taxonID	type	description	language	source
9605E23F9270FFCE96D8F90BFBF2FE51.taxon	materials_examined	Type species. Iphiculus spongiosus Adams & White, 1849, by monotypy.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9270FFCE96D8F90BFBF2FE51.taxon	discussion	Remarks. A handful of species are currently included in this genus. Its identification in the fossil record (Morris & Collins 1991; Collins et al. 2003) is largely based on the carapace outline, configuration of spines on the lateral margins and carapace surface ornamentation. In this respect it should be noted that some species of Typilobus Stoliczka, 1871, a leucosiid genus known exclusively from the fossil record, resemble extant representatives of Iphiculus. Artal & Hyžný (2016) presented an appraisal of the fossil leucosiid genus Typilobus. They pointed out the heterogeneity of the taxon, as previously emphasized by Vía Boada (1969), Müller (1993), and Feldmann et al. (2011) and reassigned Typilobus boscoi Vía Boada, 1959 to a new genus and family. More re-assignments of species now classified within the broadly defined Typilobus are likely once the genus is reviewed. It is possible that some of them will fall within the range of characters which now define Iphiculus. Current assignment of a newly described species to Iphiculus is based on striking morphological similarities with extant I. convexus (Fig. 2), especially on the general shape of carapace, short anterolateral spines and large rounded tubercles covering evenly the dorsal carapace surface. Since, the new fossil material consists only of isolated carapaces, comparison with sternum, pleon and chelipeds is not possible.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	diagnosis	Diagnosis. Carapace transversely subovate in outline, widest at level of posteriormost anterolateral spine; lateral margins bearing 6 short triangular spines; dorsal carapace surface covered evenly with large rounded tubercles: hepatic region with tubercle, gastric region with 3 pairs of tubercles, branchial region with 3 tubercles.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	etymology	Etymology. The species name is dedicated to Eliáš, son of the first author.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	materials_examined	Material examined. Holotype: near-complete carapace UMJGP Inv. No. 75.612, max. length, 15.7 mm; max. width, 19.0 mm (Figs 3 A – D). Paratype: near-complete carapace UMJGP Inv. No. 75.613, max. length, 13.0 mm (preserved portion); max. width, 17.2 mm; Fig. 3 E). Additional specimen: carapace fragment UMJGP Inv. No. 211339; Fig. 4 A – C).	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	description	Description. Small carapace; transversely subovate in outline; slightly wider than long, widest at midlength, at level of posteriormost anterolateral spine; dorsal surface moderately convex in both directions. Front not projected, slightly raised, narrow, not well preserved, presumably bilobed. Orbits small, concave, anteriorly directed. Lateral margins bearing 6 short triangular spines, anterolateral margin with 4 spines, posterolateral margin with 2 spines; corners between posterolateral, posterior margins pointed; posterior margin straight, narrow. Dorsal surface of carapace evenly covered with many large, densely packed granules, nearly identical in size (if cuticular surface preserved) or with round concave pustules (if cuticular surface not preserved). Carapace surface covered evenly with large rounded tubercles: hepatic region with 1 tubercle, gastric region with 3 pairs of tubercles, branchial region with 3 tubercles. Carapace grooves absent in anterior carapace portion, well developed in posterior carapace portion. Gastric region large, indistinctly demarcated with grooves. Cardiac region ovate in outline, strongly arched. Branchial regions broad. Intestinal region narrow without large spines. Sternum, pleon, pereiopods unknown. Occurrence. The species is known only from its type locality at Wetzelsdorf (Austria).	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	discussion	Remarks. Iphiculus eliasi sp. nov. is closest to the extant I. convexus in terms of the morphology of the dorsal carapace. The latter species, however, has a relatively wider carapace closer to the anterior half and possesses more large round tubercles on the dorsal carapace, especially on the hepatic and gastric regions (Fig. 2 A versus Fig. 2 B). Another extant species, I. spongiosus, differs from I. eliasi sp. nov. by the presence of large anterolateral spines and the possession of dorsal carapace tubercles which are more projected outwards (Ng et al. 2008: fig. 79). Similarly, all fossil species of Iphiculus so far known, i. e. I. granulatus Morris & Collins, 1991, I. miriensis Morris & Collins, 1991, and I. sexspinosus Morris & Collins, 1991, have well-developed anterolateral spines (Morris & Collins 1991: figs 15, 14 and 17, respectively) which are absent in I. eliasi sp. nov.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
9605E23F9271FFC896D8F9A5FC13FAC3.taxon	description	Vía (1941) reported an “ Iliinae, Ebaliinae? ” from the Middle Miocene of Catalonia, Spain. His figure (Vía 1941: pl. 10, fig. 75) clearly represents a leucosioid crab with striking similarities to Iphiculus eliasi sp. nov. Müller (1993: figs 5 M – N) refigured the specimen in greater detail (although still insufficient for closer comparison) and assigned it questionably to the leucosiid genus Randallia Stimpson, 1857 sensu lato (see Galil 2003). Re-examination of the specimen is needed to resolve its affinities.	en	Hyžný, Matúš, Gross, Martin (2016): A new iphiculid crab (Crustacea, Brachyura, Leucosioidea) from the Middle Miocene of Austria, with notes on palaeobiogeography of Iphiculus. Zootaxa 4179 (2): 263-270, DOI: 10.11646/zootaxa.4179.2.6
