taxonID	type	description	language	source
F2344B000629589DB9BF07898177B7A2.taxon	description	Figs 6, 7, 8, 9, 10	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
F2344B000629589DB9BF07898177B7A2.taxon	etymology	Etymology. The species epithet “ bilobus ” adjective, is used to emphasize the solenophore being strongly divided into two large lobes.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
F2344B000629589DB9BF07898177B7A2.taxon	diagnosis	Diagnosis. The species is characterized by having sub-moniliform body, poorly developed paraterga, gonopod femorites stout, somewhat waisted at the middle, basal part swollen dorsally, and separated from postfemoral region laterally by a distinct demarcation sulcus; solenophore rather simple, with only a long basal process, coiled posteriad; solenomere relatively long.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
F2344B000629589DB9BF07898177B7A2.taxon	description	Description. Size: body length 13.5 – 15.0 mm (males), 17.66 – 18.85 mm (females). Width of midbody pro- and metazona 1.04 – 1.19 mm (males), 1.51 – 1.68 mm (females), and 1.20 – 1.27 mm (males), 1.57 – 1.76 mm (females), respectively. Length of holotype 14.44 mm, width of pro- and metazona 1.10 and 1.24 mm, respectively. Coloration: generally reddish brown, except for a brownish yellow broad, longitudinal band in middle metaterga, as well as lower part of pleura and sterna. Legs pale yellow, antennae light brown, except antennomeres 6 and 7 whitish. Head (Fig. 6 C): labrum and frons densely setose, vertex sparsely setose along epicranial suture. Latter deep, distinct, running from rear margin of vertex down to labrum. Antennae slender and very long, reaching behind segment 5 if stretched laterally; antennomere 1 = 7 <6 <5 <2 = 4 in length. Collum (Fig. 6 B): a little smaller than head, surface smooth, with two rows of setae: 4 + 4 close to anterior margin and 2 + 2 nearly posterior margin. All setae broken. Body (Figs 6 A, B, 6 D, 7 A – C) submoniliform, width of ring 3 <4 <2 = 5, parallel-sided on rings 5 – 16, thereafter gradually tapering towards telson. Metatergal surface smooth, shagreened, shining, with only one row of 2 + 2 setae near anterior margin. Transverse sulcus very poorly developed, superficial. Prozona and pleura also shagreened and shining. Stricture dividing pro- and metazona rather distinct and smooth, neither striolate nor beaded. Paraterga (Figs 6 A, B, 6 D, 7 A – C) strongly reduced, almost missing, slightly more easily traceable as lateral keels on pore-bearing rings than on poreless ones. Ozopore formula normal, pores located on paraterga 5, 7, 9, 10, 12, 13, and 15 – 19. Pleurosternal carinae well-developed until segment 16, missing on subsequent ones (Figs 6 A, 7 A). Telson (Figs 7 D, 8 A): Epiproct long and curved down, with four small spinnerets at tip. Paraprocts semi-circular, each with two setae located on disk. Hypoproct roundly triangular, with two distolateral setiferous knobs (Figs 7 D, 8 A). Sternites (Figs 7 B, 8 B) sparsely setose, cross-impression with both transverse and longitudinal sulci moderately developed. Sternal cones small but obvious, front pairs even smaller than caudal pairs (Fig. 7 B). Sternite 5 with two independent small setiferous rectangular laminae between coxae 4 and two similar laminae between coxae 5 (Fig. 8 B). Legs slender and about 1.6 times as long as body height. Tarsal brushes present until ring 16, thereafter missing. Prefemora swollen dorsally. Femora without modifications. Gonopod (Figs 9, 10) rather complex. Coxite (co) cylindrical, about half length of telopodite, distoventral part sparsely setose. Prefemoral part as usual densely setose. Femorite (fe) stout, somewhat waisted at the middle, basal part swollen dorsally, and separated from postfemoral region laterally by a distinct demarcation sulcus. Solenophore (sph) rather simple, with only a long, posteriad-coiled basal process (lbp). Seminal groove (sg) running distodorsad fully on mesal side of femorite, Solenomere (sl) flagelliform, relatively long, only its proximal half sheathed by solenophore.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
5201F155866F5A0CA4FABB86A3B461A1.taxon	description	Figs 11, 12, 13, 14, 15	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
5201F155866F5A0CA4FABB86A3B461A1.taxon	etymology	Etymology. The species epithet “ gladius ” adjective, is used to emphasize the blade-shaped solenophore of the male gonopod.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
