identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
BD242A8D21B85751AEAB81C2289F235E.text	BD242A8D21B85751AEAB81C2289F235E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belobranchus belobranchus (Valenciennes 1837)	<div><p>Belobranchus belobranchus (Valenciennes, 1837)</p><p>Figs 1, 2, 3 A – D, 5 A, 6, Tables 1, 2 English name: Throat-spine Gudgeon</p><p>Eleotris belobrancha Valenciennes in Cuvier and Valenciennes 1837: 243 (type locality: Manado Island, Sulawesi, Indonesia)</p><p>Belobranchus quoyi Bleeker, 1856: 300 (type locality: Manado Island, Sulawesi, Indonesia)</p><p>Belobranchus taeniopterus Bleeker, 1856: 301 (type locality: Boleling, Bali, Indonesia)</p><p>Material examined.</p><p>Non-type specimens. 94 specimens, all specimens from   Indonesia. Java: • MZB.16307, 84.6 mm SL, male,  Cibareno, Pasir Baru, Cisolok, Sukabumi, Jawa Barat, S. Sauri et al., 23 Jun. 2002  •   MZB.16328, 63.1 mm SL, female,  Cilumajan, Pasir Baru, Cisolok, Sukabumi, Jawa Barat, S. Sauri et al., 24 May 2002  •   MZB.26872 [ex BIF.01651], 71.7 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.395&amp;materialsCitation.latitude=-6.963333" title="Search Plazi for locations around (long 105.395/lat -6.963333)">Cibareno</a>, Lebak, Banten, 6°57'48.0"S, 105°23'42.0"E, N. Hubert et al., 10 Dec. 2013  •   MZB.26873 [ex BIF.1696], 77.5, female, MZB.26906 [ex BIF 01694], 58.7 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=106.446&amp;materialsCitation.latitude=-6.946111" title="Search Plazi for locations around (long 106.446/lat -6.946111)">Citiis</a>, Sukabumi, Jawa Barat, 6°56'46.0"S, 106°26'45.6"E, N. Hubert et al., 11 Dec. 2013  •   MZB.23934, 33.9 mm SL, male,  Purworejo, Jawa Tengah, R. Hadiaty, 10 Sep. 2017  .   Bali: • MZB.26874 [ex BIF.2385], 74.8 mm SL, male, MZB.26907 [ex BIF.2384], 65.0 mm SL, female, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.752&amp;materialsCitation.latitude=-8.367001" title="Search Plazi for locations around (long 114.752/lat -8.367001)">Nbang</a>, Jembrana, Bali, 8°22'01.2"S, 114°45'07.2"E, N. Hubert et al., 15 Apr. 2014  .   Sulawesi: • MZB.4795, 67.4–106.9 mm SL, 2 specimens, male and female,  Watu Songu, Ulubongka, Poso, Sulawesi Tengah, Suyanto, 10 Nov. 1982  •   MZB.11665, 70.1–108.6 mm SL, 3, 1 male, 2 females, MZB.11676,  Lambuno, Wana Mukti, Mountong, Donggala, Sulawesi Tengah, Agus, 3 May 2002  •  MZB.11715, 45.4–52.1 mm SL, 6 specimens, 2 males, 4 females, Haryono and A. Munim, 8 Aug. 2001 •   MZB.11731, 66.1–100.2 mm SL, 8 specimens, 2 males, 6 females,  Wapo river, Lombongo, Suwawa, Gorontalo, Haryono and A. Munim, 7 Aug. 2001  •   MZB.11735, 57.3–65.2 mm SL, 4 specimens, 3 males, 1 female,  Tolumolu, Kota Selatan, Sulawesi Utara, 8 Aug. 2001  •   MZB.11964, 109.5 mm SL, female, MZB.11975, 113.0 mm SL, female, MZB.12048, 62.3 mm SL, female,  Bolaang Mongondow, Sulawesi Utara, Haryono and Hesron, 21–26 May 2002  •   MZB.20004, 58.6–77.1 mm SL, 3 specimens, 1 male, 2 females,  Masembo,  Mekongga, Tinukari, Wawo, Kolaka Utara, Sulawesi Tenggara, Jumaring, 30 Nov. 2010  •   MZB.20352, 36.4–115.4 mm SL, 17 specimens, 11 males, 6 females, MZB.20381, 52.9–96.2 mm SL, 9 specimens, 5 males, 4 females, MZB.20391, 78.9 mm SL, female, MZB.20398, 107.0– 129.5 mm SL, 3 specimens, 2 males, 1 female, MZB.20406, 