taxonID	type	description	language	source
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	description	(Figs 1 A – E)	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	materials_examined	Type locality. Taiwan, New Taipei, 0.8 km SEE of Wulai City, 24.85652 ° N 121.5536 ° E, 320 m a. s. l. Type material. HOLOTYPE: J (NMNS): ʻTAIWAN, New Taipei / 0.8 km SEE of Wulai City, Laka / Trail; 24.85652 ° N 121.5536 ° E / 5. v. 2018; 320 m; Fikáček, Leong / & Chang lgt.; 2018 - TW 11, // sifting of small leaf litter / accumulations in the secondary / lowland moist forest on a steep / slope, with sparse understoryʼ. PARATYPES: TAIWAN: NEW TAI ඉEI: 1 J (PBPC): ʻTAIWAN, New Taipei / 2.5 km SWW of Wulai City / Meilushan Mt. above Baoquing / Temple; 24.85445 ° N 121.53 ° E / 760 m; 6. v. 2018; Fikáček, Chang / & Zelený lgt.; 2018 - TW 12 // leaf litter accumulations in / primary moist forest with sparse / understory: siftingʼ; 1 J (NMPC): ʻTAIWAN: New Taipei, Fushan / Botanical Garden, ForestGEO / Vegetation Ecology plot / quadrat 1711, 665 m // 24.7514 ° N 121.555 ° E; 7. vii. 2022 / M. Fikáček; TW 2022 _ 005 B // FS 1 - 029 / Fushan / (TW 2022 _ 005 B) / OTU 687 (adult) ʼ; 1 ♀ (PBPC): TAIWAN: New Taipei / Yanmingshan National Park / Erzhiping Trail, 830 m / 25.17932 [° N] 121.51613 [° E] / 13. viii. 2021, sifting leaf / litter, HY Lin, WZ Tseng [leg.].	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	description	Description. Habitus as in Fig. 1 A. Body length 2.40 – 2.50 mm (HT 2.50 mm, mean 2.45 mm; n = 4). Colour of dorsal surface in HT and one PT ferruginous without bronze luster, elytral base, apex and anterior part of head barely lighter, in two PT upper surface slightly darker; ventral surface and legs ferruginous, mouthparts and antennae dark yellowish. Head. Clypeus bordered, with slightly and broadly protruded median part; lateral teeth bordered, moderately acute, distinctly projecting anteriorly, pointed at tip, divided from supraantennal plates by obtuse notch; clypeal field sub-quadrate, irregularly narrowed posteriorly, smooth, separated from frons by deep and broad transverse furrow; frons convex, smooth; supraantennal plates strongly convex, with blunt carina at top of convexity. Frontal furrows deep, broad, parallel to eyes mid-length then abruptly and strongly diverging posteriorly around eyes. Eyes moderately small, convex, facets distinct; genae very small, enclosing eyes in about posterior fifth. Antennae relatively long, antennomeres 5 – 10 submoniliform, about 1.2 – 1.3 times as long as wide. Neck with minute isodiametric reticulation laterally and with a few striae and punctures below eyes, constriction absent. Pronotum subglobose, in lateral view moderately flattened posteriorly; outline between lateral SP slightly convex; indistinctly attenuated anteriorly 1.12 – 1.15 (HT 1.12, mean 1.13) times as wide as long, 1.48 – 1.54 (HT 1.53, mean 1.50) times as wide as head, broadest in about posterior third, with rounded anterior angles. Anterior transverse impression deep, broadened in middle, impunctate, sparse and rough cross-striate, cross-striae especially in middle third rather deep and subparallel; median line moderately deep, deeper basally, lateral channel deep and moderately broad, gently broadened anteriorly up to anterior SP, reflexed lateral margin just surpassing posterior SP. Surface shiny, with very few, almost indistinct transverse wrinkles and sparse micropunctures, lateral channel, without reticulation. Elytra shortly ovate; disc moderately, apex more strongly convex in lateral view; 1.43 – 1.48 (HT 1.44, mean 1.46) times as long as wide, 1.18 – 1.23 (HT 1.18, mean 1.20) times as wide as pronotum. Elytra widest posterior to anterior third; base sloping to moderately rounded humeri without humeral tooth; lateral channel moderately broad from pedunculus to humeri, broadened apically; reflexed lateral margin distinct. Basal border distinct, basal tubercles and scutellar strioles absent; BSP large, isolated, situated just inside of projected extension of interval 2. Three PHSP, three PASP, two large ASP and three DSP (in interval 3, posterior one in about mid-length). Striae 1 – (2) irregular, moderately deep, stria 1 indistinctly joining BSP and joining lateral channel at apex; striae (2) – 3 irregularly, finely impressed, latero-apically mostly diminish; striae 1 – 6 (7) coarsely punctured in basal half, punctuation finer latero-apically, almost diminish, stria 8 beginning as a few very fine punctures in middle third; intervals barely convex medio-basally, flattened latero-apically. Metathoracic wings absent. Protibiae. Apical spine distinctly curved ventrolaterally; apical movable spur shorter than spine, feebly curved; distal tooth moderately large, sharp, proximal one much finer, almost indistinct. Male genitalia. Aedeagus as in Figs 1 B, C, E, median lobe in lateral view short and very broad at mid-length; apex short, broadly rounded, distinctly bent ventrad. Parameres as in Fig. 1 D, unisetose.