taxonID	type	description	language	source
CB3687DDFFA5FFA1FF4CF8E2FAC71865.taxon	description	urn: lsid: zoobank. org: act: E 5 FAC 79 E- 0647 - 404 E- 92 F 5 - 4577429 C 98 AD	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA5FFA1FF4CF8E2FAC71865.taxon	type_taxon	Type and only species. Pseudoameropterus ambiguus gen. et sp. nov.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA5FFA1FF4CF8E2FAC71865.taxon	etymology	Etymology. From the Greek pseudos, false, and Ameropterus, a genus-group name of Ascalaphidae. Gender masculine.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA5FFA1FF4CF8E2FAC71865.taxon	diagnosis	Diagnosis. May be distinguished from other genera by a set of the following hind wing characters: RP 1 profusely branched; hypostigmal cell relatively long; MP and CuA spaces relatively narrow; CuA long and incurved; branches of CuA and MP interlinked by only one row of crossveins.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA5FFA1FF4CF8E2FAC71865.taxon	discussion	Remarks. The status of Ascalaphidae as a family is an area of debate. Many studies based on morphology only have recovered a monophyletic Ascalaphidae closely related to Myrmeleontidae (e. g., Aspöck et al. 2001; Aspöck & Aspöck 2008). However, studies that have included molecular data in addition to morphology have recovered conflicting trees supporting monophyly of both Ascalaphidae and Myrmeleontidae, or showing a paraphyletic relationship of Ascalaphidae with respect to Myrmeleontidae (see Machado et al. 2018; Winterton et al. 2018; Jones 2019 for review). Recently, phylogenomic studies by Machado et al. (2018) and Winterton et al. (2018) recovered Ascalaphidae and Myrmeleontidae as being paraphyletic with respect to each other, this prompted Machado et al. (2018) to synonymize Ascalaphidae with Myrmeleontidae, dividing the family into four subfamilies and 17 tribes. But the family status of Ascalaphidae is still debated (Jones 2019; Cai et al. 2023). Here we follow Cai et al. (2023) and consider Ascalaphidae to be retained as a family with the inclusion of Stilbopteryginae as a subfamily. The long and more or less strongly incurved CuA in the hind wing as found in Pseudoameropterus gen. nov. is present in several genera of Ascalaphidae: all genera of Stilbopteryginae (Stilbopteryx Newman, 1838 and Aeropteryx Riek, 1968) and Albardiinae (Albardia van der Weele, 1903), and most genera of Ululodinae (i. e., Ameropterus Esben-Petersen, 1922, Cordulecerus Rambur, 1842 and Ululodes Currie in Smith, 1900). Interestingly, all these taxa are now distributed in the Americas (Albardiinae, Ululodinae) or Australia (Stilbopteryginae). The hind wing venation of the new genus is most similar to that of Ameropterus, especially e. g. A. trivialis (Gerstaecker, 1888) and A. mexicanus (van der Weele, 1908) (see Penny 2002: Figs 14, 16). The genus is distributed from southern Mexico in the north to Argentina in the south. However, Pseudoameropterus gen. nov. differs from that genus in that RP 1 is more proximally and profusely branched, which is branched more distally in Ameropterus (see van der Weele 1909: Figs 83 – 85; Penny 2002: Figs 12 – 15). All species of Cordulecerus (except Cordulecerus praecellens (Gerstaecker, 1885 )) differ from Pseudoameropterus ambiguus sp. nov. by the hind wings being strongly broadened proximally (see van der Weele 1909: Figs 98 – 112, 115, 116; Penny 1982: Fig. 11; Ardila-Camacho et al. 2019: Fig. 9). The species of Ululodes differ from the new species by the presence of a longitudinal crossvein connecting crossveins between RA / Sc + RA and RP below the pterostigma (see van der Weele 1909: Figs 59 – 68, 73 – 76; Penny 2002: Figs 22 – 26). However, it is possible that the new genus belongs to Stilbopteryginae or Albardiinae. The hind wing venation of Aeropteryx Riek, 1968 (Stilbopteryginae) and Pseudoameropterus gen. nov. are very similar, including the profusely branched RP 1 (see Riek 1968: Pl. 1). Moreover, a new undescribed ascalaphid species from the Green River Formation, the venation of which is most similar to that of Pseudoameropterus ambiguus sp. nov., possesses entire eyes characteristic of Stilbopteryginae and Albardiinae, while eyes are divided in Ululodinae. But the true subfamilial affinity of Pseudoameropterus gen. nov. may only be determined when more complete material is found.