identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
A56C153CFFD9FFCAFF55FB06FA94FE43.text	A56C153CFFD9FFCAFF55FB06FA94FE43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhizocephala MULLER 1862	<div><p>ORDER  RHIZOCEPHALA MÜLLER, 1862</p><p>Diagnosis (amended): The monophyly is assured by the molecularly based analyses of  Glenner &amp; Hebsgaard (2006) and Pérez-Losada et al. (2008). In addition, the following morphology-based apomorphies apply. All stages, including the nauplii and cyprids, lack any trace of an alimentary canal. The cyprids carry an aesthetasc seta and four additional setae on the fourth antennular segment. The male cyprids also carry a similar, but longer aesthetasc proximally on the third antennular segment. These are all parasites on  Crustacea with a life cycle involving an initial and completely internal phase. This starts as a stage featuring an epitheliumenveloped bladder (primordium) containing a small lump of cells (nucleus) that represent the prospective visceral sac with the ovary. The adult parasite consists of an internal ramifying and nutrient-absorbing root system and an external sac (externa) housing the reproductive organs. All parasitic stages are without any segmentation or appendages.</p><p>Remarks: The morphological diagnosis is derived from  Glenner &amp; Høeg (1994) and Høeg &amp; Rybakov (1992). The setation pattern of the antennules is subject to modification and loss within the order (Høeg &amp; Rybakov, 1996a). A detailed account of the biology and morphology is given by Høeg (1995), Høeg &amp; Lützen (1995) and Høeg et al. (2005). Larval morphology and development are treated by Høeg (1985, 1987, 1990, 1992), Martin et al. (2014),  Glenner et al. (1989, 2010) and  Glenner (2001). The existing general taxonomy is treated by Høeg &amp; Lützen (1985, 1996), Høeg &amp; Rybakov (1992), Øksnebjerg (2000) and Høeg et al. (2019).</p></div>	https://treatment.plazi.org/id/A56C153CFFD9FFCAFF55FB06FA94FE43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFD9FFCAFCD3FDBAFA8BFBB1.text	A56C153CFFD9FFCAFCD3FDBAFA8BFBB1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triangulidae HOEG & GLENNER 2020	<div><p>FAMILY  TRIANGULIDAE HØEG &amp; GLENNER FAM. NOV. (MONOGENERIC)</p><p>Type genus:  Triangulus Smith, 1906 .</p><p>Type species:  Triangulus munidae Smith, 1906 .</p><p>LSID: urn:lsid:zoobank.org:act: A57984C8-7B8A-420F- AD6C-45E62B5445C8</p><p>Diagnosis: By the molecular data. Externa asymmetrical or symmetrical. Dorsal mesentery generally broad, ventral mesentery narrower and shorter. Receptacles, receptacle ducts and colleteric glands arranged asymmetrically within visceral mass. Receptacle ducts not ending in small papillae.</p></div>	https://treatment.plazi.org/id/A56C153CFFD9FFCAFCD3FDBAFA8BFBB1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDBFFC8FF18F9B4FB0AFEF8.text	A56C153CFFDBFFC8FF18F9B4FB0AFEF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Peltogasterellidae HOEG & GLENNER 2020	<div><p>FAMILY  PELTOGASTERELLIDAE HØEG &amp; GLENNER FAM. NOV.</p><p>Type genus:  Peltogasterella Krüger, 1912 .</p><p>Type species:  Peltogasterella sulcata (Lilljeborg, 1859) .</p><p>LSID: urn:lsid:zoobank.org:act: 43B04CD0-B6CF- 4FAA-89DA-E8A0CD422EFC</p><p>Diagnosis: By the molecular data. Cylindrical or semicylindrical colonial externae all connected to a common internal root system; externae produce only one or two broods (Høeg &amp; Lützen, 1985; Yoshida et al., 2015) and are then replaced by a new generation of externae that do not develop in a one-to-one relationship to the preceding ones.