taxonID	type	description	language	source
6D3AC134CDA052A4B049AD1273541281.taxon	description	Figs 2 I, 19, 20	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
6D3AC134CDA052A4B049AD1273541281.taxon	description	Description. Body (Fig. 2 I): Small, slender and sub-cylindrical, males up to 4.6 mm cl, females up to 4.6 mm cl. Sixth abdominal somite 0.46 – 0.50 times as long as carapace. Rostrum (Fig. 19 A): Straight, reaching to not beyond eye to end of second segment of antennular peduncle; 0.2 – 0.5 of cl; armed dorsally with 3 – 11 teeth, including 2 – 3 on carapace posterior to orbital margin, ventrally with 0 – 2 teeth; rostral formula 2 – 3 + 1 – 8 / 0 – 2; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 19 A): Well developed, on short ocular peduncle, cornea globular. Carapace (Fig. 19 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 19 B): Stylocerite reaching 1.00 – 1.19 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.30 – 1.84 times as long as second segment, second segment 1.23 – 1.34 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 19 C): Scaphocerite 2.5 – 3.0 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 19 G): Palp of first maxilliped broadly triangular, ending with no finger-like projection. Third maxilliped (Fig. 20 B): Basal segment 1.25 times as long as penultimate segment; penultimate segment as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 6 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the beginning of second segment, distal margin with long plumose setae. First pereiopod (Fig. 20 C): Chela 1.84 – 2.50 times as long as high, 1.23 – 1.41 times as long as carpus; movable finger 2.70 – 3.45 times as long as wide and 0.85 – 1.01 times as long as palm, setal brushes well developed; carpus 1.60 – 2.02 times as long as wide, slightly excavated distally; merus 1.00 – 1.33 times as long as carpus. Second pereiopod (Fig. 20 D): Chela 2.61 – 3.27 times as long as high, 0.70 – 0.86 times as long as carpus; movable finger 3.82 – 4.08 times as long as wide and 1.31 – 1.76 times as long as palm, setal brushes well-developed; carpus 4.97 – 6.15 times as long as wide, slightly excavated distally; merus 0.81 – 1.01 times as long as carpus. Third pereiopod (Fig. 20 E): Dactylus 3.58 – 3.93 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 4 – 6 spiniform setae; propodus 3.43 – 4.04 times as long as dactylus, 10.48 – 11.67 times as long as wide; carpus 0.68 – 0.96 times as long as propodus; merus 1.73 – 1.88 times as long as carpus, with 2 – 3 spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 20 F): Dactylus 3.44 – 4.01 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 29 – 52 spiniform setae; propodus 3.32 – 4.42 times as long as dactylus, 10.75 – 13.13 times as long as wide; carpus 0.33 – 0.61 times as long as propodus; merus 1.44 – 1.61 times as long as carpus, with 2 – 3 spiniform setae on posterior margin. First pleopod (Fig. 20 G): Endopod reaching 0.44 – 0.57 times as long as exopod, reniform leaf-shaped, terminal width subequal to basal width, 2.70 times as long as distally wide, inner margin relatively straight, outer margin slightly convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well developed, arising from distal 0.2 of endopod, reaching slightly beyond the end of endopod, distally with cincinuli. Second pleopod (Fig. 20 H): Appendix masculina rod-shaped, reaching 0.63 – 0.72 length of exopod; some sparse long spines basal margin with 4 short spiniform setae, and inner margin and tip with robust spiniform setae; appendix interna well-developed, reaching 0.50 – 0.56 length of appendix masculina, distally with cincinuli. Telson (Fig. 20 I): With 6 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin triangular, with 3 – 4 pairs of intermedial spiniform setae, lateral spiniform setae slightly longer, intermediate spiniform setae subequal. Exopodite of the uropod (Fig. 20 J) bears a series of 18 – 22 movable spinules along diaresis.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
6D3AC134CDA052A4B049AD1273541281.taxon	etymology	Etymology. The name is derived from its pattern, and brevilineata means having short stripes.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
6D3AC134CDA052A4B049AD1273541281.taxon	distribution	Distribution. Only occurring in the type locality, Qionghai City, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
DCE005D842CD522488F6083314BEB6E2.taxon	description	Figs 2 B, 5, 6	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
DCE005D842CD522488F6083314BEB6E2.taxon	discussion	Caridina clinata Cai, 2014: 207 – 231, Figs 10, 11.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