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA6FFA0FF4CFA55FDCE1CE1.taxon	description	urn: lsid: zoobank. org: act: 4 A 61 DA 82 - 99 EF- 410 B-BADE- 94 C 6 D 728246 F Fig. 1	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA6FFA0FF4CFA55FDCE1CE1.taxon	etymology	Etymology. From the Latin ambiguus [- a, - um], ambiguous, obscure, mysterious.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA6FFA0FF4CFA55FDCE1CE1.taxon	materials_examined	Type material. Holotype USNM PAL 626076, deposited in USNM. A nearly complete hind wing. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Lutetian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA6FFA0FF4CFA55FDCE1CE1.taxon	diagnosis	Diagnosis. As for genus.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA6FFA0FF4CFA55FDCE1CE1.taxon	description	Description. Hind wing 30.3 mm long as preserved (estimated complete length 35 – 37 mm), 8.1 mm wide. Costal space moderately broad, with all veinlets simple, closely spaced. Pterostigma indistinct, short, bound by two simple veinlets, two incorporated veins, one multibranched, one simple. Sc fused with RA in distal part of wing level with centre of the pterostigma. Seven preserved veinlets of Sc + RA connected by 10 preserved crossveins; four proximal veinlets simple or forked once, three distal veinlets strongly forked. Thirteen ra-rp crossveins preserved. RP with seven branches. RP 1 originating before mid-point of wing, profusely branched at mid-point of vein. Other branches of RP probably shallowly branched (their terminal parts not preserved). Numerous crossveins (66 preserved) throughout RP spaces. One presectoral crossvein preserved. Between R and M (radiomedial space) 17 crossveins preserved. MA slightly incurved medially, simple. MP terminally simple, stronger incurve medially than MA, pectinately branched with four branches, one of these (MP 1) rather shallowly forked, other simple. Stem of MP and its branches connected by four crossveins (one between each pair). In intramedial space (between MA and MP) 15 crossveins preserved. Between MP and CuA (mediocubital space) nine preserved widely spaced crossveins. CuA long, strongly incurved; stem of CuA shallowly forked terminally, with eight pectinate branches; one of these rather shallowly forked, other simple. Stem of CuA and three distal branches connected by three crossveins (one between each pair). One cua-cup crossvein preserved in intracubital space. CuP probably rather long, fragmentarily preserved. Anal veins not preserved.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA7FFA7FF4CF9C5FA721DC9.taxon	description	urn: lsid: zoobank. org: act: 1 A 315754 - A 548 - 4 BB 7 - 9739 - 6 A 3 C 2 E 948 EA 7	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA7FFA7FF4CF9C5FA721DC9.taxon	type_taxon	Type and only species. Minimochrysa latialata sp. nov.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA7FFA7FF4CF9C5FA721DC9.taxon	etymology	Etymology. From the Latin minimus, - a, - um, tiny, smallest, and Chrysopa, a genus-group name, in reference to smallest chrysopid genus. Gender feminine.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA7FFA7FF4CF9C5FA721DC9.taxon	diagnosis	Diagnosis. Differs from other nothochrysine genera by combination of following forewing character states: extremely short (≤ 5 mm); broadly-rounded apically; distal branches of RP originating at almost right angles; RA almost reaching wing apex.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA7FFA7FF4CF9C5FA721DC9.taxon	discussion	Remarks. The Nothochrysinae affinity is based on a very long RA and long Sc. In Chrysopinae, RA is much shorter, terminating at C well proximad of wing apex. In Limaiinae, Sc is much shorter than RA. The venation of Apochrysinae is very different from that of this specimen. In most other genera of Nothochrysinae, the forewing is apically subacute or at least has a more oval apex, and the distal branches of RP originate more obliquely. In those genera that have a similar broadly-rounded forewing apex and distal branches of RP originating at almost right angles (e. g., Kimachrysa Tjeder, 1966), their RA is very short.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA0FFA6FF4CFECCFA771FFD.taxon	description	urn: lsid: zoobank. org: act: 36 FE 8 A 47 - EF 3 A- 4718 - 8374 - 1 DE 02063 E 5 FE Fig. 2	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA0FFA6FF4CFECCFA771FFD.taxon	materials_examined	Type material. Holotype USNM PAL 622725, deposited in USNM. A distal part of a forewing. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Lutetian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA0FFA6FF4CFECCFA771FFD.taxon	etymology	Etymology. From the Latin latus, broad, and ala, wing, in reference to broad distal part of the forewing (broadly-rounded apically).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA0FFA6FF4CFECCFA771FFD.taxon	description	Description. Forewing 2.5 mm long as preserved (estimated complete length ca. 4.5 – 5.0 mm), 1.4 mm wide as preserved. Forewing broadly rounded apically. Costal space with 12 preserved simple crossveins, narrowing distally. No crossveins present in subcostal space. Sc long terminating well before apex of wing. RA long almost reaching apex of the wing. Radial space wide with eight regularly spaced ra-rp crossveins preserved. RP with eight preserved branches, majority simple, four forked distally near wing margin. RP crossveins forming two gradate series. Basal portion of wing not preserved.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA0FFA6FF4CFECCFA771FFD.taxon	discussion	Remarks. Minimochrysa latialata sp. nov. is the smallest known chrysopid species. The species of the smallest extant genera have a minimum forewing length of 6 mm, e. g., the chrysopine Suarius Navás, 1914 and the nothochrysine Kimochrysa Tjeder, 1966 (Brooks & Barnard 1990). The smallest known fossil chrysopid species Pseudochrysopa harveyi Makarkin & Archibald, 2013 from the early Eocene Driftwood Canyon (Canada) with a minimum forewing length of ca. 8.2 mm is nearly twice as long as the new species (Makarkin & Archibald 2013).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA6FF4CFCE5FE5518A1.taxon	discussion	Remarks. The genus Palaeochrysa was recently revised to include only one species P. stricta Scudder, 1890 from the late Eocene of Florissant (Makarkin et al. 2022). Two more species in this genus are described below from the Kishenehn Formation. Makarkin et al. (2022) diagnosed the genus with the following characters: in the fore- and hind wings (1) two gradate series are present in the radial space; (2) inner gradate series is arranged in a strongly broken line; (3) Psm is nearly straight, slightly zigzagged; (4) Psm is formed by five branches of RP; in the forewing (5) im is elongate; and in the hind wing (6) three branches of CuA proximad fusion with MP. Two of the species (P. stricta and P. greenwalti sp. nov.) share the whole set of these character states. The character states (2) and (4) are slightly different in P. minor sp. nov.: in this species the inner gradate series is arranged in a curved line, but not broken, and Psm is formed by three branches of RP (most likely related to its small size). We found a new feature distinguishing Palaeochrysa from most other genera of Nothochrysinae. This genus possesses a proximal cell between two gradate series of crossveins in the hind wing (located at the termination of Psm; labelled gc 1 in Figs. 3 C and 4 E). The cell is acute anteriorly as the very oblique crossvein of the inner series and a part of a RP branch (which are limited the cell anteriorly) coincide in the same direction, which continues the Psm. All gradate cells are principally similar in shape at least in other fossil genera from North America. Therefore, the assignment of Palaeochrysa greenwalti sp. nov. to Palaeochrysa is undoubted, while P. minor sp. nov. is very probable.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	description	urn: lsid: zoobank. org: act: A 0 BCCD 17 - AA 0 C- 4961 - 89 AD-BBC 32456 AC 1 C Fig. 3	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	materials_examined	Type material. Holotype USNM PAL 622159, deposited in USNM. One nearly complete forewing, one incomplete forewing, and one incomplete hind wing. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. Middle Eocene (Lutetian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	etymology	Etymology. From the surname of Dale E. Greenwalt, in recognition of his work on the Kishenehn Formation.