</p></div>	https://treatment.plazi.org/id/A56C153CFFDBFFC8FF18F9B4FB0AFEF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDAFFC9FCF5FD37FA49FB5E.text	A56C153CFFDAFFC9FCF5FD37FA49FB5E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyascidae HOEG & GLENNER 2020	<div><p>POLYASCIDAE HØEG &amp; GLENNER FAM. NOV.</p><p>Type genus:  Polyascus Glenner, Lützen &amp; Takahashi, 2003 .</p><p>Type species:  Polyascus (syn.  Sacculina)  polygeneus (Lützen &amp; Takahashi, 1997) .</p><p>LSID: urn:lsid:zoobank.org:act: A733BE00-3B81- 4C57-BC0D-2A7DE85BDF89</p><p>Diagnosis: By the molecular data. External morphology resembling that in the  Sacculinidae, but external cuticle smooth or almost smooth, normally without excrescences. One or two globular receptacles located at base of stalk, outside ovary. Two receptacle ducts with short, slightly sinuous tubes lined by heavy cuticle. Multiple externae can form by budding from the root system, but are only obligatory in species of  Polyascus .</p></div>	https://treatment.plazi.org/id/A56C153CFFDAFFC9FCF5FD37FA49FB5E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDAFFC9FF37FD9CFD4EFC51.text	A56C153CFFDAFFC9FF37FD9CFD4EFC51.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sacculinidae LILLJEBORG 1861	<div><p>FAMILY  SACCULINIDAE LILLJEBORG 1861, AMENDED</p><p>Type genus:  Sacculina Thompson, 1836 .</p><p>Type species:  Sacculina carcini Thompson, 1836 .</p><p>Diagnosis: By the molecular data. Externa laterally compressed. Mantle opening more or less opposite stalk, situated on the anterior margin or displaced to the left side. Thin dorsal mesentery only, extending from the stalk to the mantle opening, but often shorter or absent. Colleteric glands with a number of branched tubes, situated in the central part of the lateral surface of the visceral mass. Receptacles placed either dorsally or far posteriorly in the visceral mass, or in the basal region of the stalk (from Øksnebjerg, 2000).</p></div>	https://treatment.plazi.org/id/A56C153CFFDAFFC9FF37FD9CFD4EFC51	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDDFFCEFF6CFAFCFC3AFB15.text	A56C153CFFDDFFCEFF6CFAFCFC3AFB15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parasacculina HOEG & GLENNER 2020	<div><p>PARASACCULINA HØEG &amp; GLENNER GEN. NOV.</p><p>Type species:  Polyascus (syn.  Sacculina)  shiinoi (Lützen et al., 2016) .</p><p>LSID: urn:lsid:zoobank.org:act: EE2FD3C8-F9C1- 49EE-AAE5-80176863160B</p><p>Diagnosis: As for the family, but, unlike  Polyascus, without obligatory colonial externae.</p><p>Etymology: The name is chosen to indicate the close morphological similarity to species of  Sacculina .</p><p>Species:  Parasacculina leptodiae (Guerin-Ganivèt, 1911),  Parasacculina oblonga (Lützen &amp; Yamaguchi,</p><p>1999),  Parasacculina shiinoi (Lützen et al., 2016), Pa r a s a c c u l i n a s i n e n s i s (B o s c h m a, 1 9 3 3) a n d  Parasacculina yatsui (Boschma, 1936) .</p><p>Remarks: The genus is paraphyletic, and this is at present the only practical solution. It consists of species formerly placed in  Sacculina that by molecular data do not belong in the redefined  Sacculinidae, but form a monophyletic unit together with, but not within, the monophyletic genus  Polyascus . The type species has recently been investigated by both molecular and morphological methods, and type specimens are available and in good condition.  Parasacculina (syn.  Sacculina)  yatsui has previously (e.g.  Glenner et al., 2010) been identified erroneously as  Sacculina confragrosa Boschma, 1933 (see Kobayashi et al., 2018). A molecular analysis of the true  Sacculina confragrosa has not yet been published.