taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818118/files/figure.png	https://doi.org/10.5281/zenodo.15818118	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818122/files/figure.png	https://doi.org/10.5281/zenodo.15818122	FIGURE 3. Habrophlebia abietis sp. nov., mouthparts. A, labrum. B, right mandible. C, maxilla. D, labium. E, maxillary palp. F, hemi hypopharynx. Scale bars: A, E, F: 100 µm; B, C, D: 200 µm.	FIGURE 3. Habrophlebia abietis sp. nov., mouthparts. A, labrum. B, right mandible. C, maxilla. D, labium. E, maxillary palp. F, hemi hypopharynx. Scale bars: A, E, F: 100 µm; B, C, D: 200 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818124/files/figure.png	https://doi.org/10.5281/zenodo.15818124	FIGURE 4. Habrophlebia abietis sp. nov., thorax.A, fore femur. B, hind femur. C–D, tarsal claw. E, setae on dorsal face of hind femur. Scale bars: A, B: 200 µm, C, E: 50 µm.	FIGURE 4. Habrophlebia abietis sp. nov., thorax.A, fore femur. B, hind femur. C–D, tarsal claw. E, setae on dorsal face of hind femur. Scale bars: A, B: 200 µm, C, E: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818128/files/figure.png	https://doi.org/10.5281/zenodo.15818128	FIGURE 5. Habrophlebia abietis sp. nov., abdomen. A, gill I. B, gill III. C, gill VII. D–H, posterior margin of abdominal tergites IX, VIII, VII, VI, and V respectively. Scale bar: 50 µm.	FIGURE 5. Habrophlebia abietis sp. nov., abdomen. A, gill I. B, gill III. C, gill VII. D–H, posterior margin of abdominal tergites IX, VIII, VII, VI, and V respectively. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818136/files/figure.png	https://doi.org/10.5281/zenodo.15818136	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF84D625F8C1FA48FD5C116F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818142/files/figure.png	https://doi.org/10.5281/zenodo.15818142	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818118/files/figure.png	https://doi.org/10.5281/zenodo.15818118	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818130/files/figure.png	https://doi.org/10.5281/zenodo.15818130	FIGURE 6. Habrophlebia linae sp. nov., mouthparts. A, labrum. B, right mandible. C, maxilla. D, labium. E, maxillary palp. F, hemi hypopharynx. Scale bars: A, E, F: 100 µm; B, C, D: 200 µm.	FIGURE 6. Habrophlebia linae sp. nov., mouthparts. A, labrum. B, right mandible. C, maxilla. D, labium. E, maxillary palp. F, hemi hypopharynx. Scale bars: A, E, F: 100 µm; B, C, D: 200 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818132/files/figure.png	https://doi.org/10.5281/zenodo.15818132	FIGURE 7. Habrophlebia linae sp. nov., thorax. A, fore femur. B, hind femur. C–D, tarsal claw. E, setae on dorsal face of hind femur. Scale bars: A, B: 200 µm; C, E: 50 µm.	FIGURE 7. Habrophlebia linae sp. nov., thorax. A, fore femur. B, hind femur. C–D, tarsal claw. E, setae on dorsal face of hind femur. Scale bars: A, B: 200 µm; C, E: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818134/files/figure.png	https://doi.org/10.5281/zenodo.15818134	FIGURE 8. Habrophlebia linae sp. nov., abdomen. A, gill I. B, gill III. C, gill V. D–G, posterior margin of abdominal tergites IX, VIII, VII, and VI respectively. Scale bar: 100 µm.	FIGURE 8. Habrophlebia linae sp. nov., abdomen. A, gill I. B, gill III. C, gill V. D–G, posterior margin of abdominal tergites IX, VIII, VII, and VI respectively. Scale bar: 100 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818136/files/figure.png	https://doi.org/10.5281/zenodo.15818136	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF89D622F8C1FB65FE2C17EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818142/files/figure.png	https://doi.org/10.5281/zenodo.15818142	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818118/files/figure.png	https://doi.org/10.5281/zenodo.15818118	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	FIGURE 2. Habrophlebia spp., nymphal habitus. A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia vaillantorum Thomas, 1986. Scale bars: 1 mm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818138/files/figure.png	https://doi.org/10.5281/zenodo.15818138	FIGURE 10. Habrophlebia vaillantorum Thomas, 1986. A, claw. B, setae on dorsal face of hind femur. C–D, posterior margin of abdominal tergites IX and VIII respectively. Scale bars: 50 µm.	FIGURE 10. Habrophlebia vaillantorum Thomas, 1986. A, claw. B, setae on dorsal face of hind femur. C–D, posterior margin of abdominal tergites IX and VIII respectively. Scale bars: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818142/files/figure.png	https://doi.org/10.5281/zenodo.15818142	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818140/files/figure.png	https://doi.org/10.5281/zenodo.15818140	FIGURE 11. Habrophlebia spp., setae on dorsal face of hind femur. A, Habrophlebia fusca (Curtis, 1834). B, Habrophlebia lauta McLachlan, 1884. C, Habrophlebia djurdjurensis Kechemir, Sartori & Lounaci, 2020. D, Habrophlebia hassainae Benhadji & Sartori, 2018. Scale bar: 50 µm.	