58.5–63.8 mm SL, 3 specimens, 1 male, 2 females, MZB.20415, 51.0– 100.3 mm SL, 3 males,  Tinukari, Wawo, Kolaka Utara, Sulawesi Tenggara, R. Hadiaty et al., 28 Jun. – 3 Jul. 2011  •   MZB.22703, 84.1 mm SL, male,  Bonepuso, Bulagi Selatan, Banggai Kepulauan, S. Sauri, 20 Sep. 2014  •   MZB.24579, 101.4 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.26508&amp;materialsCitation.latitude=-3.3620832" title="Search Plazi for locations around (long 120.26508/lat -3.3620832)">Marinding</a>, Luwu, Sulawesi Selatan, 3°21'43.5"S, 120°15'54.3"E, seine net, D. Lumbantobing, 12 Aug. 2016  .   Alor: • MZB.5346, 115.3 mm SL, male,  Alor, Nusa Tenggara Timur, Sudarmanu, Apr. 1984  .   Halmahera: • MZB.18812, 85.3 mm SL, male, MZB.18830, 61.3–79.7 mm SL, 3 males, MZB.21174, 32.8 mm SL, male, MZB.21184, 93.4 mm SL, female, MZB.21209, 71.8–98.4 mm SL, 5 specimens, 1 male, 4 females, MZB.21219, 58.2 mm SL, female, MZB.26661 [ex MZB.18747], 63.3–130.7 mm SL, 7 specimens, 1 male, 6 females, MZB.26662 [ex MZB.18820], 71.8 mm SL, female, Ake Jira, Weda Tengah, R. Hadiaty et al., 4 Feb. 2010 – 16 Jul. 2011  .   Papua: • MZB.15631, 95.2 mm SL, female,  Waigeo, Raja Ampat, Papua, R. Hadiaty et al., 4 Jun. 2007  •   MZB.17098, 65.3–97.6 mm SL, 2 males, MZB.17118, 59.7–61.0 mm SL, male and female, MZB.17140, 91.9 mm SL, male,  Batanta, Raja Ampat, Papua Barat, R. Hadiaty et al., 30 July – 5 May 2008  .</p><p>Diagnosis.</p><p>A species of  Belobranchus with the following combination of characters: head relatively depressed; interorbital width 4.8–8.0 % (mean 6.0 %) of SL; jaw length 10.3–16.7 % (14.1 %) of SL; caudal-peduncle depth 11.9–15.0 % (13.6 %) of SL; body with 3 dark-brown bands seaparated by pale bands, with many dark horizontal lines (one per scale row) along lateral surface; dorsal and lateral surfaces of head with many small whitish spots; 1 or 2 distinct oblique brown lines extending from eye to posterior edge of operculum (not always visible); first dorsal fin typically uniformly mottled dark brown or with thin yellowish distal edge; largest recorded specimen 130.7 mm SL.</p><p>Distribution and habitat.</p><p>Belobranchus belobranchus has been recorded from the South China Sea (Randall and Lim 2000), Indonesia: Java, Bali, Lombok, Flores, Alor, Timor, Buton, Sulawesi, and Halmahera islands, and West Papua (Roberts 1993; Tjakrawidjaja 2002; Larson and Pidgeon 2004; Larson et al. 2007; Keith et al. 2012; Tweedley et al. 2013; Dahruddin et al. 2016; Miesen et al. 2016; this study); the Philippines (Jamandre 2023); Japan (Sakai et al. 2001; Nakabo 2002); Papua New Guinea (Fricke et al. 2014; Amick and Toko 2021) (Fig. 6).  Belobranchus belobranchus is an amphidromous species, characterized by migration between freshwater and coastal environments (Keith et al. 2012; Sahami et al. 2019); its habitat predominantly encompasses the lower regions of rivers, typically at altitudes to 5 m, and it has been observed in muddy to clear river systems featuring sandy to rocky or gravel bottoms (Parenti 2021). Like most eleotrids, this fish is carnivorous (Kiruba-Sankar et al. 2018).</p></div>	https://treatment.plazi.org/id/BD242A8D21B85751AEAB81C2289F235E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Afrisal, Muhammad;Nurjirana, Nurjirana;Haryono, Haryono;Mokodongan, Daniel Frikli;Wibowo, Kunto;Moore, Abigail Mary;Hubert, Nicolas	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
