taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
98575F6FFFA4FFA9FF39FF684872EA82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823209/files/figure.png	https://doi.org/10.5281/zenodo.15823209	FIGURE 1. General overview of the internal structures and morphology of Lagenopolycystis, based on L. peresi. A. Photograph of live specimen. B. Micrograph of the anterior end of the same specimen, showing the proboscis in detail. C. Free-hand drawing of the same specimen. Scale not available forA–C. D. Schematic reconstruction of the female atrial organs. E. Detail of the sclerotised tube linked to the seminal receptacle. F. Micrograph of the female atrial organs in a live specimen. G. Micrograph of the male atrial organs in a live specimen. H. Drawing of the sclerotised prostate stylet with the attached accessory stylet, with blue arrows indicating measurement directions. I. Photograph of the prostate stylet and accessory stylet in a live specimen.	FIGURE 1. General overview of the internal structures and morphology of Lagenopolycystis, based on L. peresi. A. Photograph of live specimen. B. Micrograph of the anterior end of the same specimen, showing the proboscis in detail. C. Free-hand drawing of the same specimen. Scale not available forA–C. D. Schematic reconstruction of the female atrial organs. E. Detail of the sclerotised tube linked to the seminal receptacle. F. Micrograph of the female atrial organs in a live specimen. G. Micrograph of the male atrial organs in a live specimen. H. Drawing of the sclerotised prostate stylet with the attached accessory stylet, with blue arrows indicating measurement directions. I. Photograph of the prostate stylet and accessory stylet in a live specimen.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA4FFA9FF39FF684872EA82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823223/files/figure.png	https://doi.org/10.5281/zenodo.15823223	FIGURE 7. Lagenopolycystis mutabilis sp. nov. A–D. Camera lucida drawings of the stylets from A. the holotype (FMNH http:// id.luomus.fi/KV.547), and B–D. reference specimens (same scale) (B: HU XXIII.4.29; C: HU XXIII.4.30; D: HU XXIII.4.35). E–F. Micrographs of the stylets from E. the holotype, and F. a reference specimen (HU XXIII.4.29).	FIGURE 7. Lagenopolycystis mutabilis sp. nov. A–D. Camera lucida drawings of the stylets from A. the holotype (FMNH http:// id.luomus.fi/KV.547), and B–D. reference specimens (same scale) (B: HU XXIII.4.29; C: HU XXIII.4.30; D: HU XXIII.4.35). E–F. Micrographs of the stylets from E. the holotype, and F. a reference specimen (HU XXIII.4.29).	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA4FFA9FF39FF684872EA82.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823221/files/figure.png	https://doi.org/10.5281/zenodo.15823221	FIGURE 6. Lagenopolycystis conglobata sp. nov. A–F. Camera lucida drawings of the stylets from A. the designated holotype (FMNH http://id.luomus.fi/KV.546), and B–F. reference specimens (B: HU XXIII.3.46, C:HU XXIII.4.13, D: HU XXIII.3.47; E: HU XXIII.4.16, F: HU XXIII.4.01; same scale). For A–B., the sclerotised tube of the seminal receptacle is also drawn. G–I Micrographs of the stylet in fixed specimens, including G. the holotype (H: HU XXIII.3.44; I: HU XXIII.3.45). J–L. Micrographs of a live specimen from Sardinia (now HU XXIII.4.09), including J., an overview of the entire specimen, K. the proboscis, L. the stylet and seminal vesicle.	FIGURE 6. Lagenopolycystis conglobata sp. nov. A–F. Camera lucida drawings of the stylets from A. the designated holotype (FMNH http://id.luomus.fi/KV.546), and B–F. reference specimens (B: HU XXIII.3.46, C:HU XXIII.4.13, D: HU XXIII.3.47; E: HU XXIII.4.16, F: HU XXIII.4.01; same scale). For A–B., the sclerotised tube of the seminal receptacle is also drawn. G–I Micrographs of the stylet in fixed specimens, including G. the holotype (H: HU XXIII.3.44; I: HU XXIII.3.45). J–L. Micrographs of a live specimen from Sardinia (now HU XXIII.4.09), including J., an overview of the entire specimen, K. the proboscis, L. the stylet and seminal vesicle.