taxonID	type	description	language	source
9B5FBC4801439E63FC6217F5FD3E1F37.taxon	description	COMPOSITION. Iotacypha Shcherbakov, Bashkuev et Shmakov, 2024; Burmacypha Zherikhin, 2000; Retiptera Cumming, Engel, Lian et Ulitzka, 2024; all from mid-Cretaceous Burmese amber.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801439E63FC6217F5FD3E1F37.taxon	discussion	REMARKS. Tillyard [1935] assigned Lophioneuridae Tillyard, 1921 (= Cyphoneuridae Carpenter, 1932) along with Zoropsocidae to Psocoptera. Becker-Migdisova [1961] united these two families. Vishniakova [1981] retained Lophioneurinae and Zoropsocinae as subfamilies of Lophioneuridae, but our data confirm that there is no reason to keep this distinction even at the subfamily level [Shcherbakov et al., 2024]. The three currently known iotacyphine genera have the same very distinctive habitus, but differ strikingly in their forewing venation. How to cite this article: Shcherbakov D. E., Shmakov A. S. 2024. Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber // Russian Entomol. J. Vol. 33. No. 4. P. 415 – 418. doi: 10.15298 / rusentj. 33.4.01	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801439E63FC6217F5FD3E1F37.taxon	materials_examined	TYPE SPECIES. Iotacypha zherikhini Shcherbakov, Bashkuev et Shmakov, 2024 REVISED DIAGNOSIS. Minute subbrachypterous lophioneurids. Forewings weakly elongated, strongly sclerotized, deeply convex; veins carinate, beset with strong curved setae; membrane finely areolate; Sc absent; RA long sigmoidal, leaving short R + M stem before separation of M (short RP + M stem developed); arculus (basal m-cu crossvein) between R + M and CuA; veins at wing apex Y-shaped (apical cell petiolate); discal cell bordered by 4 cells; clavus short and broad. Hind wing about half as wide as forewing, with long R + M + CuA stem, R and M separating distally; R, M and CuA subtransverse; anal area areolate. Antennae 7 - segmented, at most 1 / 2 as long as forewing, scape and pedicel stout, flagellum moderately slen- der, with simple seta-like sense cones. Ocelli far apart, lateral ocelli close to eyes, median ocellus near clypeus. Pronotum short, saddle-shaped. Hind coxae enlarged. Tibiae with rows of setae. Tarsi 2 - segmented; two slender curved claws. Genitalia very small.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801439E63FC6217F5FD3E1F37.taxon	description	COMPOSITION. Type species and Iotacypha vishniakovae sp. n.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801439E63FC6217F5FD3E1F37.taxon	discussion	REMARKS. Similar to Burmacypha Zherikhin, 2000 and Retiptera Cumming, Engel, Lian et Ulitzka, 2024 in convex areolate forewings and very simple hindwing venation, but in these two genera the veins at the forewing apex are H-shaped (the apical cell is not petiolate), the discal cell is bordered by 7 cells, and the antennae are at least 2 / 3 forewing length with a filiform flagellum. Additionally, Retiptera differs from Iotacypha in the forewing discal cell constricted about midlength and the hind wing 1 / 6 as wide as the forewing, and Burmacypha differs from Iotacypha in the forewing RA short transverse (rather than long sigmoidal) and the clavus much reduced.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801419E63FF48173EFC571EF0.taxon	description	Figs 1 – 9.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801419E63FF48173EFC571EF0.taxon	materials_examined	MATERIAL. Holotype PIN 5608 / 174, female — Burmese amber, Hukawng Valley, Kachin State, Myanmar; mid-Cretaceous (Albian – Cenomanian).	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801419E63FF48173EFC571EF0.taxon	description	DESCRIPTION. Very small, robust lophioneurids; female body length 0.65 mm (as preserved). Subbrachypterous. Forewings 0.7 mm long, rather broad (1.9: 1), widest beyond 2 / 3 wing length, broadly rounded apically, steeply tectiform in repose (their apical margins held vertically and meeting along midline), deeply convex, strongly sclerotized (especially proximally); wing base pale; membrane translucent, covered with stub-like microtrichia and fine dark hexagonal areolation, areoles increase and somewhat fade towards apex; veins dark, raised, beset with strong curved setae (including distal C). Costal margin sinuate about midlength (at least in anterior view); costal vein submarginal, with well-developed areolate precostal carina reaching RP 2. Sc absent. RA strongly sigmoidal, almost as long as wing width, distant from C distally, leaving short R + M stem about its midlength (short RP + M stem developed). Arculus (basal m-cu crossvein) between R + M and CuA closing broad basal cell. RP and M both forked before 2 / 3 wing length, RP 1 and M 2 transverse, oblique RP 2 and M 1 fused for a short distance, forming petiole of apical cell enclosing wing apex. RP 1 apex not far from RA; discal cell longer than each of two preapical cells; apical and anterior preapical cells transverse, posterior preapical cell elongated. Clavus short and broad, with two anal veins. Hind wings narrow, broadly rounded at apex; veins dark; R + M + CuA stem dividing distal to 1 / 2 wing length, R and M separating distal to 1 / 3 wing length; R, M and CuA subtransverse; claval furrow distinct; clavus with areolation near base. Antennae 7 - segmented, about 1 / 2 as long as forewing; scape and pedicel stout; flagellum moderately slender; first and last flagellomeres shortest, terminal flagellomere with two very long curved sense cones (much surpassing its apex) and three unequal apical setae. Eyes protruding, rounded; ommatidia few in number, loosely arranged. Cranium rounded; lateral ocelli near eyes; median ocellus near base of clypeus. Mouth cone wide, directed ventrally and slightly anteriorly; maxillary palps moderately long, stout, probably 3 - segmented, 2 nd segment long and massive, 3 rd segment minute; labial palps short, unsegmented. Pronotum transverse, saddle-shaped; mesoscutum subtriangular. Legs rather short, hind tibia no longer than 1 / 4 forewing length. Hind coxae largest. Tibiae with rows of setae. Tarsi 2 - segmented, basitarsus about 1 / 2 distitarsus length; right fore tarsus deformed or malformed, longer than any other tarsus and appearing 3 - segmented; two slender curved claws with sub-basal tooth. Female abdomen as long as head + thorax, widest and more sclerotized proximally. Last two sternites with small rounded median projection on posterior margin. Ovipositor rudimentary, concealed by last sternite.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
9B5FBC4801419E63FF48173EFC571EF0.taxon	etymology	ETYMOLOGY. After paleoentomologist Valentina N. Vishniakova for her great contribution to our knowledge of lophioneurids.	en	Shcherbakov, Dmitry E., Shmakov, Alexei S. (2024): Another new species of Iotacyphinae (Lophioneuridae, Thysanoptera s. l.) from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (4): 415-418, DOI: 10.15298/rusentj.33.4.01, URL: http://zoobank.org/5790d6b8-416b-4ee6-9da1-d789cf7869cc