DCE005D842CD522488F6083314BEB6E2.taxon	description	Description. Body (Fig. 2 B): Males up to 4.3 mm cl, females up to 4.9 mm cl. Sixth abdominal somite 0.49 – 0.55 times as long as carapace. Rostrum (Fig. 5 A): Short, reaching to the end of the second segment of antennular peduncle, sloping ventral anteriorly; 0.4 – 0.5 of cl; armed dorsally with 11 – 16 teeth, including 3 – 5 on carapace posterior to orbital margin, ventrally with 1 – 3 teeth; rostral formula 3 – 5 + 8 – 11 / 1 – 3; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 5 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 5 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 5 B): Stylocerite reaching 0.75 – 0.89 times as long as basal segment of antennular peduncle; basal segment shorter or as long as combined length of second and third segments, 1.61 – 1.92 times as long as second segment; second segment 1.16 – 1.31 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 5 C): Scaphocerite 3.3 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 5 G): Palp of first maxilliped broadly triangular, ending in finger-like projection. Third maxilliped (Fig. 6 B): Basal segment as long as penultimate segment; penultimate segment 1.01 – 1.11 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 4 – 5 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the middle of second segment, distal margin with long plumose setae. First pereiopod (Fig. 6 C): Chela 2.01 – 2.29 times as long as high, 1.32 – 1.69 times length of carpus; movable finger 3.00 – 3.63 times as long as wide and 0.96 – 1.33 times as long as palm, setal brushes well-developed; carpus 1.37 – 1.73 times as long as wide, slightly excavated distally; merus 0.91 – 1.29 times as long as carpus. Second pereiopod (Fig. 6 D): Chela 2.68 – 2.95 times as long as high, 0.86 – 0.88 times as long as carpus; movable finger 3.62 – 4.10 times as long as wide and 1.47 – 1.93 times as long as palm, setal brushes well-developed; carpus 4.30 – 4.44 times as long as wide, slightly excavated distally; merus 0.91 – 1.01 times as long as carpus. Third pereiopod (Fig. 6 E): Dactylus 3.21 – 4.28 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 5 spiniform setae; propodus 3.60 – 4.28 times as long as dactylus, 9.06 – 11.12 times as long as wide; carpus 0.65 – 0.69 times as long as propodus; merus 1.87 – 2.07 times as long as carpus, with 4 spiniform setae on the posterior margin. Ischium sometimes with 1 small movable spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 6 F): Dactylus 3.10 – 4.28 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 25 – 29 spiniform setae; propodus 3.82 – 4.94 times as long as dactylus, 11.22 – 13.56 times as long as wide; carpus 0.53 – 0.55 times as long as propodus; merus 1.53 – 1.57 times as long as carpus, with 3 – 4 spiniform setae on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 6 G, H): Endopod of male first pleopod subtriangular, 0.34 – 0.38 times as long as exopod, 2.27 – 2.44 times as long as wide; appendix interna well developed, arising from distal 0.33 of endopod, reaching slightly beyond the end of endopod, distally with cincinuli. Second pleopod (Fig. 6 I): Appendix masculina club-shaped, reaching 0.57 – 0.64 length of exopod, inner margin and tip bearing nearly equal long and stout spiniform setae; appendix interna well developed, reaching 0.25 length of appendix masculina, distally with cincinuli. Telson (Fig. 6 J): Tapering posterior, with a projection, with 5 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae. Exopodite of the uropod (Fig. 6 K) bears a series of 16 – 18 movable spinules along diaresis. Eggs: 0.57 – 0.62 mm × 0.98 – 1.02 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
DCE005D842CD522488F6083314BEB6E2.taxon	etymology	Etymology. Caridina caii is dedicated to Cai Y. X., who not only discovered the species but also made outstanding contributions to carcinology.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
DCE005D842CD522488F6083314BEB6E2.taxon	distribution	Distribution. Only occurring in Wuzhishan City, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
D80D72DF8A905F4B8DB6017279F8D0FF.taxon	description	Figs 2 A, 3, 4	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
D80D72DF8A905F4B8DB6017279F8D0FF.taxon	description	Description. Body (Fig. 2 A): Small, slender and sub-cylindrical, males up to 4.4 mm cl, females up to 5.3 mm cl. Sixth abdominal somite 0.40 – 0.53 times as long as carapace. Rostrum (Fig. 3 A): Short, slightly sloping downwards, reaching to the beginning to the end of the second segment of antennular peduncle; 0.4 – 0.5 of cl; armed dorsally with 12 – 16 teeth, including 4 – 6 on carapace posterior to orbital margin, ventrally with 1 – 5 teeth; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 3 A): Well developed, on short ocular peduncle, cornea globular. Carapace (Fig. 3 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 3 B): Stylocerite reaching 0.67 – 0.93 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.54 – 1.70 times as long as second segment; second segment 1.12 – 1.49 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 3 C): Scaphocerite 3.1 – 3.2 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 3 G): Palp of first maxilliped broadly triangular, ending in finger-like projection. Third maxilliped (Fig. 4 B): Basal segment 0.89 – 1.09 times as long as penultimate segment; penultimate segment as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 6 – 9 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the middle of second segament, distal