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	diagnosis	Diagnosis. Similar to Palaeochrysa stricta differing from it by smaller size [forewing length ca. 11.2 mm in P. greenwalti sp. nov.; 15.5 mm in P. stricta] and proximal-most crossvein of inner gradate series in hind wing (after RP 5) shifted posteriorly, so that it terminates on Psm [terminates on RP 5 far anterior to Psm in P. stricta]; it differs from P. minor sp. nov. by larger size and wider forewing [length / width ratio ca. 2.95 in P. greenwalti sp. nov.; ca. 3.17 in P. minor sp. nov.].	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	description	Description. Forewing ca. 11.2 mm long, 3.8 mm wide. Costal space moderately broad. Subcostal veinlets simple, relatively closely spaced. Sc long (its termination not discernible). Subcostal space narrow, crossveins not detected. RA terminated at margin slightly proximad wing apex. RA space broad, narrowed distad, with 14 regularly spaced crossveins. RP with 13 pectinate branches. RP 1 to RP 5 terminated at Psm; two distal branches simple; forking of most other branches unclear, at least two in distal portion forked once in one of forewings. Psm straight, only slightly zigzagged. M poorly preserved proximally, forked slightly distad origin of RP. im incompletely preserved (its shape probably similar to that of other species of the genus: narrow, elongate). Presumable MA forked once distad Psc; presumable MP forked at Psc. Crossvein 2 m-cu long, located in proximal part of im. Psc straight, slightly zigzagged only distally. CuA presumably with three branches. CuP forked, with both branches widely spaced. 1 A partially preserved. Two gradate series of crossveins nearly parallel; inner series forming distinct angle with Psm at RP 5, with six preserved crossveins after RP 5 (probably eight in life); outer series continuing Psc, with nine crossveins after RP 5. Hind wing 9.5 mm long as preserved, 2.9 mm wide as preserved. Costal space not preserved. Sc preserved only basally. RA not preserved in distal part. RA with 10 preserved crossveins. RP with 11 preserved pectinate branches (probably 12 in life). RP 1 (presumably) to RP 5 terminating at Psm; two distal branches simple; other branches forked once. Psm straight, almost not zigzagged. Proximal part of M not preserved; presumable MA forked distad Psc; presumable MP forked at Psc. Psc straight, zigzagged only in distal part. Presumable CuA with two preserved branches. Anal veins not preserved. Two gradate series of crossveins: inner series convex, forming distinct angle with Psm at RP 5, with seven preserved crossveins before RP 5 (eight in life); outer series continuing Psc, with six preserved crossveins before RP 5 (eight in life).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA1FFA4FF4CFA29FE611F29.taxon	discussion	Remarks. The apex of the forewing of P. greenwalti sp. nov. appears more rounded than in the other species. No body parts are preserved and some parts of the forewing venation and the basal part of the hind wing are difficult to interpret.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	description	urn: lsid: zoobank. org: act: ECE 45 B 39 - 18 B 0 - 45 CC- 97 C 1 - 07942 C 4 A 2 BC 7 Fig. 4	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	materials_examined	Type material. Holotype USNM PAL 722497, deposited in USNM. An incomplete, crumpled specimen. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Lutetian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	etymology	Etymology. From the Latin parvus, - a, - um (comparative minor), meaning “ small (smaller) ”.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	diagnosis	Diagnosis. Differs from two other species by smaller size, subcostal veinlets widely spaced [closely spaced in other species], and three branches of RP terminating on Psm [five in other species].	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	description	Description. Body very poorly preserved, details not discernible. Forewing ca. 9.5 mm long, ca. 3.0 mm wide. Costal space relative narrow. Subcostal veinlets simple, relatively widely spaced. Distal parts of Sc, RA not preserved. Subcostal space narrow, crossveins not detected. RA space with five detected, widely spaced crossveins. RP with nine pectinate branches. RP 1 to RP 3 terminated at Psm; two or three distal branches simple; RP 1 deeply forked at Psc and its proximal branch shallowly forked in left forewing; other branches rather shallowly onсe forked. Basal crossvein 1 r-m very short, connecting