taxonID	type	description	language	source
AF0B87BCFE23FFC7FE1FFE34FCE9FC5A.taxon	description	(figures 1 – 13)	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE23FFC7FE1FFE34FCE9FC5A.taxon	materials_examined	Type material One female holotype (USNM 1010499) and one male allotype (USNM 1010500) preserved in alcohol, 10 dissected female (EMUCOP- 0799 - 08, EMUCOP- 0799 - 09, EMUCOP- 0799 - 10, EMUCOP- 0799 - 11, EMUCOP- 0799 - 12, EMUCOP- 0799 - 13, EMUCOP- 0799 - 14, EMUCOP- 0799 - 15, EMUCOP- 0799 - 16, EMUCOP- 0799 - 17) and five dissected male (EMUCOP- 0799 - 03, EMUCOP- 0799 - 04, EMUCOP- 0799 - 05, EMUCO- 0799 - 06, EMUCOP- 0799 - 07) paratypes, six female and three male paratypes (USNM 1010501) and six female (EMUCOP- 0799 - 01) and two male (EMUCOP- 0799 - 02) paratypes preserved in alcohol. July 1997; coll. A. Rocha-Olivares, J. W. Fleeger and D. Foltz.	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE23FFC7FE1FFE34FCE9FC5A.taxon	materials_examined	Type locality Salton Sea, California (33 ° 13 ’ N, 115 ° 52 ’ W).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE23FFC7FE1FFE34FCE9FC5A.taxon	etymology	Etymology The species was named in honour of Dr Deborah Dexter (San Diego State University, Department of Biology), who provided us with Cletocamptus specimens from the Salton Sea (California).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE23FFC7FE1FFE34FCE9FC5A.taxon	description	Description Female. Habitus (figure 1) tapering posteriorly; total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 540 to 715 M m (mean, 605 M m, N ~ 12; holotype, 620 M m). Rostrum set off, triangular, with pair of setules subapically. Cephalic shield ornamented with fine and long spinules along margin dorsolaterally (figure 1 A, B). Dorsal and lateral surface of free thoracic somites (P 2 – P 4 bearing somites) ornamented with transverse rows of minute spinules, with longitudinal row of small spinules close to posterior margin and with long spinules along posterior margin (figures 1 A, B, 2). Dorsal and lateral surface of first urosomite (P 5 bearing somite) with transverse rows of minute spinules, with row of small spinules close to posterior margin and with relatively longer spinules along posterior margin (figures 1 A, B, 3). First and second genital somites distinct dorsally and laterally (figure 1 A, B), completely fused ventrally (figure 4 A); dorsal and lateral surface of first and second genital somite with transverse rows of minute spinules, with row of long spinules along posterior margin of both somites, and with relatively longer spinules laterally, ventrally ornamented with spinules as shown in figure 4 A. Fourth and fifth urosomite as in second genital somite dorsally (figures 1 A, B, 3), ventral surface ornamented with transverse rows of spinules as illustrated in figure 4 A. Dorsal surface of anal somite (figure 1 A, C) ornamented with transverse rows of minute spinules and with dorsolateral strong spinules close to joint with caudal rami; rounded anal operculum furnished with spinules close to posterior margin (figures 1 A, C, 3, 5 H); with median row of strong spinules laterally. Caudal rami (figures 1 A, B, C, D, 5 H, I) about 1.5 times longer than wide; dorsal and ventral surface smooth, except for inner set of spinules close to insertion site of seta VII and close to posterior margin dorsally and ventrally; with seven elements in all. Antennule (figure 5 A) six-segmented; surface of segments smooth except for two rows of spinules on first segment. Armature formula, 1 - (1), 2 - (8), 3 - (5), 4 - (1 z [1 zae]), 5 - (1), 6 - (9 z [1 zae]). Antenna (figure 5 B) with small coxa. Allobasis armed with two abexopodal setae. Free endopodal segment ornamented with inner strong spinules proximally and subdistally; with two lateral inner spines and a slender seta (the latter arrowed in figure 5 B), and five distal elements. Exopod (figure 5 B, C) one-segmented; about seven times longer than wide; ornamented with few spinules, and armed with one lateral and two apical smooth setae. Mandible (figures 5 D, 6) robust, ornamented with rows of spinules proximally; chewing edge with four strong teeth, four elements as in figure 5 D, one pyriform element and one lateral pinnate strong seta. Palp (see figure 6) one-segmented, with two long setae unequal in length, with one small seta arising nearby. Maxillule (figure 5 E) robust; arthrite of praecoxa ornamented with few spinules, armed with one surface seta, seven distal spines and one lateral strong seta, the latter ornamented with long spinules (see insert in figure 5 E). Coxa ornamented with some spinules and armed with two slender setae. Basis ornamented with some median spinules, and armed with three apical setae; exopod and endopod represented by three setae each. Maxilla (figure 5 F): syncoxa ornamented with minute spinules along inner and outer margin, and close to joint with allobasis; with two endites, each bearing three setae (see insert in figure 5 F). Allobasis drawn into strong claw bearing one accompanying seta. Endopod represented by three setae. Maxilliped (figure 5 G) subchelate. Syncoxa ornamented with two rows of spinules and armed with a small