534C89B20A955B0D858A50231C77CE62.text	534C89B20A955B0D858A50231C77CE62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belobranchus segura Keith, Hadiaty & Lord 2012	<div><p>Belobranchus segura Keith, Hadiaty &amp; Lord, 2012</p><p>Figs 1, 2, 3 E – H, 4, 5 B, 6, Tables 1, 2 New English name: Segura Gudgeon</p><p>Belobranchus segura Keith, Hadiaty &amp; Lord, 2012: 480, figs 1, 2, 3 A, 3 C (type locality: Ake Jira, Leililef Waibulen, Halmahera, Indonesia)</p><p>Material examined.</p><p>Holotype. MZB.20786, 72.1 mm SL, male,  Ake Jira, Leililef Waibulen, Halmahera, Indonesia, R. Hadiaty et al., 26 Jan. 2010.</p><p>Paratypes. 9 specimens, 40–72 mm SL, all specimens collected from Maluku Island, Indonesia . •   MZB.18658, 61–66 mm SL, 2 males,  Ake Kobe, Halmahera, R. Hadiaty et al., 23 Jan. 2010  •  MZB.18684, 64–72 mm SL, 3 males, same data as holotype •   MZB.18747, 55 mm SL, female, MZB.18751, 40–44 mm SL, 2 females, MZB.18820, 44 mm SL, female,  Ake Jira, Weda Tengah, Halmahera, R. Hadiaty et al., 4–7 Feb. 2010  .</p><p>Non-type materials.</p><p>78 specimens, 33.4–79.8 mm SL, all specimens collected from   Indonesia. Java: • MZB.23934, 33.9 mm SL, male,  Purworejo, Jawa Tengah, R. Hadiaty, 10 Sep. 2017  •   MZB.26667 (ex BIF.1483), 42.1 mm SL, male, MZB.26890 [ex BIF.1482], 39.1 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.630005&amp;materialsCitation.latitude=-6.591389" title="Search Plazi for locations around (long 105.630005/lat -6.591389)">Cibeber</a>, Pandeglang, Banten, 6°35'29.0"S, 105°37'48.0"E, N. Hubert et al., 7 Dec. 2013  .   Bali: • MZB.26668 (ex BIF.2389), 56.3 mm SL, female, MZB.26891 (ex BIF.2386), 50.2 mm SL, female, MZB.26892 (ex BIF.2374), 55.1 mm SL, male, MZB.26893 (ex BIF.2375), 61.5 mm SL, male, MZB.26894 (ex BIF.2387), 52.6 mm SL, female, MZB.26895 (ex BIF.2373), 59.0 mm SL, male, MZB.26896 (ex BIF.2376), 60.8 mm SL, male, MZB.26897 (ex BIF.2388), 35.9 mm SL, female, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.752&amp;materialsCitation.latitude=-8.367001" title="Search Plazi for locations around (long 114.752/lat -8.367001)">Nbang</a>, Jembrana, Bali, 8°22'01.2"S, 114°45'07.2"E, N. Hubert et al., 15 Apr. 2014  •   MZB.26670 (ex BIF.2460), 52.0 mm SL, female, MZB.26898 (ex BIF.2457), 60.3 mm SL, male, MZB.26899 (ex BIF.2456), 65.1 mm SL, male, MZB.26900 (ex BIF.2459), 55.3 mm SL, male, MZB.26901 (ex BIF.2454), 69.2 mm SL, male, MZB.26902 (ex BIF.2455), 68.9 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=114.784&amp;materialsCitation.latitude=-8.360001" title="Search Plazi for locations around (long 114.784/lat -8.360001)">Yeh Sumbul</a>, Jembrana, Bali, 8°21'36.0"S, 114°47'02.4"E, N. Hubert et al., 16 Apr. 2014  .   Sulawesi: • MZB.11715, 41.3–52.1 mm SL, 6 specimens, 2 males, 4 females, MZB.11717, 46.1–68.0 mm SL, 6 specimens, 3 males, 3 females, MZB.11727, 37.8–62.2 mm SL, 6 specimens, 4 males, 2 females,  Baliohuto, Lombongo, Suwawa, Gorontalo, Haryono and A. Munim, 6–8 Aug. 2001  •   MZB.12048, 62.3 mm SL, female,  Molong, Pusian Dumoga, Bolaang Mongondow, Sulawesi Utara, Haryono and Hesron, 26 May 2002  •   MZB.26663 (ex MZB.11676), 47.3–69.4 mm SL, 11 specimens, 6 males, 5 females,  Mountong, Donggala, Sulawesi Tengah, Agus, 2 May 2002  •   MZB.26664 (ex MZB.1197), 61.2–76.1 mm SL, male and female,  Bolaang Mongondow, Sulawesi Utara, Haryono and Hesron, 22 May 2002  •   MZB.26669 (ex MZB.22703), 49.1–50, 4 mm SL, male and female,  Bonepuso, Bulagi Selatan, Banggai Kepulauan, S. Sauri, 20 Sep 2014  .   