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA2FFABFF39FEB54C50EB8E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823213/files/figure.png	https://doi.org/10.5281/zenodo.15823213	FIGURE 2. Lagenopolycystis peresi A–C. Camera lucida drawings of the stylets of reference material (A, C: SMNH-54926, 2 specimens; B. HU XXIII.2.37), with A. also including a depiction of the sclerotised tube adjacent to the entrance of the ovo-vitelloduct, highlighting the strong sphincter at its base. D–E. Drawings of stylets from newly collected material from the Mediterranean (D: HU XXIII.2.30, E: HU XXIII.2.29, F: specimen lost, G: HU XXIII.2.26). All stylets are drawn to the same scale. H. Micrograph of the stylet of a reference specimen (SMNH-54927). I–L. Micrographs of a live specimen from Sardinia (now HU XXIII.2.31), including I. an overview of the entire specimen, J. the proboscis, K. the pharynx, and L. the stylet.	FIGURE 2. Lagenopolycystis peresi A–C. Camera lucida drawings of the stylets of reference material (A, C: SMNH-54926, 2 specimens; B. HU XXIII.2.37), with A. also including a depiction of the sclerotised tube adjacent to the entrance of the ovo-vitelloduct, highlighting the strong sphincter at its base. D–E. Drawings of stylets from newly collected material from the Mediterranean (D: HU XXIII.2.30, E: HU XXIII.2.29, F: specimen lost, G: HU XXIII.2.26). All stylets are drawn to the same scale. H. Micrograph of the stylet of a reference specimen (SMNH-54927). I–L. Micrographs of a live specimen from Sardinia (now HU XXIII.2.31), including I. an overview of the entire specimen, J. the proboscis, K. the pharynx, and L. the stylet.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA0FFAAFF39FDB94E79EEBE.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823215/files/figure.png	https://doi.org/10.5281/zenodo.15823215	FIGURE 3. Lagenopolycystis articulata sp. nov. A. Micrograph of a live specimen from Sardinia. B–F. Micrograph and camera lucida drawings of the stylets, including B–C. the holotype (FMNH http://id.luomus.fi/KV.543), and D–F. reference specimens (D: HU XXIII.2.50; E: XXIII.2.48, XXIII.2.49). All drawings same scale. B–D. also include a drawing of the sclerotised duct adjacent to the seminal receptacle.	FIGURE 3. Lagenopolycystis articulata sp. nov. A. Micrograph of a live specimen from Sardinia. B–F. Micrograph and camera lucida drawings of the stylets, including B–C. the holotype (FMNH http://id.luomus.fi/KV.543), and D–F. reference specimens (D: HU XXIII.2.50; E: XXIII.2.48, XXIII.2.49). All drawings same scale. B–D. also include a drawing of the sclerotised duct adjacent to the seminal receptacle.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA1FFA4FF39FAE94C3AE81A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823217/files/figure.png	https://doi.org/10.5281/zenodo.15823217	FIGURE 4. Lagenopolycystis azorensis sp. nov. A–C. Micrographs and camera lucida drawings of the stylets A–B. the holotype (FMNH http://id.luomus.fi/KV.544) and C. a reference specimen (HU XXIII.3.13; same scale). D–G. Micrographs of a live specimen from Sardinia (HU XXIII.3.16), including D. an overview of the entire specimen, E. the pharynx, F. the stylet, and G. the proboscis.	FIGURE 4. Lagenopolycystis azorensis sp. nov. A–C. Micrographs and camera lucida drawings of the stylets A–B. the holotype (FMNH http://id.luomus.fi/KV.544) and C. a reference specimen (HU XXIII.3.13; same scale). D–G. Micrographs of a live specimen from Sardinia (HU XXIII.3.16), including D. an overview of the entire specimen, E. the pharynx, F. the stylet, and G. the proboscis.