anterior trace of RP and MA within im at proximal one-third length. Psm straight, only slightly zigzagged. M forked slightly distad origin of RP. im narrow, elongate (3.8 times as long as wide). Crossvein 2 m-cu long, connecting im and CuA at nearly proximal one-fifth of im. Psc straight, slightly zigzagged distally. CuA probably with two simple branches. CuP deeply forked. Two intracubital crossveins; c 1: c 2 length ratio 0.82: 1 in right wing. A 1 simple. A 2 strongly arched, simple. Two gradate series of crossveins nearly parallel; inner series with six crossveins after RP 3; outer series continuing Psc, with six crossveins after RP 3. Hind wing 6.8 mm long as preserved, 2.3 mm wide as preserved. Costal space very narrow. Sc preserved proximally. Subcostal space appears to be very narrow. RA long, terminating at margin probably near wing apex. RA space with eight regularly spaced crossveins. RP with nine pectinate branches. RP 1 to RP 3 terminated at Psm; three distal branches simple; forking of other branches mostly unknown. Psm straight, slightly zigzagged. M proximally fused with RP. Presumable MA forked distad Psc; presumable MP forked at Psc. Psc slightly zigzagged. CuA with three simple branches; probably all originating before fusion of CuA with MP. CuP, A 1, A 2 simple. Two nearly parallel gradate series of crossveins: inner series with four preserved crossveins after RP (probably six in life); outer series continuing Psc, with six crossveins after RP 3.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA3FFABFF4CFDACFCDA1991.taxon	discussion	Remarks. Palaeochrysa minor sp. nov. is the smallest member of the genus and has a more elongate forewing than the other species. The small size is reflected in the venation with it having only three branches of RP terminating on Psm [five in the other species of the genus].	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFACFFA9FF4CFAC0FC131CB9.taxon	description	Fig. 5	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFACFFA9FF4CFAC0FC131CB9.taxon	materials_examined	Material. Specimen USNM PAL 624842, deposited in USNM. An incomplete specimen. Locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Luteian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFACFFA9FF4CFAC0FC131CB9.taxon	description	Description. Body poorly preserved, fine details not discernable. Head poorly preserved, only both eyes well discernible (0.5 mm wide). Prothorax 0.6 mm long, mesothorax 1.5 mm long, metathorax 0.8 mm long. Legs and abdomen not preserved. Forewing fragmentary, poorly preserved, ca. 6.8 mm long (as preserved; estimated complete length ca. 8 mm), 2.7 mm wide (as preserved; estimated width ca. 3 mm). Recurrent humeral veinlet well developed, with four preserved branches. Costal space narrowing distally, with ca. 15 partially preserved costal veinlets, mainly forked; trichosors present on costal margin. Sc and RA partially preserved; subcostal crossveins not detected. RP poorly preserved, with five to six branches separately originating on RA. RP 1 at least deeply-forked once near origin. Three gradate series of crossveins partially preserved. Second series consists of two preserved crossveins (right wing), between M and CuP. Third (‘ inner’) series consists of eight preserved crossveins anterior to CuA (probably nine in life) (left wing); fourth (‘ outer’) series with four preserved crossveins (left wing). M forked nearly opposite fork of RP 1; posterior branch probably forked again at inner gradate series. Cubital veins fragmentarily preserved; proximal branch of CuA originating at second gradate series. Anal veins not preserved. Hind wing fragmentarily preserved, 5.5 mm long (as preserved; estimated complete length ca. 7 mm), 1.8 mm wide (as preserved; estimated complete width ca. 2.4 mm). Costal space narrow, narrowest at mid-way point. Costal veinlets partially preserved, proximally simple, distally few forked; trichosors preserved on costal margin. Subcostal space narrow. Sc and RA partially preserved, not fused; reaching wing margin before apex. RA distally with three to four branches. RP partially preserved, at least with five branches. Third (‘ outer’) gradate series with five preserved crossveins. Cubital and anal veins not preserved.