seta on inner distal corner. Basis without armature and ornamented with one anterior and one posterior row of spinules, and with few spinules close to joint with endopod. The latter drawn into long and slender claw with one accompanying small seta. P 1 (figure 7 A): praecoxa ornamented with spinules close to joint with coxa. The latter ornamented with anterior transverse rows of spinules, and with posterior long and slender spinules in outer distal corner. Basis ornamented with median row of spinules, and with stronger spinules at base of exopod, between rami and at base of inner spine of basis. Exopod three-segmented. Endopod two-segmented, nearly as long as exopod. Armature formula as in table 1. P 2 (figure 7 B): coxa ornamented with median row of small spinules and with strong spinules close to outer distal corner anteriorly, and with some spinules close to outer distal corner posteriorly. Basis ornamented with spinules between rami and at base of endopod, and with stronger spinules at base of exopod; outer element spine-like. Exopod three-segmented and ornamented as in figure 7 B. Endopod twosegmented, reaching about the middle of second exopodal segment; first segment small, slightly wider than long and ornamented with outer and inner spinules; second segment ornamented with long spinules as shown in figure 7 B, and armed with one inner spine, one apical and one inner seta. Armature formula as in table 1. P 3 (figure 8 A): praecoxa ornamented with transverse row of spinules close to joint with coxa. The latter as in P 2. Basis as in P 2 except for seta-like outer element. Exopod as in P 2. Endopod as in P 2 except for comparatively shorter second segment. Armature formula as in table 1. P 4 (figure 8 B): praecoxa, coxa and basis as in P 3. Exopod as in P 3, except for armature formula of third segment (without inner seta). Endopod two-segmented, barely reaching tip of first exopodal segment; first segment small, slightly wider than long; second segment ornamented with inner and outer slender spinules and armed with two apical setae. P 5 (figure 4 B): exopod and baseoendopod fused. Baseoendopodal lobe longer than exopod, ornamented with sets of inner spinules, with spinules at base of apical seta and with some outer spinules; armed with one outer, one apical and four inner setae; relative length of setae as in figure 4 B. Exopod ornamented with spinules as shown in figure 4 B, and armed with five setae in all, plus outer seta of basis. P 6 (figure 4 A) represented by median plate in anterior half of first genital somite, each vestigial leg represented by a long and slender seta. Copulatory pore in the middle of genital double segment. Male. Total body length measured from tip of rostrum to posterior margin of caudal rami, ranging from 515 to 650 M m (mean, 562.8 M m, N ~ 9; allotype, 605 M m). Habitus (figure 9 A, B), anal segment and caudal rami (figure 9 C, D) as in female dorsally, except for genital somite and for two rows of spinules on anal operculum. Ventral ornamentation of third, fourth and fifth urosomites (figure 12 A) coarser than in female. Antennule (figures 12 B, 13) six-segmented; subchirocer. Last segment with two acute teeth (figure 13). Antenna, mandible, maxillule, maxilla and maxilliped (not illustrated) as in female. P 1 (figure 10 A) as in female except for dimorphic projection on inner distal corner of basis. P 2 (figure 10 B) as in female except for relatively stouter outer spines of male exopod, dimorphic inner spine on male ENP 2, and relatively shorter apical setae on second endopodal segment. Exopod of P 3 (figure 11 A) as in female except for relatively stouter outer spines. Endopod three-segmented; second segment with inner distal apophysis reaching far beyond third endopodal segment; second segment barely reaching tip of first exopodal segment, with one inner and one apical seta relatively shorter than their female homologues. P 4 (figure 11 B) as in female, except for stouter outer spines of exopod and relatively shorter setae of second endopodal segment. Both P 5 fused (figure 12 A); exopod and baseoendopod fused. Exopod ornamented with spinules at base of outer seta of basis, at base of apical elements and along inner margin; with four setae. Endopodal lobe ornamented with spinules along outer and inner margin, and at base of apical setae; with three elements in all. P 6 (figure 12 A) represented by plate; without armature. Variability Male. Paratype (EMUCOP- 0799 - 05) possesses an aberrant right P 4. Female. Paratype (EMUCOP- 0799 - 10) possesses two inner setae on left P 3 ENP 2; paratype (EMUCOP- 0799 - 14) possesses two inner setae on left P 4 EXP 2, and five setae on right P 5 BENP; paratype (EMUCOP- 0799 - 15) possesses one inner seta on right P 4 EXP 3, and five setae on right P 5 BENP; paratype (EMUCOP- 0799 - 16) possesses an aberrant outer spine on right P 1 EXP 3, three setae on left P 4 ENP 2, and five setae on both P 5 BENP; paratype (EMUCOP- 0799 - 09) possesses five setae on right P 5 BENP; paratype (EMUCOP- 0799 - 18) possesses a reduced inner seta on left P 5 BENP (see table 2).