</p><p>FAMILY  POLYSACCIDAE LÜTZEN &amp; TAKAHASHI, 1996 (MONOGENERIC)</p><p>Type genus:  Polysaccus Høeg &amp; Lützen, 1993 .</p><p>Ty p e s p e c i e s: Po l y s a c c u s (s y n. Th o m p s o n i a)  mediterraneus (Caroli, 1929) .</p><p>Diagnosis: By morphology as in the study by Øksnebjerg (2000). The molecular analysis clearly separates the only analysed species,  Polysaccus japonicus Høeg &amp; Lützen, 1993, from all other families recognized here.</p></div>	https://treatment.plazi.org/id/A56C153CFFDDFFCEFF6CFAFCFC3AFB15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDDFFCEFF19FC92FDF9FA9D.text	A56C153CFFDDFFCEFF19FC92FDF9FA9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polyascus Glenner, Lutzen & Takahashi 2003	<div><p>POLYASCUS GLENNER, LÜTZEN &amp; TAKAHASHI, 2003</p><p>Type species:  Polyascus polygeneus (Lützen &amp; Takahashi, 1997) .</p><p>Diagnosis (amended): By the molecular data. Externa morphology as for the family, but with obligatory multiple (colonial) externae that are replaced in successive generations</p><p>Species:  Polyascus gregarius (Okada &amp; Miyashita, 1935),  Polyascus planus (Boschma, 1933) and  Polyascus polygeneus (Lützen &amp; Takahashi, 1997) .</p><p>Remarks: The diagnosis is amended from  Glenner et al. (2003) by specifying also the obligatory presence of multiple externae, which separates it from the new genus  Parasacculina defined below.</p></div>	https://treatment.plazi.org/id/A56C153CFFDDFFCEFF19FC92FDF9FA9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDCFFCFFF03FEE2FD16FCE9.text	A56C153CFFDCFFCFFF03FEE2FD16FCE9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clistosaccidae HOEG & RYBAKOV 1992	<div><p>FAMILY  CLISTOSACCIDAE HØEG &amp; RYBAKOV, 1992</p><p>Type genus:  Clistosaccus Lilljeborg, 1861 .</p><p>Type species:  Clistosaccus paguri Lilljeborg, 1861 .</p><p>Diagnosis (amended): By molecular data. Morphology-based apomorphies are a cypris larvae with the fourth antennular segment carrying only a single bifurcate seta situated at the apex. A reduced, near-identical pattern of setation on the cypris carapace is depicted by Jensen et al. (1994) and not found elsewhere in rhizocephalans. The second pair of lattice organs (LO 2) in the cyprids is shaped like a hairpin, another morphology unknown elsewhere in rhizocephalans.</p></div>	https://treatment.plazi.org/id/A56C153CFFDCFFCFFF03FEE2FD16FCE9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDCFFCFFF01FA3CFAA7F9EB.text	A56C153CFFDCFFCFFF01FA3CFAA7F9EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Thompsoniidae HOEG & RYBAKOV 1992	<div><p>FAMILY  THOMPSONIIDAE HØEG &amp; RYBAKOV, 1992</p><p>Type genus:  Thompsonia Kossmann, 1874 .</p><p>Type species:  Thompsonia globosa Kossmann, 1874 .</p><p>Diagnosis (amended): By molecular data. Morphology-based characters are numerous globular or club-shaped colonial externae without receptacles that emerge simultaneously from the host. and each releases only a single brood. These externae are replaced through several generations of new externae that often appear in increasing numbers. The cyprids carry one bifurcated seta and some smaller setae and structures apically on the fourth segment. The second pair of lattice organs is not different from the remaining four pairs.</p><p>H o s t s: A n o m u r a, B r a c h y u r a, C a r i d e a a n d Stomatopoda.</p><p>Genera: Diplothylacus Høeg &amp; Lützen, 1993 (four spp.),  Jensia Boyko &amp; Williams, 2015 (two spp.),  Thompsonia (five spp.) amd  Thylacoplethus Coutière, 1902 (13 spp.).</p><p>Remarks: The family comprises taxa that originate at node 13 in Figure 3. The family is monophyletic in all relevant analyses based on molecular data.  