taxonID	type	description	language	source
B6408157FF84D625F8C1FA48FD5C116F.taxon	description	https: // zoobank. org / E 4401307 - 9441 - 4 E 04 - 8 AED-BB 92312185 FD Figs 2 A, 3 – 5, 9 A – B, 12 A	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF84D625F8C1FA48FD5C116F.taxon	materials_examined	Material examined. Holotype: one nymph in ethanol (GBIFCH 00892900), MOROCCO, Chefchaouen Province, Talassemtane National Park, S 7, Oued Afeska, Loc. Afeska; 35 ° 09 ' 53 '' N, 5 ° 11 ' 12 '' W; alt. 1237 m; 30. VII. 2022, S. El Yaagoubi leg., MZL. Paratypes. MOROCCO, Chefchaouen Province, same data as holotype, 3 nymphs in ethanol (GBIFCH 00892901), 1 nymph on slide (GBIFCH 01314414), MZL, and 3 nymphs in ethanol, 2 nymphs on slide, LESCB; same locality as holotype, 2. IV. 2021, 5 nymphs in ethanol (GBIFCH 00892902), 1 nymph on slide (GBIFCH 00970949), MZL, and 3 nymphs in ethanol; 2 nymphs on slide, LESCB, S. El Yaagoubi leg.; same locality as holotype, 4. VI. 2021, 2 nymphs in ethanol, 1 nymphs on slide, S. El Yaagoubi, leg. LESCB; same locality as holotype, 2. VI. 2022, 11 nymphs in ethanol (GBIFCH 00892903), 2 nymphs on slide (GBIFCH 01314449, GBIFCH 01118435), MZL, and 12 nymphs in ethanol, 1 nymph on slide, S. El Yaagoubi leg., LESCB; same locality as holotype, 4. VII. 2022, 2 nymphs in ethanol (GBIFCH 00892904), 1 nymph on slide (GBIFCH 01144260), S. El Yaagoubi leg., MZL; same locality as holotype, 9. VI. 2024, 2 nymphs in ethanol (GBIFCH 00892905), S. El Yaagoubi leg., MZL; same locality as holotype, 22. IV. 2024, 14 nymphs in ethanol (GBIFCH 00892906), 2 nymphs on slide (GBIFCH 01550988 – GBIFCH 01550989), P. Derleth & M. Sartori leg., MZL.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF84D625F8C1FA48FD5C116F.taxon	description	Description. Nymph. Coloration and dimensions. Body length of final instar, excluding caudal filaments, 5 to 6 mm for male and 5.5 to 6.5 mm for female. Cerci as long as body length. General dark brown coloration with greyish to light brown markings mainly on abdominal terga. The whole cuticle is shagreened. Head. General coloration light brown; paler area around the compound eyes and lateral ocelli; between ocelli, a dark brown mark not reaching the clypeus distally, and extending laterally in front of the compound eyes in males; vertex sutures light brownish, barely visible (Fig. 2 A). Upper portion of male compound eyes red brownish. Antenna with pedicel brownish, scape and filament yellowish. Labrum rectangular (Fig. 3 A), 1.7 – 1.9 x wider than long; dorsal surface covered distally with scattered stout setae, proximally with long and thin setae; anterior margin with a row of stout, long and spatulate setae medially; emargination narrow, U-shaped with ca eight flat denticles; ventral surface with two bundles of stout, long setae medially. Mandibles similar to other Habrophlebia species (Fig. 3 B). 7 – 10 long and thin setae below the mola of the right mandible. Maxilla stocky, subapical row of 7 – 8 pectinate setae (Fig. 3 C); maxillary palp with three segments, segment 1 and 2 subequal in length, and much longer than segment 3; segment 3 triangular, ca 1.6 × longer than wide at base; all setae on palp segments stout and entire (Fig. 3 E). Hypopharynx with highly developed superlinguae terminated by a membranous digitation (Fig. 3 F). Labium (Fig. 3 D) with rhomboid glossae, outer margin and apex covered by short, broad setae; paraglossae enlarged laterally, covered with fine and long setae on the dorsal surface; with long, thick setae on the outer margin; labial palp with three segments; inner margin of segment 1 highly dilated near the middle, about 1.2 × longer than the maximum width, segments 2 as long as segment 3, ca 0.9 × length of segment 1; segment 3 ca 1.8 × longer than wide at base, conical