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE35FFCAFE03FBB5FEACFD28.taxon	description	(figures 14 – 23)	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE35FFCAFE03FBB5FEACFD28.taxon	materials_examined	Type material One female holotype (USNM 1010502) and one male allotype (USNM 1010503) preserved in alcohol, 12 dissected male (EMUCOP- 0698 - 15, EMUCOP- 0698 - 16, EMUCOP- 0698 - 17, EMUCOP- 0698 - 18, EMUCOP- 0698 - 19, EMUCOP- 0698 - 20, EMUCOP- 0698 - 21, EMUCOP- 0698 - 22, EMUCOP- 0698 - 23, EMUCOP- 0698 - 24, EMUCOP- 0698 - 25, EMUCOP- 0698 - 26) and 12 dissected female paratypes (EMUCOP- 0698 - 03, EMUCOP- 0698 - 04, EMUCOP- 0698 - 05, EMUCOP- 0698 - 06, EMUCOP- 0698 - 07, EMUCOP- 0698 - 08, EMUCOP- 0698 - 09, EMUCOP- 0698 - 10, EMUCOP- 0698 - 11, EMUCOP- 0698 - 12, EMUCOP- 0698 - 13, EMUCOP- 0698 - 14), 20 female and nine male paratypes preserved in alcohol (USNM 1010504), and 13 female (EMUCOP- 0698 - 01) and eight male (EMUCOP- 0698 - 02) paratypes preserved in alcohol. June 1998, coll. A. Rocha-Olivares, J. W. Fleeger and D. Foltz. Type locality Brine seep, Fred Stimpson Wildlife Refuge, Jackson, Alabama, USA (31 ° 23 ’ N, 87 ° 53 ’ W).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE35FFCAFE03FBB5FEACFD28.taxon	etymology	Etymology The species was named in honour of the Fred T. Stimpson Wildlife Refuge (Alabama, USA).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE35FFCAFE03FBB5FEACFD28.taxon	description	Description Female. Habitus (figure 14 A, B) tapering posteriorly; total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 415 to 660 M m (mean, 529 M m, N ~ 26; holotype, 550 M m). Rostrum set off, triangular, with pair of setules subapically. Cephalic shield ornamented with small spinules along posterior margin dorsolaterally (figure 14 A, B). Dorsal and lateral surface of free thoracic somites (P 2 – P 4 bearing somites) ornamented with transverse rows of minute spinules, and with very small spinules along posterior margin. Dorsal and lateral surface of first urosomite (P 5 bearing somite) ornamented as preceding somite. First and second genital somites distinct dorsally and laterally, completely fused ventrally (figure 15 A); dorsal and lateral surface of first and second genital somite with transverse rows of minute spinules, with row of stronger spinules along posterior margin of both somites, and with relatively longer spinules laterally; ventrally ornamented with spinules as shown in figure 15 A. Fourth and fifth urosomite as in second genital somite dorsally, except for more abundant transverse spinular rows; ventral surface ornamented with few median spinules as illustrated in figure 15 A, and with longer spinules along posterior margin. Dorsal surface of anal somite (figure 14 A) ornamented with transverse rows of minute spinules and with dorsolateral strong spinules close to joint with caudal rami; rounded anal operculum furnished with small spinules close to posterior margin. Caudal rami (figure 14 C, D) about 1.5 times longer than wide; with few spinular rows dorsally and ventrally (figures 14 C, D, 15 A); with seven elements in all. Antennule (figure 16 A) six-segmented; surface of segments smooth except for two rows of spinules on first segment. Armature formula, 1 - (1), 2 - (8), 3 - (5), 4 - (1 z [1 zae]), 5 - (1), 6 - (9 z [1 zae]). Antenna (figure 16 B) with allobasis armed with two abexopodal setae. Free endopodal segment ornamented with inner strong spinules proximally and subdistally; with two lateral inner spines and one slender seta (the latter arrowed in figure 16 B), and five distal elements. Exopod one-segmented; about seven times longer than wide; ornamented with few spinules and armed with one lateral and one apical smooth seta. Mandible (figure 16 C) robust, ornamented with rows of spinules proximally; chewing edge with four strong teeth, four elements as in figure 16 C, one pyriform element and one lateral pinnate strong seta. Palp small, one-segmented, with two long setae (see also figure 17). Maxillule (figure 16 D) robust; arthrite of praecoxa ornamented with few spinules, armed with a surface seta, seven distal spines and one lateral slender seta (see insert in figure 16 D). Coxa ornamented with some spinules and armed with two slender setae. Basis ornamented with some median spinules, and armed with three apical setae; exopod and endopod represented by three setae each. Maxilla (figure 16 E): syncoxa ornamented with minute spinules along inner and outer margin; with two endites, each bearing three setae. Allobasis drawn into strong claw bearing an accompanying seta. Endopod represented by three setae. Maxilliped (figure 16 F) subchelate. Syncoxa ornamented with spinules and armed with a small seta on inner distal corner. Basis without armature and ornamented with longitudinal spinules anteriorly and posteriorly. Endopod drawn into long and slender claw with one accompanying small seta. P 1 (figure 18 A): praecoxa ornamented with spinules close to joint with coxa. The latter ornamented with anterior transverse rows of spinules, and with posterior long and slender spinules in outer distal corner. Basis ornamented with median row of spinules, and with stronger spinules at base of exopod, between rami and at base of inner spine of