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFAFFFA4FF39FD4D4873EDF4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823219/files/figure.png	https://doi.org/10.5281/zenodo.15823219	FIGURE 5. Lagenopolycystis canariensis sp. nov. A–D. Camera lucida drawings and micrographs of the stylets from A–B. the designated holotype (FMNH http://id.luomus.fi/KV.545), and C–D. reference specimens (C: HU XXIII.3.20, D: HU XXIII). E–I. Micrographs of live specimens, including E., an overview of an entire specimen, F. the proboscis (E–F: HU XXIII.3.27), G. the stylet and seminal vesicle, H. the pharynx, and I. the ovary (G–I: HU XXIII.3.55).	FIGURE 5. Lagenopolycystis canariensis sp. nov. A–D. Camera lucida drawings and micrographs of the stylets from A–B. the designated holotype (FMNH http://id.luomus.fi/KV.545), and C–D. reference specimens (C: HU XXIII.3.20, D: HU XXIII). E–I. Micrographs of live specimens, including E., an overview of an entire specimen, F. the proboscis (E–F: HU XXIII.3.27), G. the stylet and seminal vesicle, H. the pharynx, and I. the ovary (G–I: HU XXIII.3.55).	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFACFFA1FF39F97449DAEC72.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823221/files/figure.png	https://doi.org/10.5281/zenodo.15823221	FIGURE 6. Lagenopolycystis conglobata sp. nov. A–F. Camera lucida drawings of the stylets from A. the designated holotype (FMNH http://id.luomus.fi/KV.546), and B–F. reference specimens (B: HU XXIII.3.46, C:HU XXIII.4.13, D: HU XXIII.3.47; E: HU XXIII.4.16, F: HU XXIII.4.01; same scale). For A–B., the sclerotised tube of the seminal receptacle is also drawn. G–I Micrographs of the stylet in fixed specimens, including G. the holotype (H: HU XXIII.3.44; I: HU XXIII.3.45). J–L. Micrographs of a live specimen from Sardinia (now HU XXIII.4.09), including J., an overview of the entire specimen, K. the proboscis, L. the stylet and seminal vesicle.	FIGURE 6. Lagenopolycystis conglobata sp. nov. A–F. Camera lucida drawings of the stylets from A. the designated holotype (FMNH http://id.luomus.fi/KV.546), and B–F. reference specimens (B: HU XXIII.3.46, C:HU XXIII.4.13, D: HU XXIII.3.47; E: HU XXIII.4.16, F: HU XXIII.4.01; same scale). For A–B., the sclerotised tube of the seminal receptacle is also drawn. G–I Micrographs of the stylet in fixed specimens, including G. the holotype (H: HU XXIII.3.44; I: HU XXIII.3.45). J–L. Micrographs of a live specimen from Sardinia (now HU XXIII.4.09), including J., an overview of the entire specimen, K. the proboscis, L. the stylet and seminal vesicle.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFAAFFA0FF39F9A54902EDB4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823223/files/figure.png	https://doi.org/10.5281/zenodo.15823223	FIGURE 7. Lagenopolycystis mutabilis sp. nov. A–D. Camera lucida drawings of the stylets from A. the holotype (FMNH http:// id.luomus.fi/KV.547), and B–D. reference specimens (same scale) (B: HU XXIII.4.29; C: HU XXIII.4.30; D: HU XXIII.4.35). E–F. Micrographs of the stylets from E. the holotype, and F. a reference specimen (HU XXIII.4.29).	FIGURE 7. Lagenopolycystis mutabilis sp. nov. A–D. Camera lucida drawings of the stylets from A. the holotype (FMNH http:// id.luomus.fi/KV.547), and B–D. reference specimens (same scale) (B: HU XXIII.4.29; C: HU XXIII.4.30; D: HU XXIII.4.35). E–F. Micrographs of the stylets from E. the holotype, and F. a reference specimen (HU XXIII.4.29).	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA9FFA2FF39FF6849C9ECE2.