5201F155866F5A0CA4FABB86A3B461A1.taxon	diagnosis	Diagnosis. The species is characterized by having sub-moniliform body, poorly developed paraterga, gonopod femorite stout, enlarged distad, swollen dorsally. distal part with an acute spine distomesally; curved posteriad, large blade-shaped with an acute terminal, basal part with a rectangular process; solenomere short but longer than solenophore.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
5201F155866F5A0CA4FABB86A3B461A1.taxon	description	Description. Size: body length 25.97 – 28.02 mm (males), 33.33 mm (female). Width of midbody pro- and metazona 2.15 – 2.36 mm (males), 3.08 mm (female), and 2.30 – 2.48 mm (males), 3.24 mm (female), respectively. Length of holotype 25.97 mm, width of pro- and metazona 2.15 mm and 2.30 mm, respectively. Coloration: generally brownish yellow, except prozona and anterior halves of metatzona brownish. Legs pale yellow, antennae light brown. Head (Fig. 11 C): labrum and frons densely setose, vertex sparsely setose along epicranial suture. Latter deep, distinct, running from rear margin of vertex down to labrum. Antennae slender and very long, reaching behind segment 6 if stretched laterally; antennomere 1 = 7 <6 <5 <2 = 4 in length. Collum (Fig. 11 A, B): a little wider than head, surface smooth, with two rows of setae: 4 + 4 close to anterior margin and 2 + 2 nearly posterior margin. All setae broken. Body (Figs 11 – 12) submoniliform, width of ring 4 <3 <2 = 5, parallel-sided on rings 5 – 16, thereafter gradually tapering towards telson. Metatergal surface smooth, shagreened, shining, with only one row of 2 + 2 setae near anterior margin. Transverse sulcus very poorly developed, superficial, possibly visible only on metaterga 5 – 18. Prozona and pleura also shagreened and shining. Stricture dividing pro- and metazona rather distinct and smooth, neither striolate nor beaded. Paraterga (Figs 11 A, B, 11 D, 12 A, 12 C) strongly reduced, almost missing, slightly more easily traceable as lateral keels on pore-bearing segments than on poreless ones. Ozopore (Fig. 11 B, 11 D) formula normal, pores located on paraterga 5, 7, 9, 10, 12, 13, and 15 – 19. Pleurosternal carinae well-developed until segment 16, missing on subsequent ones. Telson (Figs 12 C – D, 13 A): Epiproct long and curved down, with four small spinnerets at tip. Paraprocts semi-circular, each with two setae located on disk. Hypoproct roundly trapeziform, with two distolateral setiferous knobs (Fig. 13 A). Sternites (Figs 12 B, 13 B) sparsely setose, cross-impression with both transverse and longitudinal sulci moderately developed. Sternal cones obvious, front pairs slightly stouter, caudal pairs somewhat slenderer and longer. Sternite 5 with an elevated bifid lamina between coxae 4 and a ventrad-elevated setiferous process between coxae 5 (Fig. 13 B). Legs: slender and about 1.6 times as long as body height. Tarsal brushes present until ring 15, thereafter missing. Prefemora not swollen dorsally. Femora without modifications. Gonopod (Figs 14 – 15) rather complex. Coxite (co) stout, about half length of telopodite, distoventral part sparsely setose. Prefemoral part as usual densely setose. Femorite (fe) stout, enlarged distad, swollen dorsally, with a basal spine and an acute spine (as) distomesally; separated from postfemoral region laterally by a distinct demarcation sulcus. Solenophore complex, curved posteriad, large blade-shaped with an acute terminal; basal part with a rectangular process (rp). Seminal groove running distodorsad fully on mesal side of femorite, Solenomere (sl) flagelliform, longer than solenophore, only its proximal half sheathed by solenophore.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
C626BB634F46505586511BC635C6234C.taxon	description	Figs 1, 2, 3, 4, 5	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
C626BB634F46505586511BC635C6234C.taxon	etymology	Etymology. The species epithet “ lobus ” adjective, is used to emphasize the large lobe on gonopod femorite.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
C626BB634F46505586511BC635C6234C.taxon	diagnosis	Diagnosis. The species is characterized by having submoniliform body, poorly developed paraterga, gonopod femorite enlarged distad, with a large membraneous lobe and a distal spiniform process on dorsal side; solenophore complex curved down, with two basal spines and a distal laminate lobe; tip strongly bifid; solenomere relatively long.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