basis. Exopod three-segmented. Endopod two-segmented, somewhat shorter than exopod. Armature formula as in table 1. P 2 (figure 18 B): praecoxa as in P 1. Coxa ornamented with median row of small spinules and with strong spinules close to outer distal corner anteriorly, and with some spinules close to outer distal corner posteriorly. Basis ornamented with median row of spinules close to inner distal corner, and at base of exopod; outer element of basis seta-like. Exopod three-segmented and ornamented as in figure 18 B. Endopod two-segmented, reaching proximal quarter of second exopodal segment; first segment small, about as long as wide and ornamented with outer and inner spinules; second segment ornamented with long spinules as shown in figure 18 B, and armed with one inner spine, one apical and one inner seta. Armature formula as in table 1. P 3 (figure 19 A): praecoxa ornamented with transverse row of spinules close to joint with coxa. The latter and basis as in P 2. Exopod as in P 2, except for two inner setae on third segment. Endopod as in P 2, armed with three inner, one apical and one outer seta. Armature formula as in table 1. P 4 (figure 19 B): praecoxa, coxa and basis as in P 3. Exopod as in P 3, except for armature formula of third segment. Endopod two-segmented, very small; first segment small, slightly wider than long; second segment ornamented with inner and outer slender spinules and armed with two apical setae. P 5 (figure 15 B): exopod and baseoendopod fused. Baseoendopodal lobe longer than exopod, ornamented with sets of inner spinules, and with spinules on distal part of ramus; armed with one outer, one apical and four inner setae; relative length of setae as in figure 15 B. Exopod ornamented with spinules as shown in figure 15 B, and armed with five setae in all, plus outer seta of basis. P 6 (figure 15 A) represented by median plate in anterior half of first genital somite, each vestigial leg represented by a short and thick seta. Copulatory pore in the middle of genital double segment. Male. Total body length measured from tip of rostrum to posterior margin of caudal rami, ranging from 310 to 540 M m (mean, 406 M m, N ~ 18; allotype, 540 M m). Habitus (figure 20 A, B), anal segment and caudal rami (figure 20 C, D) as in female dorsally, except for longer spinules along posterior margin of cephalic shield and free thoracic somites. Ventrally as in female except for genital somite and median spinules on fourth urosomite. Antennule (figure 21 B) six-segmented; subchirocer. Last segment with two acute teeth. Antenna, mandible, maxillule, maxilla and maxilliped (not illustrated) as in female. P 1 (figure 22 A), as in female except for dimorphic projection on inner distal corner of basis. P 2 (figure 22 B): praecoxa, coxa, basis and exopod as in female, except for somewhat stouter outer spines of male exopod. Endopod two-segmented; first segment small; second segment reaching about proximal third of second exopodal segment, ornamented with spinules along inner and outer margin, and armed with one outer dimorphic spine, one apical seta and two inner slender setae. P 3 (figure 23 A) as in female except for somewhat stouter outer spines of male exopod and for dimorphic three-segmented endopod. First endopodal segment small; second segment with inner distal apophysis reaching slightly beyond third endopodal segment; second segment barely reaching tip of first exopodal segment, with one inner and one apical seta comparatively shorter than their female homologues. P 4 (figure 23 B) as in female, except for somewhat stouter outer spines of exopod and relatively shorter setae of second endopodal segment. Both P 5 fused (figure 21 A); exopod and baseoendopod fused. Ornamented with spinules at base of outer seta of basis, at base of exopodal and endopodal setae. Exopod with four, endopod with three setae. P 6 (figure 21 A) represented by plate; without armature. Variability Male. Paratype (EMUCOP- 0698 - 16) was found with an aberrant outer seta of left P 2 ENP 2, and right P 2 ENP 2 possesses two inner setae; paratype (EMUCOP- 0698 - 20) possesses two setae only on right P 5 BENP; paratype (EMUCOP- 0698 - 15) possesses two inner setae on right P 2 ENP. Female. Paratype (EMUCOP- 0698 - 13) was found with inner setae of left P 5 BENP closer to adjoining seta than in right ramus; four female paratypes (EMUCOP- 0698 - 10, EMUCOP- 0698 - 12, EMUCOP- 0698 - 03, EMUCOP- 0698 - 06) possess one inner seta on left P 2 ENP 2 and two inner setae on right P 2 ENP 2; two paratypes (EMUCOP- 0698 - 05, EMUCOP- 0698 - 14) possess one inner seta on right P 2 ENP 2 and two setae on left P 2 ENP 2; paratype (EMUCOP- 0698 - 07) possesses an aberrant left P 1 EXP 3; paratype (EMUCOP- 0698 - 08) possesses six setae on P 5 EXP; paratype (EMUCOP- 0698 - 11) lacks inner seta on P 1 ENP 1 (see table 2).