margin with long plumose setae. First pereiopod (Fig. 4 C): Chela 2.11 – 2.38 times as long as high, 1.26 – 1.42 times as long as carpus; movable finger 3.14 – 3.92 times as long as wide and 1.02 – 1.30 times as long as palm, setal brushes well developed; carpus 1.71 – 2.03 times as long as wide, slightly excavated distally; merus 0.99 – 1.21 times as long as carpus. Second pereiopod (Fig. 4 D): Chela 2.83 – 3.29 times as long as high, 0.66 – 0.79 times as long as carpus; movable finger 3.97 – 4.81 times as long as wide and 1.46 – 1.58 times as long as palm, setal brushes well developed; carpus 5.42 – 6.30 times as long as wide, slightly excavated distally; merus 0.81 – 0.93 times as long as carpus. Third pereiopod (Fig. 4 E): Dactylus 3.00 – 4.20 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 5 – 6 spiniform setae; propodus 2.78 – 4.62 times as long as dactylus, 9.11 – 10.05 times as long as wide; carpus 0.64 – 0.70 times as long as propodus; merus 1.41 – 2.05 times as long as carpus, with 3 spiniform setae on the posterior margin. Ischium sometimes with 1 small movable spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 4 F): Dactylus 4.53 – 5.71 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 41 – 51 spiniform setae; propodus 2.94 – 3.43 times as long as dactylus, 11.53 – 13.43 times as long as wide; carpus 0.49 – 0.69 times as long as propodus; merus 1.49 – 1.73 times as long as carpus, with 2 – 3 spiniform setae on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 4 G): Endopod in male 0.46 – 0.67 times as long as exopod, oblong-reniform, 2.40 – 3.04 times as long as distally wide, tip rounded; appendix interna well developed, arising from distal 0.33 of endopod, reaching slightly beyond the end of endopod, distally with cincinuli. Second pleopod (Fig. 4 H): Appendix masculina club-shaped, reaching 0.60 – 0.70 length of exopod, some short spiniform setae on basal part, and inner margin and tip bearing nearly equal long and stout spiniform setae; appendix interna well developed, reaching 0.52 – 0.63 length of appendix masculina, distally with cincinuli. Telson (Fig. 4 I): With 4 – 5 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae. Exopodite of the uropod (Fig. 4 J) bears a series of 18 – 21 movable spinules along diaresis. Eggs: 0.65 – 0.70 mm × 1.07 – 1.20 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
D80D72DF8A905F4B8DB6017279F8D0FF.taxon	distribution	Distribution. Hainan Island, China and Vietnam.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
75A7515A0C615997B9AA16DE8AD43330.taxon	description	Figs 2 G, 15, 16	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
75A7515A0C615997B9AA16DE8AD43330.taxon	description	Description. Body (Fig. 2 G): Small, slender and sub-cylindrical, males up to 2.7 mm cl, females up to 3.4 mm cl. Sixth abdominal somite 0.44 – 0.48 times as long as carapace. Rostrum (Fig. 15 A): Straight, reaching to the end of the first segment to the end of the second segment of the antennular peduncle; 0.3 – 0.4 of cl; armed dorsally with 9 – 11 teeth, including 3 – 5 on carapace posterior to orbital margin, ventrally with 1 tooth; rostral formula 3 – 5 + 6 / 1; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 15 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 15 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 15 B): Peduncle reaching slightly short of scaphocerite; stylocerite reaching 0.76 – 0.96 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.70 – 1.77 times as long as second segment; second segment 1.35 – 1.66 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 15 C): Scaphocerite 3.1 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 15 G): Palp of first maxilliped broadly triangular, ending in finger-like projection. Third maxilliped (Fig. 16 B): Basal segment 0.95 – 1.06 times as long as penultimate segment; penultimate segment 0.85 – 0.94 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 5 – 6 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond middle of second segment, distal margin with long plumose setae. First pereiopod (Fig. 16 C): Chela 1.88 – 2.27 times as long as high, 1.26 – 1.44 times as long as carpus; movable finger 2.37 – 3.33 times as long as wide and 0.86 – 0.93 times as long as palm, setal brushes well-developed; carpus 1.36 – 2.11 times as long as wide, slightly excavated distally; merus 0.95 – 1.13 times as long as carpus. Second pereiopod (Fig. 16 D): Chela 2.31 – 2.49 times as long as high, 0.78 – 0.84 times as long as carpus; movable finger 3.59 – 4.50 times as long as wide and 1.28 – 1.80 times as long as palm, setal brushes well developed; carpus 4.64 – 6.23 times as long as wide; slightly excavated distally, merus 0.81 – 0.92 times as long as carpus. Third pereiopod (Fig. 16 E): Dactylus 3.59 – 4.49 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 4 – 6 spiniform setae; propodus 3.41 – 3.87 times as long as dactylus, 10.74 – 11.80 times as long as wide; carpus 0.67 – 0.68 times as long as propodus; merus 1.80 – 1.85 times as long as carpus, with 3 – 4 spiniform setae on posterior margin. Ischium sometimes