C626BB634F46505586511BC635C6234C.taxon	description	Description. Size: body length 31.34 – 32.11 mm (males), 31.29 – 32.78 mm (females). Width of midbody pro- and metazona 2.13 – 2.18 mm (males), 2.88 – 3.21 mm (females), and 2.45 – 2.56 mm (males), 3.2 – 3.47 mm (females), respectively. Length of holotype 31.34 mm, width of pro- and metazona 2.18 and 2.56 mm, respectively. Coloration generally dark brown, except for yellowish brown middle parts of metaterga, as well as pleura and sterna. Legs pale yellow, antennae light brown. Head: labrum and frons densely setose, vertex sparsely setose along epicranial suture. Latter deep, distinct, running from rear margin of vertex down to labrum. Antennae slender and very long, reaching to segment 4 if stretched laterally. Antennomere 1 = 7 <6 <3 = 4 = 5 <2 in length. Collum (Fig. 1 A, B): a little wider than head, surface smooth, with two rows of setae: 4 + 4 close to anterior margin and 2 + 2 nearly posterior margin. Body (Figs 1 B, D, 2 A, C) submoniliform, width of body ring 3 <4 <2 = 5, parallel-sided on body rings 5 – 16, thereafter gradually tapering towards telson. Metatergal surface smooth, shagreened, shining, with only one row of 2 + 2 setae near anterior margin (broken in preserved specimens). Transverse sulcus poorly developed, superficial on metaterga 5 – 18. Prozona and pleura also shagreened and shining. Stricture dividing pro- and metazona rather distinct and smooth, neither striolate nor beaded (Figs 1, 2). Paraterga (Figs 1 A, B, D, 2 A, C, D) strongly reduced, almost missing, slightly more easily traceable as lateral keels on pore-bearing body rings than on poreless ones. Ozopore formula normal, pores located on paraterga 5, 7, 9, 10, 12, 13, and 15 – 19. Pleurosternal carinae well-developed until body ring 16, missing on subsequent ones (Figs 1 A, 2 A, C). Telson (Figs 2 C, D, 3 A): Epiproct long and curved down, with four small spinnerets at tip. Hypoproct subtrapeziform, with two distolateral setiferous knobs (Fig. 3 A). Sternites (Figs 1 C, 2 B) sparsely setose, cross-impression with both transverse and longitudinal sulci moderately developed. Sternal cones obvious, front pairs slightly stouter, caudal pairs somewhat slenderer and longer (Fig. 2 B). Sternite 5 with an elevated bifid lamina between coxae 4 and a ventrad-elevated setiferous process between coxae 5 (Figs 1 C, 3 B). Legs slender and long, about 1.7 times as long as body height. Tarsal brushes present until ring 17, thereafter missing. Prefemora swollen dorsally. Femora without modifications. Gonopod (Figs 4, 5) rather complex. Coxite (co) stout, about half length of telopodite, distoventral part sparsely setose. Prefemoral part (pref) as usual densely setose. Femorite (fe) stout, enlarged distad, with a basal straight spine (bsp), a large membraneous lobe (lo), and a small distal spiniform process (dsp) on dorsal side, separated from postfemoral region laterally by a distinct demarcation sulcus. Solenophore very complex, curved down, with two basal spines (bs 1 and bs 2) and a spiniform process (sp) in middle; tip strongly bifid. Seminal groove (sg) running distodorsad fully on mesal side of femorite. Solenomere (sl) flagelliform, relatively long, only its proximal half sheathed by solenophore.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
67C21A3196F15D8AB4E1F8848C3609A6.taxon	type_taxon	Type species. Anoplodesmus anthracinus Pocock, 1895 (p. 798, fig. 5), by subsequent designation.	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
67C21A3196F15D8AB4E1F8848C3609A6.taxon	diagnosis	Diagnosis. The genus is easily recognized by the following characters: paraterga modest or strongly reduced, legs usually with tarsal brushes, sterna usually with four cones, gonopod more or less elaborate, with or without a distofemoral process, both lamina medialis and lamina lateralis well-developed with several obvious lobes, solenomere as long as solonophore or extremely long (Golovatch and Semenyuk 2010). The concept of this genus was discussed more clearly by Golovatch (1999) and Golovatch and Semenyuk (2010).	en	Nguyen, Anh D., Vu, Tam T. T., Nguyen, Thu Anh T., Le, Son X. (2025): Mountainous millipedes in Vietnam. IV. Three new Anoplodesmus Pocock, 1895 in northern Vietnam (Diplopoda, Polydesmida, Paradoxosomatidae), with notes on its phylogenetics. Zoosystematics and Evolution 101 (3): 919-934, DOI: 10.3897/zse.101.150442