FIGURE 11. Habrophlebia spp., setae on dorsal face of hind femur. A, Habrophlebia fusca (Curtis, 1834). B, Habrophlebia lauta McLachlan, 1884. C, Habrophlebia djurdjurensis Kechemir, Sartori & Lounaci, 2020. D, Habrophlebia hassainae Benhadji & Sartori, 2018. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818136/files/figure.png	https://doi.org/10.5281/zenodo.15818136	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	FIGURE 9. Habrophlebia spp., egg structure in optical microscopy (A, B, C, E) and scanning electronic microscopy (D, F, G). A–B, Habrophlebia abietis sp. nov. C–D, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. E-G, Habrophlebia linae sp. nov. from Madissouka (E, F) and oued Stah (G).	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818142/files/figure.png	https://doi.org/10.5281/zenodo.15818142	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	FIGURE 12. Habrophlebia spp., setae on hind tibia (outer margin up, inner margin down). A, Habrophlebia abietis sp. nov. B, Habrophlebia linae sp. nov. C, Habrophlebia dakkii El Alami, Sartori & Vuataz, 2023. D, Habrophlebia vaillantorum Thomas, 1986. E, Habrophlebia eldae Jacob & Sartori, 1984. F, Habrophlebia lauta McLachlan, 1884. Scale bar: 50 µm.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15818144/files/figure.png	https://doi.org/10.5281/zenodo.15818144	FIGURE 13. Bayesian (BEAST) maximum clade credibility COI tree of the genus Habrophlebia in the West Palearctic. Branch tips labelled with GBIF codes indicate newly sequenced specimens; the other codes correspond to sequences obtained from various sources (see Materials and methods). Brown vertical boxes indicate species delimitation hypothesis (MOTUs) based on the ASAP, mPTP, and GMYC methods. Species names (where available) and the country (-region) of origin are shown adjacent to their respective clades, with the two newly described species and their associated GBIF codes highlighted in bold. For specimens from the Rif region (the two upper clades), squares to the right represent five key morphological characters (claw: number of denticles on tarsal claw; tergite: shape and size of posterior spines on abdominal segment VII; tibia: shape and size of bristles on hind tibia; gill: number of filaments on upper and lower lamellae of gills II–VI; egg: egg chorionic structure), colorcoded to reflect species affiliation (light green: Habrophlebia abietis sp. nov.; dark green: H. linae sp. nov.; blue: H. dakkii). The “composite clade” (highlighted in red) comprises specimens of H. abietis sp. nov., H. linae sp. nov., H. dakkii, and their morphologically intermediate forms (i.e., putative hybrids, represented by squares of different colors on the same line). Circles on branches indicate Bayesian posterior probabilities> 0.9. Outgroup branches are presented in grey with their corresponding labels and genus name.	FIGURE 13. Bayesian (BEAST) maximum clade credibility COI tree of the genus Habrophlebia in the West Palearctic. Branch tips labelled with GBIF codes indicate newly sequenced specimens; the other codes correspond to sequences obtained from various sources (see Materials and methods). Brown vertical boxes indicate species delimitation hypothesis (MOTUs) based on the ASAP, mPTP, and GMYC methods. Species names (where available) and the country (-region) of origin are shown adjacent to their respective clades, with the two newly described species and their associated GBIF codes highlighted in bold. For specimens from the Rif region (the two upper clades), squares to the right represent five key morphological characters (claw: number of denticles on tarsal claw; tergite: shape and size of posterior spines on abdominal segment VII; tibia: shape and size of bristles on hind tibia; gill: number of filaments on upper and lower lamellae of gills II–VI; egg: egg chorionic structure), colorcoded to reflect species affiliation (light green: Habrophlebia abietis sp. nov.; dark green: H. linae sp. nov.; blue: H. dakkii). The “composite clade” (highlighted in red) comprises specimens of H. abietis sp. nov., H. linae sp. nov., H. dakkii, and their morphologically intermediate forms (i.e., putative hybrids, represented by squares of different colors on the same line). Circles on branches indicate Bayesian posterior probabilities> 0.9. Outgroup branches are presented in grey with their corresponding labels and genus name.	2025-06-03	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel		Zenodo	biologists	Alami, Majida El;Vuataz, Laurent;Yaagoubi, Sara El;Sartori, Michel			
