identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B84387E3FF91FFFAFCA0F91275A7D895.text	B84387E3FF91FFFAFCA0F91275A7D895.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spirostreptida Brandt 1833	<div><p>Order Spirostreptida Brandt, 1833</p><p>Remarks: Our fossil specimen clearly belongs to the order Spirostreptida based on the following characters: Architecture of anterior body-rings; body-ring 4 legless (Figs. 1C, 2A, B)); gnathochilarial characters: stipites always separated by mentum and reaching base of gnathochilarium (Fig. 2E); absence of setae from frontal setae as well as setation at body rings (present in Julida).</p></div>	https://treatment.plazi.org/id/B84387E3FF91FFFAFCA0F91275A7D895	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Wesener, Thomas;RÜhr, Peter T.	Wesener, Thomas, RÜhr, Peter T. (2025): Internal gonopod reconstruction in an amber-preserved millipede from the Cretaceous: LaeviglyphiUlUS patriCKMUelleri n. gen., n. sp. (Diplopoda, Spirostreptida, Cambalopsidae). Swiss Journal of Palaeontology (16) 144 (1): 1-8, DOI: 10.1186/s13358-025-00353-w, URL: https://doi.org/10.1186/s13358-025-00353-w
B84387E3FF92FFFAFF1AF9C173ABDB75.text	B84387E3FF92FFFAFF1AF9C173ABDB75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cambalidea Cook 1895	<div><p>Suborder Cambalidea Cook, 1895</p><p>Remarks: Our amber specimen is identified as a member of the extant suborder Cambalidea based on the following characters: Anterior and posterior gonopod well-developed (Fig. 3B–G); gnathochilarium divided into eumentum and promentum (Fig. 2E).</p></div>	https://treatment.plazi.org/id/B84387E3FF92FFFAFF1AF9C173ABDB75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Wesener, Thomas;RÜhr, Peter T.	Wesener, Thomas, RÜhr, Peter T. (2025): Internal gonopod reconstruction in an amber-preserved millipede from the Cretaceous: LaeviglyphiUlUS patriCKMUelleri n. gen., n. sp. (Diplopoda, Spirostreptida, Cambalopsidae). Swiss Journal of Palaeontology (16) 144 (1): 1-8, DOI: 10.1186/s13358-025-00353-w, URL: https://doi.org/10.1186/s13358-025-00353-w
B84387E3FF92FFFDFCA0F9E175FEDCE0.text	B84387E3FF92FFFDFCA0F9E175FEDCE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cambalopsidae Cook 1895	<div><p>Family Cambalopsidae Cook, 1895</p><p>(See figure on next page.)</p><p>Fig. 2 Laeviglyphiulus patrickmuelleri gen. nov., sp.nov., holotype, ZFMK-MYR6696 (BuB903), micro-CT scan reconstructions. A Anterior body, lateral view. B Anterior body,ventral view. C Anterior body, lateral view, with inner structures visible. D Anterior body, lateral view, with inner structures visible. E Head,ventral view. F Head lateral view. G Head, lateral view. H First leg pair, oral view. I. First leg pair, anal view. Numbers refer to body ring, podomere or antennomere number; Apo =apodeme; ca =cardines of mandible; Co = collum (body ring 1); cp =coxal process;gn =gnathochilarium; go =anterior and posterior gonopod; gu= gula; IL =incisura lateralis;L1 =leg pair 1; La = labrum; LL =lamellae linguales of gnathochilarium ;</p><p>Me =mentum PM= promentum; O =ommatidia (eyes);st =stipes of mandible; St = sternite with stigma openings; syn= syncoxite. Not to scale</p><p>Remarks: Our amber specimen is identified as a member of the extant family Cambalopsidae based on the following characters: ozopore starting at body ring 5</p><p>(Fig. 1C). Anterior gonopods without flagella (Fig. 3B–G) posterior gonopods reduced in size (Fig. 3B–G).</p></div>	https://treatment.plazi.org/id/B84387E3FF92FFFDFCA0F9E175FEDCE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Wesener, Thomas;RÜhr, Peter T.	Wesener, Thomas, RÜhr, Peter T. (2025): Internal gonopod reconstruction in an amber-preserved millipede from the Cretaceous: LaeviglyphiUlUS patriCKMUelleri n. gen., n. sp. (Diplopoda, Spirostreptida, Cambalopsidae). Swiss Journal of Palaeontology (16) 144 (1): 1-8, DOI: 10.1186/s13358-025-00353-w, URL: https://doi.org/10.1186/s13358-025-00353-w