shape, with 6 – 8 long and stout setae on dorsal face and inner margin with one row of longer and robust setae. Thorax. Pro- and mesonotum medium brown, with greyish or brownish maculae, two light brown oval or rounded large maculae on the prothorax (Fig 2 A). Legs light to medium brown; dorsal surface of femora and tarsi with greyish maculae. Fore legs (Fig. 4 A) femora elongated, ca 2.4 × longer than wide, dorsal surface covered with long, entire and pointed setae; fore tibiae subequal in length to femora, outer margin with few thin and long setae, inner margin with several rows of long stout and feathered setae; tarsi 0.5 × length of tibiae, outer margin with long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca 2.7 × longer than wide, dorsal surface of femora with more numerous and slightly longer stout, pointed and feathered setae; tibiae and femora of subequal length; tarsi 0.5 × length of tibiae. Hind legs (Fig. 4 B) femora 3 × longer than wide, dorsal surface covered with stout, pointed and feathered setae (Fig. 4 E); ventral surface with few stout setae; hind tibiae as long as hind femora, outer margin with very few long, stout, pointed setae; inner margin with stout, long, pointed, scattered setae; tarsi 0.4 × length of tibiae, outer margin without or with 1 – 2 long and thin setae, inner margin with long, stout, pointed setae. Claws (Fig. 4 C) of all legs slightly hooked, with 11 to 12 denticles, the last four apically reduced in size (Fig. 4 D). Abdomen. Tergites greyish to dark brown (Fig. 2 A). Tergites I – VIII with a dark brown band posteriorly tergites IX – X uniformly medium to light brown. Sternum I entirely greyish brown; sternites II−VIII greyish to lighter brown with posterior bands brown, and two parasagittal nervous ganglia dark brown; sternite IX greyish. Posterolateral expansions only on segments VIII and IX. Ornamentation of the tergites (Figs. 5 D – H): posterior margin of tergite IX with well-developed narrow and pointed spines, ca 2 – 3 times longer than wide, bordered with submarginal microdenticles; tergites VII−VIII with much narrower and shorter spines, bordered with submarginal microdenticles; tergites V−VI with needle-shaped spines, tergite III – IV with minute spines, not longer than the submarginal microdenticles; tergites I−II with barely visible spines. Gills present on segments I−VII; all gills long and large; first gill (Fig. 5 A) with ventral lamella bearing 3 – 4 filaments, dorsal lamella with 3 – 4 filaments, gills II−VII (Figs. 5 B – C) with 3 – 4 and 4 – 7 filaments on the ventral and dorsal lamella respectively. Cerci and terminal filament yellowish brown, medium brown in mature nymphs (Fig. 2 A). Eggs ornamented with small depressions surrounded by reticulated ribs shaped like a small-meshed net, distributed and arranged along the whole of the chorionic surface. These ribs are made up of microtubules joined together to form hexagonal, pentagonal or irregular rosette-shaped units (Figs. 9 A – B). Imago. Unknown.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF84D625F8C1FA48FD5C116F.taxon	etymology	Etymology This new species is named abietis to remember it inhabits a region where the endemic and endangered tree Abies marocana, the Moroccan fir or Rif fir, is still present.