identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
9D2265BEB9625278AEBD6D408EE554CE.text	9D2265BEB9625278AEBD6D408EE554CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces crenatus Haqnawaz, Niazi & Khalid 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Candolleomyces crenatus Haqnawaz, Niazi &amp; Khalid sp. nov.</p>
            <p>Figs 3, 4</p>
            <p>Etymology.</p>
            <p> Species name “  crenatus ” (Latin) refers to the crenate cap margins. </p>
            <p>Holotype.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu District,  
                <a title="Search Plazi for locations around (long 70.82361/lat 30.4675)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.82361&amp;materialsCitation.latitude=30.4675">Indus Riverbed</a>
                 , (30°28'03"N, 70°49'25"E, 126 m a. s. l.), on loamy soil, rich in organic matter, under  Vachellia nilotica , 07 Aug. 2022, Muhammad Haqnawaz, Q – 13 (Holotype LAH 38257). GenBank: PQ 329548 [ITS], PQ 329553 [LSU], PQ 369432 [tef- 1 α]. 
            </p>
            <p>Diagnosis.</p>
            <p> This species is different from  Candolleomyces badiophyllus by its convex to plane, crenate cap margins, wrinkled, dull surface, light gray, with dull orange center of pileus, marginate base of stipe, and small basidiospores (7–9 × 4–5 µm). </p>
            <p>Description.</p>
            <p>Pileus 14–34 mm diameter, convex to plane, straight shape, and crenate cap margins; presence of veil elements; wrinkled, dull surface; light gray (7.5 YR 8 / 1) with dull orange (7.5 YR 7 / 3) center. Lamellae adnexed, ventricose, average, close, fimbriate, unequal, 1–5 tiers, dull reddish brown (5 YR 5 / 3). Stipe 23–35 × 3–5 mm, compressed, flexuous, tomentose, marginate base. Annulus absent. Taste and odor are not recorded.</p>
            <p>Basidiospores [150 / 3 / 2], (7 –) 7.5–8.5 (– 9) × (4 –) 4.5–5 (– 5.7) µm, Q = 1.4–1.8, Qav = 1.61, ellipsoid, thick-walled, smooth, rarely guttulate, central germpore present, reddish brown in water. Basidia (15 –) 16–23 (– 24) × (7 –) 7.5–11 (– 12) µm, clavate to broadly clavate, hyaline to yellowish in water, thick-walled, smooth, with 3–4 sterigmata. Cheilocystidia (15 –) 16–31 (– 32) × (7 –) 8–12 (– 13) µm, ellipsoid to broadly ellipsoid, clavate to broadly clavate, globose, utriform to utriform with median constriction, cylindrical, hyaline to yellowish in water, thick-walled. Pleurocystidia absent. Pileipellis an irregular epithelium, thick-walled, 18–30 × 16–24 µm, globose to subglobose and ellipsoid to broadly ellipsoid, clavate to broadly clavate cells, hyaline to light yellowish in water. Stipitipellis a cutis, hyphae subregular, branched 6–17 µm in diameter, thin-walled, septate, and hyaline to yellow in water. Caulocystidia (17 –) 18–42 (– 43) × (8 –) 9–14 (– 15) µm, utriform, clavate to broadly clavate, globose, broadly ellipsoid, hyaline in water. Clamp connections present in all tissues.</p>
            <p>Ecology and habitat.</p>
            <p> Terrestrial, caespitose, on loamy soil rich in organic matter, under  Vachellia nilotica . </p>
            <p>Additional material examined.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu,  
                <a title="Search Plazi for locations around (long 70.87639/lat 30.41861)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.87639&amp;materialsCitation.latitude=30.41861">Pirhar Gherbi</a>
                 , (30°25'07"N, 70°52'35"E, 136 m a. s. l.), on loamy soil, 11 Aug. 2023, Muhammad Haqnawaz, HA – 03 (Paratype LAH 38258). GenBank: PQ 329547 [ITS], PQ 329554 [LSU], PQ 369433 [tef- 1 α]  . 