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	diagnosis	Differential diagnosis. Reicheiodes kuankong sp. nov. is most similar to R. (Reichonippodes) nishii from Iriomote-jima Island, Japan. The new species could be distinguished by longer antennae with the antennomeres 5 – 10 about 1.2 – 1.3 times as long as wide (1.0 – 1.1 times as long as wide in R. nishii); by the pronotum having the anterior impression more roughly cross-striate medially; and especially by the elytra with its base distinctly less sloping to the more distinct humeri and with the striae latero-apically longer and more deeply punctured. The new species differs from R. taiwanensis in the body being smaller and lighter in colour (fuliginous in R. taiwanensis), in the narrower lateral channel of the pronotum (only slightly broader than reflexed lateral margin in latero-dorsal view, in contrast to twice broader in R. taiwanensis), and in the antennomeres 5 – 10 being 1.2 – 1.3 times as long as wide (1.0 – 1.1 times as long as wide in R. taiwanensis). For differences from R. ursinus sp. nov. see under that species.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	etymology	Etymology. The new species is named after Kuan Kong, the deity worshiped in Chinese folk religion, Taoism and Buddhism, and usually depicted with a red face, resembling the reddish body coloration of this species. The name is a noun in nominative singular, standing in apposition. Collection circumstances. All specimens were collected by sifting of moist leaf litter in primary lowland and lower montane forests at altitudes of 320 – 830 m a. s. l.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE4D446FF4B3A66FDEFFC37.taxon	distribution	Distribution. Endemic to Taiwan. Only known from northern Taiwan, so far known from the northernmost part of the central mountain range (Fushan and Wulai) and from Yanmingshan National Park.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE5D446FF333A65FB41F817.taxon	description	(Figs 1 K – O)	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE5D446FF333A65FB41F817.taxon	materials_examined	Type locality. Taiwan, Taitung, Haiduan, Jinping Logging Trail km 15, 23.163342 ° N 121.144153 ° E. Type material. HOLOTYPE: J (NMNS): ʻTAIWAN, Taitung, Haiduan / Jinping Logging Trail, km 15 / 23.163342 ° N 121.144153 ° E, 1520 m / 3. iii. 2024, BH Ho [leg.], TW 2024 - 004 ʼ. PARATYPES: TAIWAN: TAI- TUNG: 23 JJ 17 ♀♀ (BMNH, BHHC, NMNS, NMPC, PMFI, PBPC): same label data as holotype.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE5D446FF333A65FB41F817.taxon	description	Description. Habitus as in Fig. 1 K. Body length 2.20 – 2.60 mm (HT 2.40 mm, mean 2.40 mm; n = 30). Colour of dorsal surface ferruginous to dark ferruginous without bronze luster; ventral surface and legs ferruginous, mouthparts and antennae dark yellowish. Head. Clypeus bordered, with very slightly and broadly to moderately protruded median part; lateral teeth bordered, moderately acute, distinctly projecting anteriorly, pointed at tip, divided from supraantennal plates by obtuse notch; clypeal field sub-quadrate, smooth, separated from frons by deep and broad transverse furrow; frons convex, smooth; supraantennal plates strongly convex, with blunt carina at top of convexity. Frontal furrows deep, moderately broad, parallel to eyes mid-length then abruptly and strongly diverging posteriorly around eyes. Eyes rather small, strongly convex, facets distinct; genae small, gently enclosing eyes posteriorly. Antennae relatively short, antennomeres 5 – 10 moniliform. Neck without isodiametric reticulation, with a few punctures below eyes, constriction absent. Pronotum subglobose, in lateral view moderately flattened posteriorly; outline between lateral SP slightly convex; indistinctly attenuated anteriorly 1.09 – 1.16 (HT 1.11, mean 1.13) times as wide as long, 1.45 – 1.54 (HT 1.52, mean 1.48) times as wide as head, broadest in about posterior third, with rounded anterior angles. Anterior transverse impression deep, broadened in middle, impunctate, very sparsely and shortly cross-striate, especially in middle third; median line rather fine on disc, deeper basally, lateral channel deep and moderately broad, not broadened anteriorly up to anterior SP, reflexed lateral margin just surpassing posterior SP. Surface shiny, with very few, almost indistinct transverse wrinkles and sparse micropunctures, lateral channel shiny, without reticulation. Elytra shortly ovate; disc moderately, apex more strongly convex in lateral view; 1.47 – 1.53 (HT 