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE38FFFEFE17FD5FFE04FC7C.taxon	description	(figures 24 – 33)	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE38FFFEFE17FD5FFE04FC7C.taxon	materials_examined	Type material One female holotype (EMUCOP- 300392 - 37) preserved in alcohol and one dissected male allotype (EMUCOP- 300392 - 38) from Ensenada del Pabellón lagoon, five dissected female (EMUCOP- 300491 - 42, EMUCOP- 030192 - 69, EMUCOP- 300392 - 39, EMUCOP- 300392 - 40, EMUCOP- 300392 - 41) and four dissected male paratypes (EMUCOP- 300392 - 42, EMUCOP- 300392 - 43, EMUCOP- 230691 - 44, EMUCOP- 240691 - 41) from Ensenada del Pabellón lagoon, and one dissected female paratype (EMUCOP- 0100 - 02) from El Yugo estuary, seven female paratypes (EMUCOP- 0100 - 01) from El Yugo estuary preserved in alcohol, 27 female paratypes (EMUCOP- 300491 - 40) from Ensenada del Pabellón lagoon preserved in alcohol and one male paratype (EMUCOP- 300491 - 41) from Ensenada del Pabellón lagoon preserved in alcohol. April and June 1991 and January and March 1992 (Ensenada del Pabellón lagoon) and January 2000 (El Yugo estuary), coll. S. Gómez (Ensenada del Pabellón lagoon) and A. C. Puello-Cruz (El Yugo estuary). Type locality Ensenada del Pabellón lagoon, Sinaloa (24 ° 19 ’ – 24 ° 35 ’ N, 107 ° 28 ’ – 107 ° 45 ’ W). Other localities El Yugo estuary (23 ° 18 ’ 14 @ N, 106 ° 29 ’ W) and Urías system (23 ° 11 ’ 06 @ N, 106 ° 25 ’ 06 @ W), Mazatlán, Sinaloa.	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE38FFFEFE17FD5FFE04FC7C.taxon	etymology	Etymology The species was named after the region (Sinaloa State) where the species was found.	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE38FFFEFE17FD5FFE04FC7C.taxon	description	Description Female. Habitus (figure 24 A, B) tapering posteriorly; total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 510 to 750 M m (mean, 658 M m, N ~ 11; holotype, 560 M m). Rostrum set off, triangular, with pair of setules subapically. Cephalic shield ornamented with fine and long spinules along margin dorsolaterally (figure 24 A, B). Dorsal and lateral surface of free thoracic somites (P 2 – P 4 bearing somites) ornamented with transverse rows of minute spinules, with longitudinal row of small spinules close to posterior margin and with long spinules along posterior margin. Dorsal and lateral surface of first urosomite (P 5 bearing somite) with transverse rows of minute spinules, with row of small spinules close to posterior margin and with relatively longer spinules along posterior margin. First and second genital somites distinct dorsally and laterally, completely fused ventrally (figure 25 A); dorsal and lateral surface of first and second genital somite with transverse rows of minute spinules, with row of longer spinules along posterior margin of both somites, and with relatively longer spinules laterally, ventrally ornamented with spinules as shown in figure 25 A. Fourth and fifth urosomite as in second genital somite dorsally and laterally (figure 24 A, B), ventral surface ornamented with transverse rows of spinules as illustrated in figure 25 A. Dorsal surface of anal somite (figure 24 A, C) ornamented with proximal transverse rows of minute spinules, with transverse rows of stronger spinules, and with dorsolateral strong spinules close to joint with caudal rami; rounded anal operculum furnished with two transverse rows of strong spinules (distalmost spinules relatively smaller) close to posterior margin (figure 24 A, C). Caudal rami (figures 24 A, B, C, D, 25 A) about 1.5 times longer than wide; dorsal and ventral surface smooth, except for inner strong spinules close to site insertion of seta VII and close to posterior margin dorsally and ventrally; with seven elements in all. Antennule (figure 26 A) six-segmented; surface of segments smooth except for two rows of spinules on first segment. Armature formula, 1 - (1), 2 - (9), 3 - (5), 4 - (1 z [1 zae]), 5 - (1), 6 - (9 z [1 zae]). Antenna (figure 26 B) with small coxa ornamented with outer spinules and some inner spinules close to inner distal corner. Allobasis armed with two abexopodal setae. Free endopodal segment ornamented with inner strong spinules proximally and subdistally; with two lateral inner spines and a slender seta (the latter arrowed in figure 26 B), and five distal elements. Exopod one-segmented; about seven times longer than wide; armed with one lateral and one apical smooth setae. Mandible (figure 26 C) robust, ornamented with rows of spinules proximally; chewing edge with four strong teeth, four elements as in figure 26 C, one pyriform element and one lateral pinnate strong seta. Palp one-segmented, with two long setae unequal in length, and a small seta arising nearby. Maxillule (figure 26 D) robust; arthrite of praecoxa ornamented with few spinules, armed with a surface seta, seven distal spines and one lateral strong seta, the latter ornamented with long spinules. Coxa ornamented with some spinules and armed with two setae. Basis ornamented with some median spinules, and armed with three apical setae; exopod and endopod represented by three setae each. Maxilla (figure 26 E): syncoxa ornamented with minute spinules along inner margin; with two endites, each bearing three setae. Allobasis drawn into strong claw bearing one accompanying seta. Endopod represented by three setae. Maxilliped (figure 26 F) subchelate. Syncoxa ornamented by two rows of spinules and armed with a small seta on inner distal corner. Basis without armature and ornamented