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823225/files/figure.png	https://doi.org/10.5281/zenodo.15823225	FIGURE 8. Lagenopolycystis poena sp. nov. A–B. Micrographs of the stylet, from A. the holotype (FMNH http://id.luomus. fi/KV.548), and B. a reference specimen (HU XXIII.4.42). C–G. Camera lucida drawings of the stylets from C. the designated holotype (FMNH http://id.luomus.fi/KV.548),andD–G.reference specimens(same scale)(D: HUXXIII.4.42, E:HUXXIII.4.44, F: HU XXIII.4.43). H–K. Micrographs of a live specimen from Portugal (HU XXIII.4.42), including H., an overview of the entire specimen, I. the pharynx, J. the proboscis, and K. the vitellarium and ovary.	FIGURE 8. Lagenopolycystis poena sp. nov. A–B. Micrographs of the stylet, from A. the holotype (FMNH http://id.luomus. fi/KV.548), and B. a reference specimen (HU XXIII.4.42). C–G. Camera lucida drawings of the stylets from C. the designated holotype (FMNH http://id.luomus.fi/KV.548),andD–G.reference specimens(same scale)(D: HUXXIII.4.42, E:HUXXIII.4.44, F: HU XXIII.4.43). H–K. Micrographs of a live specimen from Portugal (HU XXIII.4.42), including H., an overview of the entire specimen, I. the pharynx, J. the proboscis, and K. the vitellarium and ovary.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA9FFBFFF39F91449BAE88A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823229/files/figure.png	https://doi.org/10.5281/zenodo.15823229	FIGURE 9. Lagenopolycystis mandelai A. Micrograph and B. camera lucida drawing of the stylet of the holotype (schematized after Willems et al., 2017). C–F. Micrographs of a live specimen, including C., an overview of the entire specimen, D. the proboscis, E. the pharynx, and F. the male copulatory system.	FIGURE 9. Lagenopolycystis mandelai A. Micrograph and B. camera lucida drawing of the stylet of the holotype (schematized after Willems et al., 2017). C–F. Micrographs of a live specimen, including C., an overview of the entire specimen, D. the proboscis, E. the pharynx, and F. the male copulatory system.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA9FFBFFF39F91449BAE88A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823233/files/figure.png	https://doi.org/10.5281/zenodo.15823233	FIGURE 10. Maps indicating the geographical distribution of the species of Lagenopolycystis: A. Worldwide, B. Localities in the Mediterranean and along the Iberian Atlantic coast.	FIGURE 10. Maps indicating the geographical distribution of the species of Lagenopolycystis: A. Worldwide, B. Localities in the Mediterranean and along the Iberian Atlantic coast.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
98575F6FFFA9FFBFFF39F91449BAE88A.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15823235/files/figure.png	https://doi.org/10.5281/zenodo.15823235	FIGURE 11. Part of the topology of Tessens et al. (2014), with taxon names updated based on recent literature (Schockaert et al., 2014, 2019) and newly established species from this study. Branches without symbols indicate maximum support. Symbols above branches represent posterior probabilities (pp), where ** denotes 0.98 <pp <1. Symbols below branches indicate bootstrap values (bp): ●● denotes 85 <bp <100, and ● denotes 70 <bp <85.	FIGURE 11. Part of the topology of Tessens et al. (2014), with taxon names updated based on recent literature (Schockaert et al., 2014, 2019) and newly established species from this study. Branches without symbols indicate maximum support. Symbols above branches represent posterior probabilities (pp), where ** denotes 0.98 <pp <1. Symbols below branches indicate bootstrap values (bp): ●● denotes 85 <bp <100, and ● denotes 70 <bp <85.	2025-07-04	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.		Zenodo	biologists	Monnens, Marlies;Schockaert, Ernest R.;Diez, Yander L.;Revis, Nathalie;Janssen, Toon;Jouk, Philippe E. H.;Tessens, Bart;Van Steenkiste, Niels W. L.;Artois, Tom J.			