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	https://treatment.plazi.org/id/9D2265BEB9625278AEBD6D408EE554CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Qirui;Haqnawaz, Muhammad;Niazi, Abdul Rehman;Khalid, Abdul Nasir	Li, Qirui, Haqnawaz, Muhammad, Niazi, Abdul Rehman, Khalid, Abdul Nasir (2025): Phylogenetic studies on the genus Candolleomyces (Psathyrellaceae, Basidiomycota) occurring in the bed of the Indus River, Punjab, Pakistan, reveal three new species. MycoKeys 112: 165-182, DOI: 10.3897/mycokeys.112.141766
B767CB0CFDDE55099FF6F7DC8D43C4B9.text	B767CB0CFDDE55099FF6F7DC8D43C4B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces undulatus Haqnawaz, Niazi & Khalid 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Candolleomyces undulatus Haqnawaz, Niazi &amp; Khalid sp. nov.</p>
            <p>Figs 5, 6</p>
            <p>Etymology.</p>
            <p> Species name “  undulatus ” (Latin) refers to the wavy cap margins. </p>
            <p>Holotype.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu District,  
                <a title="Search Plazi for locations around (long 70.87/lat 30.425833)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.87&amp;materialsCitation.latitude=30.425833">Indus Riverbed</a>
                 , (30°25'33"N, 70°52'12"E, 130 m a. s. l.), on loamy soil, rich in organic matter, under  Phoenix dactylifera , 28 Aug. 2020, Muhammad Haqnawaz, KA – 39 (Holotype LAH 38255). GenBank: PQ 329546 [ITS], PQ 329551 [LSU], PQ 369428 [tef- 1 α]. 
            </p>
            <p>Diagnosis.</p>
            <p>This species is different from its closest species by parabolic to campanulate, wavy margins, light purplish gray with light brownish gray center of pileus, appendiculate, pendant annulus, oblong to ellipsoid, globose, ovoid, cheilocystidia, and caulocystidia.</p>
            <p>Description.</p>
            <p>Pileus 10–27 mm diameter, parabolic to campanulate, straight shape and wavy cap of margins, presence of veil elements, light purplish gray (5 P 7 / 1), and light brownish gray (5 YR 7 / 2) center. Lamellae adnexed to adnate, narrow, average, close, eroded, unequal, 1–3 tiers, light reddish gray (7.5 R 7 / 1), Stipe 15–46 × 2–4 mm, concolourous to pileus, punctate, flexuous, tomentose. Annulus appendiculate and pendant. Taste and odor not recorded.</p>
            <p>Basidiospores [150 / 3 / 2], (6 –) 6.5–7.5 (– 8) × (3.5 –) 4–5 (– 6.6) µm, Q = 1.15–1.94, Qav = 1.58, ellipsoid to oblong, thick-walled, smooth, guttulate, germpore present, brownish with yellowish guttule in water. Basidia (15 –) 16–24 (– 25) × (7 –) 7.5–11 (– 12) µm, clavate to broadly clavate, rarely cylindrical, spherical with broad peduncle, hyaline in water, thick-walled, smooth, with 2–4 sterigmata. Cheilocystidia (15 –) 16–31 (– 32) × (7 –) 8–12 (– 13) µm, oblong to ellipsoid, globose, ovoid, utriform, hyaline in water, thick-walled. Pleurocystidia absent. Pileipellis an irregular epithelium, thick-walled, 18–30 × 16–24 µm diameter, globose to subglobose and ellipsoid to broadly ellipsoid, hyaline in water. Stipitipellis a cutis, hyphae subregular, branched 7–18 µm in diameter, thin-walled, septate, hyaline in water. Caulocystidia 17 –) 18–42 (– 43) × (8 –) 9–14 (– 15) µm, utriform, clavate to broadly clavate, ovoid, spheropedunculate, globose, hyaline in water. Clamp connection present in all tissues.</p>
            <p>Ecology and habitat.</p>
            <p> Terrestrial, connate, on loamy soil rich in organic matter, under  Phoenix dactylifera . </p>
            <p>Additional material examined.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu,  
                <a title="Search Plazi for locations around (long 70.87778/lat 30.460835)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.87778&amp;materialsCitation.latitude=30.460835">Noor Shah Thal</a>
                 , (30°27'39"N, 70°52'40"E, 129 m a. s. l.), on loamy soil, 11 Aug 2021, Muhammad Haqnawaz, SSC – 44 (Paratype LAH 38256). GenBank: PQ 329545 [ITS], PQ 329552 [LSU], PQ 369429 [tef- 1 α]  . 