B84387E3FF95FFFDFF1AFE747690DB00.text	B84387E3FF95FFFDFF1AFE747690DB00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laeviglyphiulus Wesener & RÜhr 2025	<div><p>Laeviglyphiulus gen. nov.</p><p>LSID urn:lsid:zoobank.org:act: 6EEBE818-77CD-43A7- A5AC-BDABEFED7AA1</p><p>Type species: Laeviglyphiulus patrickmuelleri sp. nov.</p><p>Etymology: Laeviglyphiulus, masculine, from laevis = smooth, in contrast to numerous other genera of Cambalopsidae with the presence of especially numerous carinae on the body segments.</p><p>Diagnosis: Laeviglyphiulus gen. nov. differs from the only other known member of the order from the Cretaceous, the Electrocambalidae, in the absence of setation of the tergites and head (Fig. 1A–D) (vs. present in the Electrocambalidae), as well as the absence of legs only on body ring 4 (Fig. 1C) (vs. body rings 3 and 4 in the Electrocambalidae), as well as other family-level characters (see above). Laeviglyphiulus gen. nov. shares gnathochilarium (Fig. 2E) and gonopod characteristics, in which the anterior gonopods form a flat shield-like structure (Fig. 3A–E), only with the common and diverse SE Asian extant species of the genera Hypocambala Silvestri, 1897, Plusioglyphiulus Silvestri, 1923 and Glyphiulus Gervais, 1842 . Te more complex gonopods of Laeviglyphiulus gen. nov. with a flagellum (Fig. 3A–E) are quite similar to the diverse SE Asian genus Plusioglyphiulus but also to some species of Hypocambala . Laeviglyphiulus gen. nov. differs from Plusioglyphiulus in the complete absence of any ornamentation (crests or tubercles) on the collum or other body rings (Figs. 1C,D, 2A–D). Laeviglyphiulus differs from all species of Hypocambala in the combination of absence of both tergal setation and collum crests, as well as a more strongly reduced male leg 1 (Fig. 2H, I). Te collum is also enlarged in Plusioglyphiulus, and more moderately so in the genus Glyphiulus and Hypocambala, but of normal size, if not even smaller than normal body rings (Figs. 1C, 2A–G), in Laeviglyphiulus gen. nov .. Laeviglyphiulus gen. nov. shares most characters with extant species of the morphologically diverse genus Hypocambala .</p></div>	https://treatment.plazi.org/id/B84387E3FF95FFFDFF1AFE747690DB00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Wesener, Thomas;RÜhr, Peter T.	Wesener, Thomas, RÜhr, Peter T. (2025): Internal gonopod reconstruction in an amber-preserved millipede from the Cretaceous: LaeviglyphiUlUS patriCKMUelleri n. gen., n. sp. (Diplopoda, Spirostreptida, Cambalopsidae). Swiss Journal of Palaeontology (16) 144 (1): 1-8, DOI: 10.1186/s13358-025-00353-w, URL: https://doi.org/10.1186/s13358-025-00353-w
B84387E3FF95FFFEFF1AF95476F1DF20.text	B84387E3FF95FFFEFF1AF95476F1DF20.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laeviglyphiulus patrickmuelleri Wesener & RÜhr 2025	<div><p>Laeviglyphiulus patrickmuelleri sp. nov.</p><p>LSID urn:lsid:zoobank.org:act:BFB20F5C-3A16-4F4F-93DBB282A113CB56</p><p>Material examined: Holotype: 1 male (ZFMK MYR6696). From the collection of Patrick Müller, Myanmar, Kachin State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.566666&amp;materialsCitation.latitude=26.25" title="Search Plazi for locations around (long 96.566666/lat 26.25)">Hukawng Valley</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.566666&amp;materialsCitation.latitude=26.25" title="Search Plazi for locations around (long 96.566666/lat 26.25)">Noije Bum</a> amber mine, N 26° 15 ′, E 96° 34 ′. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=96.566666&amp;materialsCitation.latitude=26.25" title="Search Plazi for locations around (long 96.566666/lat 26.25)">Transferred</a> to the ZFMK in 2017.