with 1 spine on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 16 F): Dactylus 4.37 – 5.39 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 22 – 32 spiniform setae; propodus 3.06 – 3.43 times as long as dactylus, 11.62 – 14.91 times as long as wide; carpus 0.56 – 0.57 times as long as propodus; merus 1.49 – 1.57 times as long as carpus, with 3 – 5 spiniform setae on posterior margin. Ischium sometimes with 1 spine on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 16 G, H): Endopod in male oblong-reniform, 0.49 – 0.61 times as long as exopod, 2.76 – 3.29 times as long as distally wide, distal end is distinctly narrower than the basal part, with a concave inner margin and a convex outer margin, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well developed, arising from distal 0.25 of endopod, reaching slightly beyond end of endopod, distally with cincinuli. Second pleopod (Fig. 16 I): Appendix masculina rod-shaped, reaching 0.63 – 0.71 length of exopod, bearing sparse long spiniform setae; appendix interna well developed, reaching 0.72 – 0.78 length of appendix masculina, distally with cincinuli. Telson (Fig. 16 J): Tapering posterior, with a projection, with 3 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 3 – 4 pairs of intermedial spiniform setae. Exopodite of the uropod (Fig. 16 K) bears a series of 12 – 16 movable spinules along diaresis. Eggs: 0.56 – 0.67 mm × 0.83 – 0.97 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
75A7515A0C615997B9AA16DE8AD43330.taxon	etymology	Etymology. The name originates from their type locality, referring to their habitat at the spring’s outlet, with “ fontinalis ” meaning “ spring-dweller ” in reference to their dwelling place near the water source.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
75A7515A0C615997B9AA16DE8AD43330.taxon	distribution	Distribution. Only occurring in the type locality, Qiongshan District, Haikou City, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
25171598E26953089CE50498C6720EC9.taxon	description	Figs 2 F, 13, 14	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
25171598E26953089CE50498C6720EC9.taxon	description	Description. Body (Fig. 2 F): Small, slender and sub-cylindrical, males up to 4.6 mm cl, females up to 5.0 mm cl. Sixth abdominal somite 0.46 – 0.66 times as long as carapace. Rostrum (Fig. 13 A): Short, straight, reaching to the end of the eye to the end of the first segment of antennular peduncle; 0.2 – 0.3 of cl; usually unarmed, rarely armed dorsally with 1 – 3 teeth, and 0 – 1 teeth on carapace posterior to orbital margin, ventrally unarmed; rostral formula 0 – 1 + 0 – 2 / 0; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 13 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 13 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 13 B): Peduncle reaching slightly short of scaphocerite; stylocerite reaching 0.84 – 0.93 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.31 – 1.42 times as long as second segment; second segment 1.07 – 1.29 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 13 C): Scaphocerite 2.9 – 3.2 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 13 G): Palp of first maxilliped broadly triangular, ending in a bulbous tip. Third maxilliped (Fig. 14 B): Basal segment 1.09 – 1.49 times as long as penultimate segment; penultimate segment 0.80 – 0.96 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 4 – 5 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching to the end of second segment, distal margin with long plumose setae. First pereiopod (Fig. 14 C): Chela 1.70 – 2.08 times as long as high, 1.27 – 1.49 times as long as carpus; movable finger 2.22 – 2.99 times as long as wide and 0.72 – 1.21 times as long as palm, setal brushes well-developed; carpus 1.33 – 1.67 times as long as wide, slightly excavated distally; merus 1.02 – 1.17 times as long as carpus. Second pereiopod (Fig. 14 D): Chela 2.25 – 3.03 times as long as high, 0.67 – 1.06 times as long as carpus; movable finger 3.63 – 4.45 times as long as wide and 1.11 – 1.74 times as long as palm, setal brushes well-developed; carpus 3.49 – 5.51 times as long as wide, slightly excavated distally; merus 0.87 – 1.31 times as long as carpus. Third pereiopod (Fig. 14 E): Dactylus 3.04 – 3.48 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 4 – 5 spiniform setae; propodus 3.28 – 3.91 times as long as dactylus, 7.98 – 9.62 times as long as wide; carpus 0.65 – 0.77 times as long as propodus; merus 1.71 – 1.99 times as long as carpus, with 3 spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 14 F): Dactylus 3.27 – 4.00 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 36 – 57 spiniform setae; propodus 3.35 – 4.35 times as long as dactylus, 9.24 – 12.23 times as long as wide; carpus 0.55 – 0.66 times as long as propodus; merus 1.34 – 1.51 times as long as carpus, with 2 – 3 strong spines on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 14 G): Endopod in male 0.46 – 0.56 times as long as exopod, oblong-reniform, 2.53 – 3.25 times as long as distally wide, tip rounded, inner margin slightly straight, outer