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF89D622F8C1FB65FE2C17EB.taxon	description	https: // zoobank. org / 10 E 7 AF 77 - C 3 BC- 4 BDE-B 8 A 2 - 0 FDC 4 D 8 CA 38 E Figs 2 B, 6 – 8, 9 E – G, 12 B	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF89D622F8C1FB65FE2C17EB.taxon	materials_examined	Material examined. Holotype: one nymph on slide (GBIFCH 01314432), MOROCCO, Chefchaouen Province, Talassemtane National Park, S 6, Oued Madissouka; 35 ° 10 ' 27 '' N, 5 ° 08 ' 18 '' W; alt. 1377 m; 4. VII. 2021; M. El Alami team leg.; MZL. Paratypes. MOROCCO, Chefchaouen Province, same data as holotype, 2 nymphs in ethanol (GBIFCH 00892907), 1 nymph on slide (GBIFCH 01118421), MZL, and 10 nymphs in ethanol, 2 nymphs on slide, LESCB; same locality as holotype, 10. VI. 2021, 3 nymphs in ethanol (GBIFCH 00892908), 2 nymphs on slide (GBIFCH 01144263, GBIFCH 01314491), S. El Yaagoubi leg., MZL; same locality as holotype, 2. VI. 2022, one nymph in ethanol (GBIFCH 00892909), one nymph on slide (GBIFCH 01550999), S. El Yaagoubi leg., MZL; same locality as holotype, 6. V. 2005, 10 nymphs in ethanol (GBIFCH 00892910), one nymph on slide (GBIFCH 01118434), M. El Alami leg., MZL. Larache Province, S 21, Oued Stah, Loc. Tazroute; 35 ° 16 ' 34.47 " N, 5 ° 31 ' 53.08 " W; alt. 778 m; 2. V. 2021, 2 nymphs in ethanol (GBIFCH 01523589); 1 nymph on slide (GBIFCH 01118420); S. El Yaagoubi leg., MZL. 3 nymphs in ethanol, 2 nymphs on slide, S. El Yaagoubi leg. LESCB; same locality, 24. IV. 2024, 11 nymphs in ethanol (GBIFCH 01523590), one nymph on slide (GBIFCH 01550991), S. El Yaagoubi, Y. El Kourchi & M. Sartori leg., MZL.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF89D622F8C1FB65FE2C17EB.taxon	description	Description. Nymph. Coloration. Nymph. Body length of final instar, excluding caudal filaments, 4 to 6 mm for male and 6 to 8 mm for female. Cerci as long as body length. General dark brown coloration with light brown and yellowish markings mainly on abdominal terga. The whole cuticle is shagreened. Head. General coloration yellowish to light brown; paler area round compound eyes and lateral ocelli; dark brown patch between ocelli extending laterally in front of compound eyes but not reaching clypeus distally. Upper part of male eyes reddish brown. Antenna with light brownish pedicel, scape and filament whitish to yellowish (Fig. 2 B). Labrum rectangular (Fig. 6 A), ca 2 × wider than long; dorsal surface covered distally with scattered stout setae, proximally with long and thin setae; anterior margin with a row of stout, long and spatulate setae medially; U-shaped emargination with four flat, irregularly sized denticles; ventral surface with two tufts of strong and long setae in the median area. Mandibles similar to other Habrophlebia species (Fig. 6 B). 6 – 7 long and thin setae below the mola of the right mandible. Maxilla stocky, subapical row of 6 to 8 pectinate setae (Fig. 6 C); maxillary palp with three segments, segment 1 and 2 subequal in length, and longer than segment 3; segment 3 triangular, 1.4 to 1.7 × longer than wide at base; all setae on palp segments stout and entire (Fig. 6 E). Hypopharynx with very robust superlinguae terminated by a fairly long membranous digitation (Fig. 6 F). Labium (Fig. 6 D) with rhomboid glossae, outer margin and apex with short, broad setae; paraglossae laterally enlarged, covered with fine and long setae on the dorsal surface; with long, thick setae on the outer margin; labial palp with three segments, inner margin of segment 1 strongly expanded near the middle, ca 1.2 – 1.3 × longer than maximum width, segments 2 as long as segment 3, ca 0.7 × length of segment 1; segment 3 ca 1.7 × longer than broad at base, conical shape and inner margin with a row of longer and robust setae. Thorax. Pro- and mesonotum yellowish brown, with greyish brown maculae, particularly on lateral margins and 3 light brown oval or rounded large maculae on prothorax (Fig. 2 B). Legs whitish to light brown; dorsal surface of femora washed with greyish macula; tarsi and tibiae generally lighter. Fore legs (Fig. 7 A) femora elongated, ca 2.6 × longer than wide, dorsal surface covered with long, entire and pointed setae; fore tibiae subequal in length to femora, outer margin without setae, inner margin with several rows of long stout