Jensia was formerly named ‘  Pottsia ’ Høeg &amp; Lützen, 1993, but this was a preoccupied name (see Hiller et al., 2015). Høeg &amp; Lützen (1993) gave an extensive account of the systematics, morphology and phylogeny of this family, and to this is added the paper by Hiller et al. (2015). Among all Rhizocephala,  Thompsoniidae have the widest taxonomic range of host animals.</p><p>FAMILY  CHTHAMALOPHILIDAE BOCQUET-VEDRINE, 1961</p><p>Type genus:  Chthamalophilus Bocquet-Vedrine, 1957 .</p><p>Type species:  Chthamalophilus delagei Bocquet-Vedrine, 1957 .</p><p>Diagnosis (amended): By molecular data. Morphology-based diagnosis amended from Høeg &amp; Rybakov (1992) is cypris larvae being minute (&lt;100 µm long) and completely lacking a thorax, whence they can move only by walking on the antennules. The fourth segment is reduced to a mere rudiment. The cypris carapace is furnished with only four long setae located posteriorly. The externa is always surrounded by a double layer of cuticle separated by a fluid filled space, rendering the externa surface distinctly refringent. The male organs are invaginated from the mantle into the mantle cavity as free-floating bodies enveloped in cuticle. These ‘primary spermatogenic islets’ later split into several ‘secondary islets’ devoid of cuticle and in which spermatogenesis proceeds (Høeg et al., 1990, 2019).</p></div>	https://treatment.plazi.org/id/A56C153CFFDCFFCFFF01FA3CFAA7F9EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDFFFCCFF62FBA4FE3BF989.text	A56C153CFFDFFFCCFF62FBA4FE3BF989.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Duplorbidae HOEG & RYBAKOV 1992	<div><p>FAMILY  DUPLORBIDAE HØEG &amp; RYBAKOV, 1992</p><p>Type genus:  Duplorbis Smith, 1906 .</p><p>Type species:  Duplorbis calathurae Smith, 1906 .</p><p>Diagnosis (amended): The cyprids have long and thick frontal filaments. Additional characters are externae with a mesenteric canal, as in chthamalophilids but lacking the double cuticle and refringent appearance of this family. The male organs are spermatogenic islets that originate as in the  Chthamalophilidae, but they continue to be sheathed in a cuticle that continues to increase in thickness, and spermatogenesis proceeds without any formation of secondary islets (Mourey, 1974; Høeg &amp; Rybakov, 1992).</p></div>	https://treatment.plazi.org/id/A56C153CFFDFFFCCFF62FBA4FE3BF989	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
A56C153CFFDFFFCCFCF1FD72FA89FABF.text	A56C153CFFDFFFCCFCF1FD72FA89FABF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pirusaccidae HOEG & GLENNER 2020	<div><p>FAMILY  PIRUSACCIDAE HØEG &amp; GLENNER FAM. NOV.</p><p>Type genus:  Pirusaccus Lützen, 1985 .</p><p>Type species:  Pirusaccus socialis Lützen, 1985 (by monotypy).</p><p>LSID: urn:lsid:zoobank.org:act: E13A86B8-C88E-4709-8A7A-599CC6B58C6E</p><p>Diagnosis: Colonial externae of an elongated club shape. The male organs are bodies (‘spermatogenic islets’) floating free in the mantle cavity. Mesentery present; mesenteric canal absent.</p><p>Hosts: On the galatheoid species  Galacantha (syn.  Munidopsis) rostrata A. Milne Edwards, 1880.</p></div>	https://treatment.plazi.org/id/A56C153CFFDFFFCCFCF1FD72FA89FABF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Høeg, Jens T.;Noever, Christoph;Rees, David J.;Crandall, Keith A.;Glenner, Henrik	Høeg, Jens T., Noever, Christoph, Rees, David J., Crandall, Keith A., Glenner, Henrik (2020): A new molecular phylogeny-based taxonomy of parasitic barnacles (Crustacea: Cirripedia: Rhizocephala). Zoological Journal of the Linnean Society 190: 632-653