margin slightly convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well-developed, arising from distal 0.33 of endopod, reaching slightly beyond end of endopod, distally with cincinuli. Second pleopod (Fig. 14 H): Appendix masculina rod-shaped, slightly thicker centrally, narrower distally, reaching 0.64 – 0.73 length of exopod, some short spiniform setae on basal part, and inner margin and tip bearing two rows of long spiniform setae; appendix interna well-developed, reaching 0.48 – 0.51 length of appendix masculina, distally with cincinuli. Telson (Fig. 14 I): Tapering posterior, with a projection, with 4 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae, two outer pairs subequal in length and slightly shorter than two middle pairs. Exopodite of uropod (Fig. 14 J) bears a series of 19 – 23 movable spinules along diaresis. Eggs: 0.67 – 0.80 mm × 1.10 – 1.20 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
25171598E26953089CE50498C6720EC9.taxon	etymology	Etymology. The new species is named after its type locality name: Heping Town.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
25171598E26953089CE50498C6720EC9.taxon	distribution	Distribution. Only occurring in Heping Town, Qiongzhong County, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
C623936158525894BD29E996A0F0C0E7.taxon	description	Figs 2 E, 11, 12	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
C623936158525894BD29E996A0F0C0E7.taxon	description	Description. Body (Fig. 2 E): Small, slender and sub-cylindrical, males up to 4.7 mm cl, females up to 4.6 mm cl. Sixth abdominal somite 0.46 – 0.52 times as long as carapace. Rostrum (Fig. 11 A): Short, straight, reaching to the end of the first segment to the middle of the second segment of antennular peduncle; 0.2 – 0.5 of cl; armed dorsally with 9 – 15 teeth, including 3 – 5 on carapace posterior to orbital margin, ventrally with 0 – 2 teeth; rostral formula 3 – 5 + 6 – 10 / 0 – 2; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 11 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 11 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 11 B): Stylocerite reaching 1.19 – 1.31 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.32 – 1.53 times as long as second segment; second segment 1.20 – 1.45 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 11 C): Scaphocerite 3.0 – 3.3 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 11 G): Palp of first maxilliped broadly triangular, ending in finger-like projection. Third maxilliped (Fig. 12 B): Basal segment 0.94 – 1.21 times as long as penultimate segment; penultimate segment 0.92 – 1.00 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 5 – 6 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the middle of second segment, distal margin with long plumose setae. First pereiopod (Fig. 12 C): Chela 2.00 – 2.33 times as long as high, 1.30 – 1.44 times as long as carpus; movable finger 2.41 – 3.00 times as long as wide and 0.80 – 1.21 times as long as palm, setal brushes well developed; carpus 1.51 – 2.40 times as long as wide, slightly excavated distally; merus 1.03 – 1.21 times as long as carpus. Second pereiopod (Fig. 12 D): Chela 2.44 – 2.92 times as long as high, 0.75 – 0.86 times as long as carpus; movable finger 3.87 – 4.79 times as long as wide and 1.10 – 1.54 times as long as palm, setal brushes well-developed; carpus 4.78 – 5.45 times as long as wide, slightly excavated distally; merus 0.79 – 1.03 times as long as carpus. Third pereiopod (Fig. 12 E): Dactylus 3.22 – 3.81 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 4 – 6 spiniform setae; propodus 3.29 – 4.44 times as long as dactylus, 8.32 – 10.25 times as long as wide; carpus 0.58 – 0.79 times as long as propodus; merus 1.79 – 2.25 times as long as carpus, with 3 spiniform setae on posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 12 F): Dactylus 3.27 – 4.85 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 41 – 49 spiniform setae; propodus 3.35 – 3.62 times as long as dactylus, 11.22 – 13.12 times as long as wide; carpus 0.48 – 0.73 times as long as propodus; merus 1.48 – 1.86 times as long as carpus, with 2 – 3 spiniform setae on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 12 G): Endopod in male 0.44 – 0.46 times as long as exopod, oblong-reniform, 2.40 – 2.80 times as long as distally wide, tip rounded, inner margin slight concave, outer margin slightly convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well-developed, arising from distal 0.33 of endopod, reaching slightly beyond end of endopod, distally with cincinuli. Second pleopod (Fig. 12 H): Appendix masculina rod-shaped, reaching 0.59 – 0.67 times length of exopod, some short spiniform setae on basal part, and inner margin and tip bearing nearly equal long and stout spiniform setae; appendix interna well developed, reaching 0.49 – 0.55 length of appendix masculina, distally with cincinuli. Telson (Fig. 12 I): Tapering posterior, with a projection, with 5 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae. Exopodite of the uropod (Fig. 12 J) bears a series of 19 – 26 movable spinules along diaresis. Eggs: 0.66 – 0.85 mm × 1.10 – 1.27 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