and feathered setae especially near apex; tarsi 0.5 × length of tibiae, outer margin with few long and thin setae, inner margin with long and pointed entire setae. Middle legs similar to fore legs, femora ca 2.9 × longer than wide, dorsal surface of femora with numerous and long entire and pointed setae; tibiae and femora of subequal length; tarsi 0.4 to 0.5 × length of tibiae. Hind legs (Fig. 7 B) with femora ca 3.2 × longer than wide, dorsal surface covered with stout, pointed and feathered setae (Fig. 7 E); ventral surface with few stout setae; hind tibiae as long as hind femora, outer margin with long, stout, pointed setae; inner margin with stout, long, pointed setae, entire and feathered; tarsi 0.4 – 0.5 × length of tibiae, outer margin naked or with 1 – 2 long and thin setae, inner margin with long, stout, pointed setae. Claw with 13 to 15 long, thin, pointed denticles (Fig. 7 C), the last three apically reduced in size (Fig. 7 D). Abdomen. Tergites grey to dark brownish with characteristic light patterns (Fig. 2 B). Tergites I – II dark brown, and with an oblong sagittal macula, leaving two lateral light areas; posterior marking less visible on tergites III – IV, more conspicuous on tergites V – IX; posterior light mark sometimes absent on tergites VIII – IX; tergite X light brown. Sternite I greyish brown; sternites II−VIII yellowish to light brown with lateral and posterior brown bands, and two parasagittal nervous ganglia greyish brown; sternites IX – X lighter brown. Posterolateral expansions only on segments VIII and IX. Posterior margin of tergite X with well-developed narrow and pointed spines, ca 3 – 4 times longer than wide at base; tergite IX with narrower and shorter spines; regular shape and size of posterior spines on tergites VII to V, with regular spines decreasing in size, bordered with submarginal microdenticles; tergites IV – I with barely visible spines, slightly more developed than the microdenticles bordering them (Figs. 8 D – G). Gills present on segments I – VII; all gills long and large; first gill (Fig. 8 A) with ventral lamella bearing 3 or 4 filaments, dorsal lamella with 3 – 4 filaments, gills II−VII (Figs. 8 B – C) with 3 – 4 and 3 – 7 filaments on the ventral and dorsal lamella, respectively. Cerci and terminal filament yellowish brown, light brown in mature nymphs. Eggs. General shape ovoid; chorionic surface uniformly decorated with sinuous, non-punctuated longitudinal ribs, variable in length; some connected in the equatorial or subequatorial region, anastomosed and giving rise to a Y-like configuration and others entire; micropyle in subequatorial area with margins slightly elevated (Figs. 9 E – G). Imago. Unknown.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF89D622F8C1FB65FE2C17EB.taxon	etymology	Etymology This new species is dedicated to the first author’s daughter, Lina, for her encouragement and assistance with sampling during some field trips.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	description	Figs 2 C, 10, 12 D	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	materials_examined	Material examined. 6 nymphs in ethanol (GBIFCH 00892912), 1 nymph on slide (GBIFCH 01118423), MOROCCO, High Atlas, Al Haouz Province, Assif n’Ouarzane, 31 ° 06 ' 41 '' N, 7 ° 58 ' 34 '' W, 2190 m., 16. VII. 2005, A. Abdaoui leg., MZL; 6 nymphs in ethanol (GBIFCH 01133089), same locality, 16. III. 2005, A. Abdaoui leg., MZL; 9 nymphs in ethanol (GBIFCHGBIFCH 00892911), 2 nymphs on slide (GBIFCH 01118427 – GBIFCH 01118428), MOROCCO, High Atlas, Al Haouz Province, oued Tagdet, N’fis, 30 ° 59 ' 19 '' N, 8 ° 02 ' 34 '' W, 2010 m., 3. III. 2022, M. El Battahi leg., MZL.	