taxonID	type	description	language	source
BD242A8D21B85751AEAB81C2289F235E.taxon	description	Figs 1, 2, 3 A – D, 5 A, 6, Tables 1, 2 English name: Throat-spine Gudgeon	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
BD242A8D21B85751AEAB81C2289F235E.taxon	diagnosis	Diagnosis. A species of Belobranchus with the following combination of characters: head relatively depressed; interorbital width 4.8 – 8.0 % (mean 6.0 %) of SL; jaw length 10.3 – 16.7 % (14.1 %) of SL; caudal-peduncle depth 11.9 – 15.0 % (13.6 %) of SL; body with 3 dark-brown bands seaparated by pale bands, with many dark horizontal lines (one per scale row) along lateral surface; dorsal and lateral surfaces of head with many small whitish spots; 1 or 2 distinct oblique brown lines extending from eye to posterior edge of operculum (not always visible); first dorsal fin typically uniformly mottled dark brown or with thin yellowish distal edge; largest recorded specimen 130.7 mm SL.	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
BD242A8D21B85751AEAB81C2289F235E.taxon	distribution	Distribution and habitat. Belobranchus belobranchus has been recorded from the South China Sea (Randall and Lim 2000), Indonesia: Java, Bali, Lombok, Flores, Alor, Timor, Buton, Sulawesi, and Halmahera islands, and West Papua (Roberts 1993; Tjakrawidjaja 2002; Larson and Pidgeon 2004; Larson et al. 2007; Keith et al. 2012; Tweedley et al. 2013; Dahruddin et al. 2016; Miesen et al. 2016; this study); the Philippines (Jamandre 2023); Japan (Sakai et al. 2001; Nakabo 2002); Papua New Guinea (Fricke et al. 2014; Amick and Toko 2021) (Fig. 6). Belobranchus belobranchus is an amphidromous species, characterized by migration between freshwater and coastal environments (Keith et al. 2012; Sahami et al. 2019); its habitat predominantly encompasses the lower regions of rivers, typically at altitudes to 5 m, and it has been observed in muddy to clear river systems featuring sandy to rocky or gravel bottoms (Parenti 2021). Like most eleotrids, this fish is carnivorous (Kiruba-Sankar et al. 2018).	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
534C89B20A955B0D858A50231C77CE62.taxon	description	Figs 1, 2, 3 E – H, 4, 5 B, 6, Tables 1, 2 New English name: Segura Gudgeon	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
534C89B20A955B0D858A50231C77CE62.taxon	diagnosis	Diagnosis. A species of Belobranchus with the following combination of characters: head somewhat convex; interorbital width 6.4 – 11.6 % (mean 8.2 %) of SL; jaw length 8.7 – 13.5 % (11.1 %) of SL; caudal-peduncle depth 13.2 – 16.6 % (14.7 %) of SL; body with 4 or 5 dark-brown bands separated by narrow pale bars, lacking dark horizontal lines; dorsal and lateral surfaces of head without small whitish spots; 3 or 4 oblique brown lines extending from eye to posterior edge of operculum; upper margin of first dorsal fin yellowish to orange; largest recorded specimen 79.8 mm SL.	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
534C89B20A955B0D858A50231C77CE62.taxon	distribution	Distribution and habitat. Belobranchus segura has been recorded from Indonesia: Sumatra, Java, Bali, Lombok, Sulawesi, Halmahera, Sumbawa, Flores, Timor, Papua Barat (Kottelat et al. 1993; Keith et al. 2012; Dahruddin et al. 2016; Miesen et al. 2016; Sahami and Habibie 2021; Hasan et al. 2023; this study), the Philippines (Vedra et al. 2013), Japan (Suzuki et al. 1995), Papua New Guinea (Amick and Toko 2021: fig. 2 P, Q), Solomon Islands and Vanuatu (Keith et al. 2012), and Philippines (Gonzalez et al. 2023) (Fig. 6). This species is considered amphidromous (Keith et al. 2012; Sahami and Habibie 2021) and has been found in the lower reaches of muddy to clear rivers with sandy to gravel substrate, to 5 m in altitude, close to estuaries. Like most eleotrids, this fish is considered carnivorous (Kiruba-Sankar et al. 2018).	en	Afrisal, Muhammad, Nurjirana, Nurjirana, Haryono, Haryono, Mokodongan, Daniel Frikli, Wibowo, Kunto, Moore, Abigail Mary, Hubert, Nicolas (2025): Revised diagnoses of the gudgeons Belobranchus belobranchus and B. segura (Actinopterygii, Gobiiformes, Eleotridae). ZooKeys 1232: 173-186, DOI: 10.3897/zookeys.1232.141880