Lombok: • MZB.22933, 42.0– 50.5 mm SL, 3 females,  Ormori,  Kawinda Toi, Tambora, Sumbawa, Nusa Tenggara Barat, Mulyadi, 24 Apr. 2015  .   Flores: • MZB.6359, 43.4–53.5 mm SL, 4 specimens, 2 males, 2 females,  Wae Laku, Ruteng, Flores, Nusa Tenggara Timur, Agus and Munir, 26 May 1994  .   Timor: • MZB.26903, 59.6 mm SL, male, MZB.26904, 59.6 mm SL, male, MZB.26905, 57.8 mm SL, male, MZB.26906, 44.4 mm SL, male, MZB.26908, 66.1 mm SL, male, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=124.88117&amp;materialsCitation.latitude=-9.011611" title="Search Plazi for locations around (long 124.88117/lat -9.011611)">Jenilu</a>, Kakuluk Mesak, Belu, Nusa Tenggara Timur, 9°00'41.8"S, 124°52'52.2"E, M. Afrisal et al., 23 Jan. 2023  .   Halmahera: • MZB.18700, 67.2 mm SL, male, MZB.18706, 76.4–79.8 mm SL, 3 males, MZB.18711, 56.7 mm SL, female, MZB.26909 (ex MZB.18820), 59.0– 58.1 mm SL, 2 females,  Ake Jira, Weda Tengah, R. Hadiaty et al., 27 Jan. – 7 Feb. 2010  •   MZB.21219, 58.2 mm SL, female,  Ake Kobe, Weda Tengah, R. Hadiaty et al., 7 Jul. 2011  •   MZB.26665 (ex MZB.18747), 38.4–43.8 mm SL, 2 females, MZB.26666 (ex MZB.18751), 33.4–43.0 mm SL, 4 females,  Ake Saki, Weda Tengah, R. Hadiaty et al., 7 Feb. 2010  .</p><p>Diagnosis.</p><p>A species of  Belobranchus with the following combination of characters: head somewhat convex; interorbital width 6.4–11.6 % (mean 8.2 %) of SL; jaw length 8.7–13.5 % (11.1 %) of SL; caudal-peduncle depth 13.2–16.6 % (14.7 %) of SL; body with 4 or 5 dark-brown bands separated by narrow pale bars, lacking dark horizontal lines; dorsal and lateral surfaces of head without small whitish spots; 3 or 4 oblique brown lines extending from eye to posterior edge of operculum; upper margin of first dorsal fin yellowish to orange; largest recorded specimen 79.8 mm SL.</p><p>Distribution and habitat.</p><p>Belobranchus segura has been recorded from Indonesia: Sumatra, Java, Bali, Lombok, Sulawesi, Halmahera, Sumbawa, Flores, Timor, Papua Barat (Kottelat et al. 1993; Keith et al. 2012; Dahruddin et al. 2016; Miesen et al. 2016; Sahami and Habibie 2021; Hasan et al. 2023; this study), the Philippines (Vedra et al. 2013), Japan (Suzuki et al. 1995), Papua New Guinea (Amick and Toko 2021: fig. 2 P, Q), Solomon Islands and Vanuatu (Keith et al. 2012), and Philippines (Gonzalez et al. 2023) (Fig. 6). This species is considered amphidromous (Keith et al. 2012; Sahami and Habibie 2021) and has been found in the lower reaches of muddy to clear rivers with sandy to gravel substrate, to 5 m in altitude, close to estuaries. Like most eleotrids, this fish is considered carnivorous (Kiruba-Sankar et al. 2018).</p></div>	https://treatment.plazi.org/id/534C89B20A955B0D858A50231C77CE62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Afrisal, Muhammad;Nurjirana, Nurjirana;Haryono, Haryono;Mokodongan, Daniel Frikli;Wibowo, Kunto;Moore, Abigail Mary;Hubert, Nicolas	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