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED622F8C1FCF4FA2213F0.taxon	description	Complements to the nymph description In contrast to the description by Bauernfeind & Soldán (2012), the setae on the upper face of hind femora are pointed and fringed, not pointed and entire (Fig. 10 B). Most of Habrophlebia species possess pointed and fringed setae on upper face of hind femora (Figs. 11 C – D), except for H. lauta, where they are pointed and entire (Fig. 11 B), and H. fusca, where they are entire and truncate (Fig. 11 A). Thus, the character is not discriminant for most species. The claw is described as possessing the most distal denticles reduced in size and is illustrated as such (Thomas et al. 1999, fig. 12 a – d). In fact, these denticles are diminished in width rather than length (Fig. 10 A), and, as such, are like those of H. abietis sp. nov., which also possess this character. The spines on the posterior margin of tergites differ somewhat from those by Thomas et al. (1999, fig. 14); tergite IX possesses more developed spines, making it similar to all other representatives of Habrophlebia (Fig. 10 C), although these spines are obtuse and less pointed than in most other species. Spines on tergite VII are reduced and needle-shaped, as mentioned by Thomas et al. (1999). Those on tergite VIII are small and pointed (Fig. 10 D).	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	description	Figs 9 C – D, 12 C	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	materials_examined	Material examined (not type material). 1 nymph in ethanol (GBIFCH 01123077), 2 nymphs on slide (GBIFCH 01118417 – GBIFCH 01314688) MOROCCO, Tetouan Province, oued Assemlil, 5 ° 15 ' 33 '' N, 5 ° 25 ' 58 '' W, 1068 m ,, 20. VI. 2022, S El Yaagoubi leg.; 9 nymphs in ethanol GBIFCH 01123078), 2 nymphs on slide (GBIFCH 01550986 – GBIFCH 01550987), Tetouan Province, oued Hamma, 35 ° 23 ' 23 '' N, 5 ° 29 ' 58 '' W, 200 m., 24. IV. 2024, S. El Yaagoubi, Y. El Kourchi & M. Sartori leg.; 19 nymphs in ethanol (GBIFCH 01123079), 1 nymph on slide (GBIFCH 01550993), Tetouan Province, oued Taida, 35 ° 22 ' 06 '' N, 5 ° 32 ' 15 '' W, 510 m., 24. IV. 2024, S. El Yaagoubi, Y. El Kourchi & M. Sartori leg.; 9 nymphs in ethanol (GBIFCH 01123080), 1 nymph on slide (GBIFCH 01550997), Tetouan Province, oued Tisgris, 35 ° 22 ' 05 '' N, 5 ° 32 ' 04 '' W, 500 m., 24. IV. 2024, S. El Yaagoubi, Y. El Kourchi & M. Sartori leg. (all MZL).	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
B6408157FF8ED623F8C1F8CDFBCC16EF.taxon	description	Additions to the egg description In the original description, only a light microscope picture was presented (Fig. 9 C and El Alami et al. 2023, fig. 8). The real structure of the egg, revealed by SEM (Fig. 9 D), shows that the chorion is not “ regularly decorated by numerous and small granulations ”, but by a dense mesh of adhesive layer, leaving only small “ holes ”, which were interpreted under light microscopy as “ granulations ”. Morphologically intermediate forms Based on five key diagnostic characters (number and size of denticles on claw; shape and size of posterior spines on abdominal segment VII; shape and size of bristles on hind tibia; number of filaments on upper and lower lamellae of gills II – VI; egg chorionic structure), an important proportion of the examined specimens from the Rif region presented intermediate morphological characteristics between Habrophlebia dakkii and H. abietis sp. nov., then H. dakkii and H. linae sp. nov., as well as H. abietis sp. nov. and H. linae sp. nov. (Fig. 13).	en	Alami, Majida El, Vuataz, Laurent, Yaagoubi, Sara El, Sartori, Michel (2025): Integrative taxonomy of the genus Habrophlebia Eaton, 1881 in Morocco (Ephemeroptera, Leptophlebiidae): new species, redescriptions, and morphological-mitochondrial discrepancies. Zootaxa 5642 (5): 401-426, DOI: 10.11646/zootaxa.5642.5.1, URL: https://doi.org/10.11646/zootaxa.5642.5.1