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	https://treatment.plazi.org/id/B767CB0CFDDE55099FF6F7DC8D43C4B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Qirui;Haqnawaz, Muhammad;Niazi, Abdul Rehman;Khalid, Abdul Nasir	Li, Qirui, Haqnawaz, Muhammad, Niazi, Abdul Rehman, Khalid, Abdul Nasir (2025): Phylogenetic studies on the genus Candolleomyces (Psathyrellaceae, Basidiomycota) occurring in the bed of the Indus River, Punjab, Pakistan, reveal three new species. MycoKeys 112: 165-182, DOI: 10.3897/mycokeys.112.141766
25AC64DF00CF5404A087DD1248A654FD.text	25AC64DF00CF5404A087DD1248A654FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Candolleomyces virgatus Haqnawaz, Niazi & Khalid 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Candolleomyces virgatus Haqnawaz, Niazi &amp; Khalid sp. nov.</p>
            <p>Figs 7, 8</p>
            <p>Etymology.</p>
            <p> Species name “  virgatus ” (Latin) refers to the virgate pileus surface. </p>
            <p>Holotype.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu District,  
                <a title="Search Plazi for locations around (long 70.85111/lat 30.245556)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=70.85111&amp;materialsCitation.latitude=30.245556">Indus Riverbed</a>
                 , (30°14'44"N, 70°51'04"E, 134 m a. s. l.), on soil, rich in organic matter, 29 Aug 2020, Muhammad Haqnawaz, EF – 17 (Holotype LAH 38259). GenBank: PQ 329543 [ITS], PQ 329549 [LSU], PQ 369430 [tef- 1 α]. 
            </p>
            <p>Diagnosis.</p>
            <p> This species is different from  Candolleomyces campanulatus by its parabolic to plane, distinct umbo, virgate surface of pileus, crinkled, 1–7 tiers of forking lamellae, marginate base, longitudinally striation on surface of stipe, lageniform with moniliform at top, moniliform, cylindrical, tibiiform, conical cheilocystidia. </p>
            <p>Description.</p>
            <p>Basidiomata medium-sized. Pileus 17–30 mm diameter, parabolic at young stage, campanulate to plane, umbonatus, straight shape and regular cap of margins, virgate, light brownish gray (7.5 YR 7 / 1), and dull reddish brown (5 YR 5 / 4) center. Lamellae adnate, broad, crinkled, 1–7 tiers of lamellae, branched or forking; some are not branched, grayish red (10 R 6 / 2). Stipe 40–80 × 1–4 mm, white gray (N 8 / 0), slightly flexuous, hollow, tomentose, and broad at base, marginate, longitudinally striation. Annulus absent.</p>
            <p>Basidiospores [150 / 2 / 3], (6 –) 7–9 (– 10) × (4 –) 4.5–5 (– 6) µm, Q = 1.2–1.8, Qav = 1.61, ellipsoid to broadly ellipsoid, thick-walled, smooth, guttulate, germ pore present, purplish to brownish with greenish to yellow guttule in 5 % KOH and water. Basidia (12 –) 13–29 (– 30) × (6 –) 7–9 (– 10) µm, clavate, broadly clavate, hyaline to light yellowish in 5 % KOH, thick-walled, smooth, with 2–4 sterigmata. Cheilocystidia (26 –) 27–46 (– 47) × (8 –) 9–13 (– 14) µm, utriform, lageniform with moniliform at top, moniliform, cylindrical, tibiiform, conical, hyaline to light yellowish in 5 % KOH. Pleurocystidia and Caulocystidia absent. Pileipellis an irregular epithelium; pileipellis cells 30–60 × 20–42 µm diameter, globose to subglobose and ellipsoid to broadly ellipsoid, clavate, hyaline to light yellowish in 5 % KOH. Stipitipellis a cutis, hyphae subregular, branched 6–15 µm in diameter, thin-walled, septate, hyaline to light yellowish in 5 % KOH. Clamp connection present in all tissues.</p>
            <p>Ecology and habitat.</p>
            <p>Terrestrial, gregarious, scattered, on loamy soil rich in organic matter.</p>
            <p>Additional material examined.</p>
            <p>
                  Pakistan • Punjab Province: Kot Addu,  
                <a title="Search Plazi for locations around (long 71.03472/lat 30.463057)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=71.03472&amp;materialsCitation.latitude=30.463057">Noor Shah Thal</a>
                 , (30°27'47"N, 71°02'05"E, 132 m a. s. l.), on loamy soil, 02 September 2020, Muhammad Haqnawaz, KA – 59 (Paratype; LAH 38260) GenBank PQ 329544 [ITS], PQ 329550 [LSU], PQ 369431 [tef- 1 α]  . 
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	https://treatment.plazi.org/id/25AC64DF00CF5404A087DD1248A654FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Li, Qirui;Haqnawaz, Muhammad;Niazi, Abdul Rehman;Khalid, Abdul Nasir	Li, Qirui, Haqnawaz, Muhammad, Niazi, Abdul Rehman, Khalid, Abdul Nasir (2025): Phylogenetic studies on the genus Candolleomyces (Psathyrellaceae, Basidiomycota) occurring in the bed of the Indus River, Punjab, Pakistan, reveal three new species. MycoKeys 112: 165-182, DOI: 10.3897/mycokeys.112.141766