with an anterior and a posterior row of spinules, and with few spinules close to joint with endopod. The latter drawn into long and slender claw with one accompanying small seta. P 1 (figure 27 A): praecoxa ornamented with spinules close to joint with coxa. The latter ornamented with anterior transverse rows of small spinules, with strong spinules close to outer distal corner anteriorly, and with somewhat slender spinules posteriorly. Basis ornamented with median sets of spinules, and with stronger spinules at base of exopod, between rami and at base of inner spine of basis. Exopod three-segmented. Endopod two-segmented, second segment reaching far beyond third exopodal segment. Armature formula as in table 1. P 2 (figure 27 B): praecoxa as in P 1. Coxa ornamented with median row of small spinules and with strong spinules close to outer distal corner anteriorly, and with some spinules close to outer distal corner posteriorly. Basis ornamented with spinules between rami and at base of endopod, and with stronger spinules at base of exopod; outer element spine-like. Exopod three-segmented and ornamented as in figure 27 B. Endopod two-segmented, reaching distal quarter of second exopodal segment; first segment small, somewhat wider than long and ornamented with slender inner spinules; second segment ornamented with long spinules as shown in figure 27 B, and armed with one inner spine, one apical and one inner seta. Armature formula as in table 1. P 3 (figure 28 A): praecoxa, coxa and basis ornamented as in P 2. Exopod as in P 2. Endopod as in P 2 except for relatively shorter second segment. Armature formula as in table 1. P 4 (figure 28 B): praecoxa, coxa and basis as in P 2. Exopod as in P 2, except for armature formula of third segment (without inner seta). Endopod two-segmented, barely reaching distal third of first exopodal segment; first segment small, somewhat wider than long; second segment ornamented with inner and outer slender spinules and armed with two apical setae. P 5 (figure 25 B): exopod and baseoendopod fused. Baseoendopodal lobe longer than exopod, ornamented with sets of inner spinules, with spinules at base of apical seta and with some outer spinules; armed with one outer, one apical and four inner setae; relative length of setae as in figure 25 B. Exopod ornamented with spinules as shown in figure 25 B, and armed with five setae in all, plus outer seta of basis. P 6 (figure 25 A) represented by median plate in anterior half of first genital somite, each vestigial leg represented by a very small and slender seta. Copulatory pore in the middle of genital double somite. Male. Total body length measured from tip of rostrum to posterior margin of caudal rami, ranging from 570 to 610 M m (mean, 582 M m, N ~ 4; allotype, 570 M m). Habitus (figure 29 A, B), anal segment and caudal rami (figure 29 C, D) as in female dorsally, except for fewer rows of spinules on anal operculum. Ventrally as in female except for genital somite, ornamentation of third and fourth urosomites (see figure 30) which are coarser than in female; fifth urosomite seemingly without ornamentation ventrally. Antennule (figure 31 A) six-segmented; subchirocer. Last segment with two acute teeth. Antenna, mandible, maxillule, maxilla and maxilliped (not illustrated) as in female. P 1 (figure 32 A) as in female except for dimorphic projection on inner distal corner of basis. P 2 (figure 32 B) as in female except for relatively stouter outer spines of male exopod, dimorphic inner spine on male ENP 2 and relatively shorter apical setae on second endopodal segment. Exopod of P 3 (figure 33 A) as in female except for relatively stouter outer spines. Endopod three-segmented; second segment with inner distal apophysis reaching beyond third endopodal segment; second segment reaching little beyond first exopodal segment, with one inner and one apical seta relatively shorter than their female homologues. P 4 (figure 33 B) as in female, except for stouter outer spines of exopod and relatively shorter setae of second endopodal segment. Both P 5 fused (figure 31 B); exopod and baseoendopod fused. Exopod ornamented with transverse row of strong spinules; with four setae. Endopodal lobe ornamented with sets of spinules along outer and inner margin, and at base of apical setae; with three elements in all. P 6 (figure 30) represented by plate; without armature. Variability No variability was observed in males. Female. Paratype (EMUCOP- 300491 - 42) without inner seta and with two sets of spinules on left P 4 EXP 3, and with one inner seta and without spinules on right P 4 EXP 3; paratype (EMUCOP- 300392 - 40) with five setae / spines on P 1 EXP 3 (see table 2).