C623936158525894BD29E996A0F0C0E7.taxon	etymology	Etymology. The new species is named after its discoverer Mr. Huang.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
C623936158525894BD29E996A0F0C0E7.taxon	distribution	Distribution. Only occurring in Heping Town, Qiongzhong County, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
84E116990D8758119159FC93974AEE28.taxon	description	Figs 2 D, 9, 10	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
84E116990D8758119159FC93974AEE28.taxon	description	Description. Body (Fig. 2 D): Small, slender and sub-cylindrical, males up to 4.6 mm cl, females up to 5.0 mm cl. Sixth abdominal somite 0.45 – 0.52 times as long as carapace. Rostrum (Fig. 9 A, B): Short, straight, reaching to the end of the first segment to the middle of 2 rd segment of antennular peduncle; 0.2 – 0.4 of cl; armed dorsally with 8 – 12 teeth, including 2 – 5 on carapace posterior to orbital margin, ventrally with 0 – 2 teeth; rostral formula 2 – 5 + 6 – 7 / 0 – 2; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 9 A, B): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 9 A, B): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 9 C): Stylocerite reaching 1.05 – 1.26 times as long as basal segment of antennular peduncle; basal segment as long or shorter than combined length of second and third segments, 1.48 – 1.63 times as long as second segment; second segment 1.70 – 2.01 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 9 D): Scaphocerite 3.7 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 9 H): Palp of first maxilliped broadly triangular, ending in finger-like projection. Third maxilliped (Fig. 10 B): Basal segment 0.95 – 1.26 times as long as penultimate segment; penultimate segment 0.96 – 1.15 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 5 – 6 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the beginning of second segment, distal margin with long plumose setae. First pereiopod (Fig. 10 C): Chela 2.06 – 2.25 times as long as high, 1.34 – 1.51 times as long as carpus; movable finger 2.58 – 3.32 times as long as wide and 0.87 – 1.16 times as long as palm, setal brushes well-developed; carpus 1.18 – 1.66 times as long as wide, slightly excavated distally; merus 0.95 – 1.44 times as long as carpus. Second pereiopod (Fig. 10 D): Chela 2.38 – 2.90 times as long as high, 0.83 – 0.90 times as long as carpus; movable finger 3.54 – 4.51 times as long as wide and 1.13 – 1.53 times as long as palm, setal brushes well-developed; carpus 4.13 – 4.90 times as long as wide, slightly excavated distally; merus 0.95 – 1.12 times as long as carpus. Third pereiopod (Fig. 10 E): Dactylus 2.88 – 4.03 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 5 – 6 spiniform setae; propodus 2.88 – 4.51 times as long as dactylus, 8.72 – 10.61 times as long as wide; carpus 0.61 – 0.87 times as long as propodus; merus 1.76 – 2.16 times as long as carpus, with 3 – 4 spiniform setae on the posterior margin. Ischium sometimes with 1 small movable spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 10 F): Dactylus 3.22 – 4.70 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 35 – 52 spiniform setae; propodus 3.02 – 4.94 times as long as dactylus, 9.38 – 12.47 times as long as wide; carpus 0.43 – 0.67 times as long as propodus; merus 1.41 – 1.55 times as long as carpus, with 2 – 3 spiniform setae on the posterior margin. Ischium sometimes with 1 spine on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 10 G): Endopod in male 0.50 – 0.56 times as long as exopod, oblong-reniform, distal end is distinctly narrower than the basal part, 2.77 – 3.09 times as long as wide, tip rounded, inner margin slightly concave. Outer margin slightly convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well developed, arising from distal 0.25 – 0.33 of endopod, reaching slightly beyond end of endopod, distally with cincinuli. Second pleopod (Fig. 10 H): Appendix masculina rod-shaped, wider and slightly swollen in the end, reaching 0.68 – 0.69 length of exopod, some short spiny setae on basal part, the spiniform setae at the end of appendix masculina are thicker, and the spiniform setae of middle of appendix masculina are shorter; appendix interna well developed, reaching 0.50 – 0.56 of appendix masculina, distally with cincinuli. Telson (Fig. 10 I): With 5 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae, the innermost one usually strongest and longest. Exopodite of the uropod (Fig. 10 I) bears a series of 20 – 23 movable spinules along diaresis. Eggs: 0.70 – 0.76 mm × 1.17 – 1.30 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
84E116990D8758119159FC93974AEE28.taxon	etymology	Etymology. The new species is named after its type locality name: Qiongzhong County.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
84E116990D8758119159FC93974AEE28.taxon	distribution	Distribution. Only occurring in Yinggen Town, Qiongzhong County, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
