus among the Cambay Tanytarsini. The specific diversity proportions within the tribe may thus be similar relative to the extant fauna: five Tanytarsus / eight Tanytarsini in Cambay amber vs. ~ 360 Tanytarsus / ~ 700 Tanytarsini extant species (Giłka W, Zakrzewska M, unpublished data). SUBTRIBE: ZAVRELIINA SAETHER, 1977	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB61890237010FBEAC3EDFC29.taxon	materials_examined	Type species: Stempellinella saltuum (Goetghebuer, 1921). STEMPELLINELLA POLLEX GIŁKA & ZAKRZEWSKA SP.	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
7C3D87FBB6069039723CF880C389F9FF.taxon	description	& Zakrzewska, 2013 described from Baltic amber (cf. Giłka et al., 2013; Zakrzewska & Giłka, 2014).	en	Zakrzewska, Marta, Singh, Hukam, Wagner-Wysiecka, Ewa, Giłka, Wojciech (2020): Minute and diverse in fossil sticky stuff: Tanytarsini (Diptera: Chironomidae) from early Eocene Indian Cambay amber. Zoological Journal of the Linnean Society 189: 1398-1425