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE0CFFE3FE10FB97FD4CFCD3.taxon	description	(figures 34 – 42)	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE0CFFE3FE10FB97FD4CFCD3.taxon	materials_examined	Type material One female holotype (USNM 1010505) and one male allotype (USNM 1010506) preserved in alcohol, five dissected female paratypes (EMUCOP- 1298 - 01, EMUCOP- 1298 - 04, EMUCOP- 1298 - 05, EMUCOP- 1298 - 06, EMUCOP- 1298 - 07), three female and five male paratypes (USNM 1010507), two female (EMUCOP- 1298 - 13) and four male paratypes (EMUCOP- 1298 - 14) preserved in alcohol, and seven dissected male paratypes (EMUCOP- 1298 - 02, EMUCOP- 1298 - 03, EMUCOP- 1298 - 08, EMUCOP- 1298 - 09, EMUCOP- 1298 - 10, EMUCOP- 1298 - 11, EMUCOP- 1298 - 12). December 1998 (Port Fourchon) and January 1999 (Cocodrie), coll. A. Rocha-Olivares, J. W. Fleeger and D. Foltz. Type locality Port Fourchon, Louisiana (29 ° 05.42 ’ N, 90 ° 05.8 ’ W). Other sites Cocodrie, Louisiana (29 ° 15.2 ’ N, 90 ° 39.8 ’ W); Graveline Bay, Mississippi (29 ° 15 ’ N, 91 ° 21 ’ W) (Carman et al., 2000).	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE0CFFE3FE10FB97FD4CFCD3.taxon	etymology	Etymology The species was named after the region (Port Fourchon) where the species was found.	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
AF0B87BCFE0CFFE3FE10FB97FD4CFCD3.taxon	description	Description Female. Habitus (figure 34) tapering posteriorly; total body length measured from tip of rostrum to posterior margin of caudal rami ranging from 560 to 725 M m (mean, 603 M m, N ~ 10; holotype, 567 M m). Rostrum set off, triangular, with pair of setules subapically. Cephalic shield ornamented with fine and long spinules along margin dorsolaterally. Dorsal and lateral surface of free thoracic somites (P 2 – P 4 bearing somites) ornamented with transverse rows of minute spinules, with longitudinal row of small spinules and with long spinules along posterior margin. Dorsal and lateral surface of first urosomite (P 5 bearing somite) with transverse rows of minute spinules, and with comparatively longer spinules along posterior margin. First and second genital somites distinct dorsally and laterally (figure 34 A, B), completely fused ventrally (figure 35 A); dorsal and lateral surface of first and second genital somite with transverse rows of minute spinules, with row of long spinules along posterior margin of both somites (spinules of second somite longer), and with relatively longer spinules laterally, ventrally ornamented with spinules as shown in figure 35 A. Fourth and fifth urosomite as in second genital somite dorsally, except for fewer transverse rows on fifth urosomite, ventral surface ornamented with transverse rows of spinules as illustrated in figure 35 A. Dorsal surface of anal somite (figure 34 A, C) ornamented with few transverse rows of minute spinules and with dorsolateral strong spinules close to joint with caudal rami; rounded anal operculum without ornamentation; with median row of strong spinules laterally (figure 34 D). Caudal rami (figure 34 A – D) about 1.5 times longer than wide; dorsal and ventral surface smooth, except for inner set of spinules close to insertion site of seta VII and close to posterior margin dorsally and ventrally; with seven elements in all. Antennule (figure 36 A) six-segmented; surface of segments smooth except for two rows of spinules on first segment. Armature formula, 1 - (1), 2 - (9), 3 - (6), 4 - (1 z [1 zae]), 5 - (1), 6 - (9 z [1 zae]). Antenna (figure 36 B) with small coxa ornamented with two sets of spinules. Allobasis armed with two strong abexopodal setae. Free endopodal segment ornamented with inner strong spinules proximally and subdistally, and armed with two lateral inner spines and one slender seta, and five distal elements. Exopod one-segmented; about five times longer than wide; ornamented with few spinules, and armed with one lateral and two apical smooth setae. Mandible (figure 36 C) robust, ornamented with rows of spinules proximally; chewing edge with four strong teeth, a multicuspidate tooth, a pyriform element and a lateral pinnate strong seta. Palp small, one-segmented, with two long setae unequal in length, with a small seta arising nearby. Maxillule (figure 36 D, E) robust; arthrite of praecoxa ornamented with few spinules, armed with a surface seta, seven distal spines and one lateral strong seta, the latter ornamented with long spinules. Coxa ornamented with some spinules and armed with two slender setae. Basis ornamented with some median spinules, and armed with three apical setae; exopod and endopod represented by three setae each. Maxilla (figure 36 F): syncoxa ornamented with minute spinules along inner and outer margin; with two endites, each bearing three setae. Allobasis drawn into strong claw bearing one accompanying seta. Endopod represented by three setae. Maxilliped (figure 36 G) subchelate. Syncoxa ornamented by anterior and posterior rows of spinules and armed with a small seta on inner distal corner. Basis without armature and ornamented with two longitudinal rows of spinules anteriorly and posteriorly, and with few spinules close to joint with endopod. The latter drawn into long and slender claw with one accompanying small seta. P 1 (figure 37 A): praecoxa ornamented with spinules close to joint with coxa. The latter ornamented with anterior transverse rows of spinules, with comparatively stronger spinules close to outer distal corner anteriorly, and with some spinules posteriorly. Basis ornamented with median row of spinules, and with stronger spinules at base of exopod, between rami and at base of inner spine of basis. Exopod three-segmented. Endopod