0779195C54415BC98730BFB71541D327.taxon	description	Figs 2 H, 17, 18	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
0779195C54415BC98730BFB71541D327.taxon	description	Description. Body (Fig. 2 H): Small, slender and sub-cylindrical, males up to 4.3 mm cl, females up to 4.4 mm cl. Sixth abdominal somite 0.40 – 0.51 times as long as carapace. Rostrum (Fig. 17 A): Short, straight, reaching from not beyond the eye to beginning of second segment of antennular peduncle; 0.2 – 0.3 of cl; armed dorsally with 3 – 13 teeth, including 0 – 5 on carapace posterior to orbital margin, ventrally with 1 – 2 teeth; rostral formula 0 – 5 + 3 – 8 / 1 – 2; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 17 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 17 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 17 B): Stylocerite reaching 0.79 – 0.90 times as long as basal segment of antennular peduncle; basal segment shorter than combined length of second and third segments, 1.46 – 1.74 times as long as second segment; second segment 1.16 – 1.40 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 17 C): Scaphocerite 3.3 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 17 G): Palp of first maxilliped broadly triangular, ending with no finger-like projection. Third maxilliped (Fig. 18 B): Basal segment 1.07 – 1.17 times as long as penultimate segment; penultimate segment 0.97 – 1.02 times as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 5 – 6 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching the middle of second segment, distal margin with long plumose setae. First pereiopod (Fig. 18 C): Chela 1.79 – 2.26 times as long as high, 1.21 – 1.34 times as long as carpus; movable finger 2.08 – 2.77 times as long as wide and 0.71 – 1.33 times as long as palm, setal brushes well developed; carpus 1.38 – 1.85 times as long as wide, slightly excavated distally; merus 1.03 – 1.11 times as long as carpus. Second pereiopod (Fig. 18 D): Chela 2.22 – 3.15 times as long as high, 0.66 – 0.76 times as long as carpus; movable finger 2.85 – 4.37 times as long as wide and 1.07 – 1.47 times as long as palm, setal brushes well developed; carpus 5.17 – 5.66 times as long as wide, slightly excavated distally; merus 0.88 – 0.99 times as long as carpus. Third pereiopod (Fig. 18 E): Dactylus 2.95 – 3.96 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 5 – 7 spiniform setae; propodus 3.04 – 4.24 times as long as dactylus, 8.13 – 10.05 times as long as wide; carpus 0.67 – 0.82 times as long as propodus; merus 1.42 – 1.96 times as long as carpus, with 3 spiniform setae on the posterior margin. Ischium sometimes with 1 small movable spiniform setae on the posterior margin. Third and fourth pereiopods with 1 spine on ischium. Fifth pereiopod (Fig. 18 F): Dactylus 3.37 – 4.62 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 36 – 42 spiniform setae; propodus 3.37 – 4.18 times as long as dactylus, 9.63 – 12.61 times as long as wide; carpus 0.51 – 0.61 times as long as propodus; merus 1.42 – 1.63 times as long as carpus, with 3 spiniform setae on the posterior margin. First pleopod (Fig. 18 G): Endopod reaching 0.42 – 0.45 times as long as exopod, oblong-reniform, 2.46 – 2.58 times as long as distally wide, terminal width subequal to basal width; inner margin concave, outer margin convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well developed, arising from distal 0.2 of endopod, reaching slightly beyond the end of endopod, distally with cincinuli. Second pleopod (Fig. 18 H): Appendix masculina rod-shaped, reaching 0.63 – 0.64 length of endopod; with some short spiniform setae on outer surface and some long and stout spiniform setae on distal surface; appendix interna reaching 0.50 – 0.56 length of appendix masculina. Telson (Fig. 18 I): With 4 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin triangular, with 3 – 4 pairs of intermedial plumose setae, lateral plumose setae slightly longer, intermediate plumose setae subequal. Exopodite of the uropod (Fig. 18 J) bears a series of 17 – 22 movable spinules along diaresis. Eggs: 0.71 – 0.78 mm × 1.09 – 1.23 mm in diameter.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
0779195C54415BC98730BFB71541D327.taxon	etymology	Etymology. The name is derived from its type locality, Tuncheng Town.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
0779195C54415BC98730BFB71541D327.taxon	distribution	Distribution. Only occurring in the type locality, Tuncheng Town, Tunchang County, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
2B6806B8C1445392B32A1E3C79838585.taxon	description	Figs 2 C, 7, 8	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