1.48, mean 1.50) times as long as wide, 1.21 – 1.28 (HT 1.26, mean 1.24) times as wide as pronotum. Elytra widest after anterior third; base sloping to moderately rounded humeri without tooth; lateral channel moderately broad from pedunculus to humeri, broadened apically; reflexed lateral margin distinct. Basal border distinct, basal tubercles and scutellar strioles absent; BSP mostly missing, in nine PT present on one elytron. Three PHSP, three PASP, two large ASP and two DSP (in anterior part of interval 3, posterior DSP missing). Striae 1 – (2) irregular, moderately deep, stria 1 joining lateral channel at apex; striae (2) 3 – 5 irregularly, finely impressed, latero-apically mostly diminish; striae 1 – 4 punctures rough in basal half, latero-apically punctuation finer, stria 5 – 7 punctures fine latero-apically, diminish on apex, stria 8 indistinct; inner intervals barely convex medio-basally, flattened latero-apically. Metathoracic wings absent. Protibiae. Apical spine moderately curved ventrolaterally; apical movable spur shorter than spine, feebly curved; distal tooth moderately large, sharp, proximal one much finer, almost indistinct. Male genitalia. Aedeagus as in Figs 1 L, M, O. Medi- an lobe in lateral view moderately broad, rather narrow at mid-length, ventral margin almost evenly regularly concave, apex short, broadly rounded, indistinctly bent ventrad, apex in ventral view broadly rounded at tip, distinctly turned right. Parameres as in Fig. 1 N, unisetose. Differential diagnosis. Reicheiodes ursinus sp. nov. is similar to Reicheiodes (Reichonippodes) kuankong sp. nov. The new species can be distinguished by shorter antennae with the antennomeres 5 – 10 about as long as broad, by the elytra with only two DSP and none or unilaterally one BSP, and by having finer striae punctuation latero-apically.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE5D446FF333A65FB41F817.taxon	etymology	Etymology. Ursinus is a Latin adjective derived from Ursus (bear). We name the new species in this way as its type locality is also famous for the population of the Taiwanese bear (Ursus thibetanus formosanus Swinhoe, 1864). Collection circumstances. All specimens were collected by sifting moist leaf litter in the primary montane forest.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE5D446FF333A65FB41F817.taxon	distribution	Distribution. Endemic to Taiwan. Only known from the type locality on the eastern slope of the central mountain range above the Taitung Valley.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE3D440FF2A3ED3FE0BFB46.taxon	description	(Figs 1 F – J)	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE3D440FF2A3ED3FE0BFB46.taxon	materials_examined	Type material examined. HOLOTYPE: ♀ (NMPC): ʻTaiwan, Taichung / Dasyueshan forest / 24.2334 [° N] 120.9664 [° E] / 2140 m; 10. viii. 2013, sift. / TW 01, V. Grebennikovʼ. Additional material examined. TAIWAN: TAICHUNG: 4 JJ 6 ♀♀ (ADWA, PBPC): ʻTaiwan, Taichung; 178 / Xueshan rd. (to Daxueshan / Nat. Forest Recr. Area) / Prim. Forest, 24 ° 13 ’ 36.4 ” N / 120 ° 58 ’ 34.7 ” E, 1700 m [sic!, the altitude at the GPS position given is 1990 m!], 30. v. 2019 / leg. Berger-Dostalʼ; 2 JJ (NMNS, NMPC): Taiwan, Taichung / Daxueshan Nat. Forest Area / at Rd. 1 S of Shaolaishan Mt. / 24.22601 [° N] 120.975772 [° E] / 28. vi. 2023 / Fikáček, Růžička, Ho, Hu, Král [leg.] / TW 2023 - 031 // sifting of small accumulations / of leaf litter in a montane forest / with huge trees. Supplementary description. Male as in Fig. 1 F. Measurements (n = 13, including holotype). Body length 2.60 – 2.95 mm (mean 2.75 mm, HT 2.75 mm); pronotum 1.10 – 1.18 (mean 1.13, HT 1.18) times as wide as long, 1.50 – 1.59 (mean 1.55, HT 1.59) times as wide as head; Elytra 1.40 – 1.48 (mean 1.45, HT 1.40) times as long as wide, 1.21 – 1.28 (mean 1.24, HT 1.22) times as wide as pronotum. Male genitalia. Aedeagus as in Figs 1 G, H, J; median lobe in lateral view moderately long, moderately broad at mid-lenght, apex short, very broadly rounded, barely bent ventrad. Parameres as in Fig. 1 I, unisetose. Collection circumstances. All specimens were collected by sifting leaf litter in a primary montane forest.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
A44087B8FFE3D440FF2A3ED3FE0BFB46.taxon	distribution	Distribution. Endemic to Taiwan. The species is only known from the Daxueshan area in the central mountain range, from two close localities along the Daxueshan Rd. at altitudes 1990 – 2140 m a. s. l.	en	Bulirsch, Petr, Magrini, Paolo, Ho, Bin-Hong, Fikáček, Martin (2025): Species of the genus Reicheiodes from Taiwan (Coleoptera: Carabidae: Scaritinae). Acta Entomologica Musei Nationalis Pragae 65 (1): 61-70, DOI: 10.37520/aemnp.2025.005, URL: https://doi.org/10.37520/aemnp.2025.005