two-segmented, second segment reaching beyond third exopodal segment. Armature formula as in table 1. P 2 (figure 37 B): coxa ornamented with median row of small spinules and with strong spinules close to outer distal corner anteriorly, and with some spinules close to outer distal corner posteriorly. Basis ornamented with median set of spinules, with spinules between rami and with comparatively stronger spinules at base of exopod; outer element spine-like. Exopod three-segmented and ornamented as in figure 37 B. Endopod two-segmented, reaching beyond the middle of second exopodal segment; first segment small, slightly wider than long and ornamented with some inner spinules; second segment ornamented with long spinules as shown in figure 37 B, and armed with three elements in all. Armature formula as in table 1. P 3 (figure 38 A): praecoxa as in P 1. Coxa ornamented with transverse rows of spinules, with strong spinules close to outer distal corner anteriorly and with slender and long spinules on outer distal corner posteriorly. Basis as in P 2 except for seta-like outer element. Exopod as in P 2. Endopod as in P 2 except for relatively shorter second segment reaching proximal third of second exopodal segment. Armature formula as in table 1. P 4 (figure 38 B): praecoxa, coxa and basis as in P 3. Exopod as in P 3, except for armature formula of third segment (without inner seta). Endopod two-segmented, barely reaching tip of first exopodal segment; first segment small, slightly wider than long; second segment ornamented with inner and outer slender spinules and armed with two apical setae. P 5 (figure 35 B): exopod and baseoendopod fused. Baseoendopodal lobe longer than exopod, ornamented with sets of inner and outer spinules, with spinules at base of apical seta; armed with one outer, one apical and four inner setae; relative length of setae as in figure 35 B. Exopod ornamented with spinules as shown in figure 35 B, and armed with five setae in all, plus outer seta of basis. P 6 (figure 35 A) represented by median plate in anterior half of first genital somite, each vestigial leg represented by a long and slender seta. Copulatory pore in the middle of genital double segment. Male. Total body length measured from tip of rostrum to posterior margin of caudal rami, ranging from 430 to 640 M m (mean, 539 M m, N ~ 15; allotype, 640 M m). Habitus (figure 39 A, B), anal segment and caudal rami (figure 39 C, D) as in female dorsally, except for some minute spinules along posterior margin of anal operculum. Ventrally as in female except for genital somite (figure 40 A). Ventral ornamentation of third, fourth and fifth urosomites (figure 40 A) coarser than in female. Antennule (figure 40 B) six-segmented; subchirocer. Last segment with two acute teeth. Antenna, mandible, maxillule, maxilla and maxilliped (not illustrated) as in female. P 1 (figure 41 A) as in female except for comparatively shorter endopod and dimorphic projection on inner distal corner of basis. P 2 (figure 41 B) as in female except for relatively stouter outer spines of male exopod, dimorphic inner spine on male ENP 2, and relatively shorter apical setae on second endopodal segment. Exopod of P 3 (figure 42 A) as in female except for relatively stronger outer spines. Endopod three-segmented; second segment with inner distal apophysis reaching beyond third endopodal segment, the latter reaching tip of first exopodal segment, with one inner and one apical seta relatively shorter than their female homologues. P 4 (figure 42 B) as in female, except for stouter outer spines of exopod and relatively shorter setae of second endopodal segment. Both P 5 fused (figure 40 B); exopod and baseoendopod fused. Exopod ornamented with spinules at base of outer seta of basis, at base of apical elements and along inner margin; with four setae. Endopodal lobe ornamented with spinules along outer and inner margin, and at base of apical setae; with three elements in all. P 6 (figure 40 A) represented by plate; without armature. Variability Male. Paratype (EMUCOP- 1298 - 03) possesses a reduced apical seta on right P 1 ENP 2, and an abnormally swollen apical element on right P 2 EXP 3; paratype (EMUCOP- 1298 - 09) possesses an aberrant right P 1 ENP 2, with two setae; the gap between innermost and adjoining seta of P 5 BENP is larger in right P 5 (EMUCOP- 1298 - 09); paratype (EMUCOP- 1298 - 10) possesses two and three setae on right and left P 5 BENP, respectively; paratype (EMUCOP- 1298 - 11) possesses one inner seta on left P 2 ENP 1 and four setae on P 5 BENP. Female. Paratype (EMUCOP- 1298 - 05) possesses a reduced inner seta on left P 4 ENP 2; paratype (EMUCOP- 1298 - 06) with three setae on both P 4 ENP 2; paratype (EMUCOP- 1298 - 07) possesses an abnormal inner distal seta on left P 1 ENP 2 and a reduced seta on right P 4 ENP 2.	en	Gómez, Samuel, Fleeger, John W., Rocha-Olivares, Axayácatl, Foltz, David (2004): Four new species of Cletocamptus Schmankewitsch, 1875, closely related to Cletocamptus deitersi (Richard, 1897) (Copepoda: Harpacticoida). Journal of Natural History 38 (21): 2669-2732, DOI: 10.1080/0022293031000156240, URL: http://dx.doi.org/10.1080/0022293031000156240