2B6806B8C1445392B32A1E3C79838585.taxon	description	Description. Body (Fig. 2 C): Small, slender and sub-cylindrical, males up to 3.4 mm cl, females up to 3.4 mm cl. Sixth abdominal somite 0.41 – 0.48 times as long as carapace. Rostrum (Fig. 7 A): Short, reaching to the end of the first segment to the middle of the second segment of antennular peduncle, sloping ventral anteriorly; 0.3 – 0.5 of cl; armed dorsally with 12 – 15 teeth, including 3 – 5 on carapace posterior to orbital margin, ventrally with 2 – 4 teeth; rostral formula 3 – 5 + 9 – 10 / 2 – 4; lateral carina dividing rostrum into two unequal parts, continuing posteriorly to orbital margin. Eyes (Fig. 7 A): Well-developed, on short ocular peduncle, cornea globular. Carapace (Fig. 7 A): Smooth, glabrous; antennal spine acute, fused with inferior orbital angle; pterygostomian margin broadly rectangular, slightly produced forward; pterygostomian spine absent. Antennule (Fig. 7 B): Peduncle reaching slightly short of scaphocerite; stylocerite reaching 0.86 – 0.96 times as long as basal segment of antennular peduncle; basal segment shorter or as long as combined length of second and third segments, 1.72 – 1.96 times as long as second segment; second segment 1.00 – 1.10 times as long as third segments; all segments with marginal plumose setae. Antenna (Fig. 7 C): Scaphocerite 3.0 – 3.2 times as long as wide, outer margin straight, asetose, ending in a strong sub-apical spine, inner and anterior margins with long plumose setae. First maxilliped (Fig. 7 G): Palp of first maxilliped broadly triangular, ending in a finger-like projection. Third maxilliped (Fig. 8 B): Basal segment 0.91 – 1.10 times as long as penultimate segment; penultimate segment as long as distal segment, ending in a large claw-like spine surrounded by simple setae, preceded by 3 – 5 thin spines on distal third of posterior margin, proximally a clump of long and short simple, serrate setae; exopod reaching beyond the middle of second segment, distal margin with long plumose setae. First pereiopod (Fig. 8 C): Chela 1.90 – 2.26 times as long as high, 1.21 – 1.55 times as long as carpus; movable finger 2.55 – 3.45 times as long as wide and 0.86 – 1.10 times as long as palm, setal brushes well developed; carpus 1.49 – 1.94 times as long as wide, slightly excavated distally; merus 0.95 – 1.16 times as long as carpus. Second pereiopod (Fig. 8 D): Chela 2.30 – 2.70 times as long as high, 0.75 – 0.80 times as long as carpus; movable finger 3.50 – 3.81 times as long as wide and 1.24 – 1.53 times as long as palm, setal brushes well-developed; carpus 4.59 – 5.41 times as long as wide, slightly excavated distally; merus 0.74 – 1.00 times as long as carpus. Third pereiopod (Fig. 8 E): Dactylus 3.36 – 3.90 times as long as wide, terminating in a prominent claw-like spine surrounded by simple setae, followed by 5 spiniform setae; propodus 3.27 – 3.83 times as long as dactylus, 8.65 – 9.43 times as long as wide; carpus 0.60 – 0.69 times as long as propodus; merus 1.80 – 2.29 times as long as carpus, with 3 – 4 spiniform setae on the posterior margin. Fourth pereiopod: Proportion and spination similar to third pereiopod. Fifth pereiopod (Fig. 8 F): Dactylus 3.13 – 4.24 times as long as wide, ending in prominent claw-like spine surrounded by simple setae, followed by a row of 29 – 31 spiniform setae; propodus 3.30 – 4.24 times as long as dactylus, 9.16 – 11.29 times as long as wide; carpus 0.54 – 0.58 times as long as propodus; merus 1.42 – 1.61 times as long as carpus, with 3 spiniform setae on the posterior margin. First four pereiopods with epipod. First pleopod (Fig. 8 G): Endopod in male 0.49 – 0.51 times as long as exopod, oblong-reniform, 2.72 – 3.34 times as long as distally wide, tip rounded, inner margin concave, bearing nearly equal short spine setae, outer margin slightly convex, long pappose setae on outer and distal margins, medium-length setae on inner margin; appendix interna well developed, arising from distal 0.33 of endopod, reaching slightly beyond end of endopod, distally with cincinuli. Second pleopod (Fig. 8 H): Appendix masculina rod-shaped, reaching 0.69 – 0.71 length of exopod, some short spiniform setae on basal part, and inner margin and tip bearing nearly equal long and stout spiniform setae; appendix interna well-developed, reaching 0.55 – 0.57 length of appendix masculina, distally with cincinuli. Telson (Fig. 8 I): Tapering posterior, with a projection, with 4 pairs of short spiniform setae dorsally and one pair of short spiniform setae dorsolaterally; posterior margin with 4 pairs of intermedial spiniform setae, close in length. Exopodite of the uropod (Fig. 8 J) bears a series of 14 – 18 movable spinules along diaresis. Eggs: 0.58 – 0.60 mm × 0.92 – 0.98 mm in diameter. Females of this small-bodied species attain sexual maturity and carry eggs at approximately tl 11 mm.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
2B6806B8C1445392B32A1E3C79838585.taxon	etymology	Etymology. The new species is named after its type locality name: Wenchang City.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
2B6806B8C1445392B32A1E3C79838585.taxon	distribution	Distribution. Only occurring in Chongxing Town, Wenchang City, Hainan Island.	en	Hou, Junjie, Zhang, Jiping, Chen, Bing, Zhang, Yixuan, Chen, Wenjian, Guo, Zhaoliang (2025): Integrative taxonomy reveals the Caridina clinata Cai, Nguyen & Ng, 1999 species complex (Crustacea, Decapoda, Atyidae), with descriptions of eight new species from Hainan Island, China. Zoosystematics and Evolution 101 (4): 2295-2336, DOI: 10.3897/zse.101.172207