</p><p>Amber description: Amber flat, semi-circle, 19.6 mm × 18.8 mm × 3.5 mm, polished, yellow, clear transparent.</p><p>Syninclusions: Remains of a broken up immature Blattodea, soil particles, faecal pellets, 1 very small Staphylinidae Coleoptera, 1 small Curculionidae Coleoptera</p><p>(Fig. 1A, B).</p><p>Derivation of name: patrickmuelleri, noun in apposition, after the amber collector Patrick Müller, who not only opened his large collection for scientific study, but also agreed to donate the unique specimen described here to a public Museum.</p><p>Diagnosis: See genus diagnosis as the genus is currently monotypic. Small-bodied (10 mm long), thin (0.7 mm wide) juliform millipede with 41 body rings and one apodous ring. Tergites not ornamented, but ventral part of pleurotergite with seven or eight prominent longitudinal striae. Gula of gnathochilarium undivided. Eyes well-developed with 24 ommatidia in well-defined rows. Cardo of mandible much shorter than prominent stipes, similar to Choctella Chamberlin, 1918 ( Cambalidea, Choctellidae). First leg pair in males strongly reduced, with short, uncate process, telopodite consisting of four reduced podomeres. Legs 1 and 2 lacking an apical spine.</p><p>Description:</p><p>Ca. 10 mm long, 0.6–0.7 mm wide. 41 + 1 body rings.</p><p>Head: Eyes 24 ommatidia in 5 well-defined rows</p><p>(Figs. 1C, 2A, C, G). Apical half of antennae damaged or missing. Gnathochilarium mentum divided into eumentum and promentum (Fig. 2E). Mandible with very prominent stipes, cardo much smaller and almost hidden from lateral view (Fig. 2A–G).</p><p>Body rings: Collum small, surface smooth, no striae. lateral edges well-rounded, ending well below body ring 2 (Figs. 1C, 2A–D). Pro- and metazona of body rings ventrally with 7 or 8 strong longitudinal ridges (Fig. 2A–D). Ozopores starting at body ring 5. Surface of body-rings smooth except for longitudinal striae on pleurotergite</p><p>(Fig. 2A–D).</p><p>Legs: Leg 1 greatly reduced to a fused coxite with a telopodite consisting of four cylindrical podomeres (Fig. 2H, I). Coxae each with a wide, apically tapering triangular process, process as long as podomere 1. Leg 2 flattened and shorter than following ones, tarsus with a claw, no tarsal spine. Legs 3 and onwards elongated, much longer than width of body rings. A single ventral apical spine present on each podomere, and two pair of long ventral spines on tarsus visible in some legs.</p><p>Telson: Preanal ring well-rounded, not projecting. Anal valves narrow, not located posteriorly but ventrally of the preanal scale. Anal lips not visible, valves glabrous but with micropunctation (Fig. 1D). Subanal scale inconspicuous.</p><p>Gonopods: Anterior gonopod with well-visible sternite</p><p>(Fig. 3B, C, E). Coxite massive, shield-like. Coxite process short, well rounded (Fig. 3A–F). Telopodite digitiform, located laterally of coxite and not projecting beyond</p><p>(Fig. 3A–F). Posterior gonopod less well-preserved, separated into a basal coxite and apical telopodite (Fig. 3D– F). Telopodite with a visible start of a free flagellum clearly visible (Fig 3D–F). Coxite apically with 2 bumps</p><p>(Fig. 3D). Telopodite apico-laterally with a swollen retrorse tip (Fig. 3D, F), and more laterally with a short ledge below flagellum (Fig. 3D).</p></div>	https://treatment.plazi.org/id/B84387E3FF95FFFEFF1AF95476F1DF20	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Wesener, Thomas;RÜhr, Peter T.	Wesener, Thomas, RÜhr, Peter T. (2025): Internal gonopod reconstruction in an amber-preserved millipede from the Cretaceous: LaeviglyphiUlUS patriCKMUelleri n. gen., n. sp. (Diplopoda, Spirostreptida, Cambalopsidae). Swiss Journal of Palaeontology (16) 144 (1): 1-8, DOI: 10.1186/s13358-025-00353-w, URL: https://doi.org/10.1186/s13358-025-00353-w