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFACFFA9FF4CFAC0FC131CB9.taxon	discussion	Remarks. This specimen obviously belongs to a genus of the Megalomus - group (Drepanepteryginae, Drepanacrinae or Megalominae). These hemerobiids share the wing character states such as a dense venation; a deeply forked CuP in the forewing; and a fully developed CuP in the hind wing (see Makarkin et al. 2016 for detailed discussion). The venation of the specimen is fragmentarily preserved, and it is therefore difficult to place into a particular genus. However, the preserved venation is most similar to that of Proneuronema Makarkin et al. 2016, in particular the presence of at least once deeply-forked RP 1. This genus was widely distributed in the Eocene in North America, Europe, and East Asia (see Makarkin & Perkovsky 2020).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFAFFF4CFB40FD6D1C71.taxon	description	urn: lsid: zoobank. org: act: 67388462 - CDFA- 46 FA- 8803 - B 2978 E 50605 C	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFAFFF4CFB40FD6D1C71.taxon	type_taxon	Type and only species. Macrostigmoraphia diluta sp. nov.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFAFFF4CFB40FD6D1C71.taxon	etymology	Etymology. From the Greek macros, long, stigma, spot, and Raphidia, a genus-group name of Raphidiidae. Gender feminine.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFAFFF4CFB40FD6D1C71.taxon	diagnosis	Diagnosis. May be distinguished from other genera by configuration of pterostigma: it is very long (10 times as long as wide), very pale, and lacks any incorporated veinlets or crossveins.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFAFFF4CFB40FD6D1C71.taxon	discussion	Remarks. The specimen has been placed within Raphidiidae based on the combination of 2 sc-r being far distad the termination of Sc [Inocelliidae: located near termination of Sc], closing the proximal end of pterostigma [Inocelliidae: sc-r not closing pterostigma proximally]. The long pterostigma closed proximally by crossvein 2 sc-r and often without incorporated veinlets is also present in the genera of the Cretaceous tribe Nanoraphidiini (Mesoraphidiidae). In general, their venation is very similar to that of some Raphidiidae, particularly the late Eocene subfamily Succinoraphidiinae Aspöck & Aspöck, 2004 (Perkovsky & Makarkin 2019). Apart from the long pterostigma, the rest of the forewing venation of Macrostigmoraphia gen. nov. is typical of the Eocene Raphidiidae. The venation of all species of Nanoraphidiini (except one wing of Rhynchoraphidia burmana Liu et al., 2016: Fig. 6 B) differs from it, in particular by the simple (not forked) anterior trace of RP distad of 3 ra-rp [two branches are present in Macrostigmoraphia gen. nov.]. It cannot be ruled out, however, that the new genus may theoretically belong to Nanoraphidiini or another group of Mesoraphidiidae, but currently there is insufficient data for this conclusion. The few crossveins and a lack of an enriched venation separate the specimen from Baissopteridae. The slight bend of RA proximad 3 ra-rp might be indicative of the last distal veinlet, closing the pterostigma distally (indiscernible by preservation). In this case, 3 ra-rp would be located distad the pterostigma, but it is very unlikely as this crossvein is located within the pterostigma in all known Eocene Raphidiidae (see Archibald & Makarkin 2021). In Nanoraphidiini, both conditions are present: 3 ra-rp may be located proximad or distad the pterostigma (see Makarkin 2023). If the indiscernible veinlet exists, then RA has two distal veinlets, but all known Nanoraphidiini only have one. Therefore, if this is either Raphidiidae or Nanoraphidiini, the very long pterostigma is characteristic of the genus, not an artefact.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFA8FF4CFF2CFEDC1ED6.taxon	description	Fig. 6	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFA8FF4CFF2CFEDC1ED6.taxon	materials_examined	Material examined. Specimen USNM PAL 621094, deposited in USNM. A fragmentary wing. Locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Luteian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFA8FF4CFF2CFEDC1ED6.taxon	description	Description. Wing ca. 5.8 mm long as preserved (estimated complete length approximately 8 – 10 mm). Costal space relatively narrow. Subcostal veinlets simple, closely spaced. RA space narrow; one crossvein well discernible. Branches of RP closely spaced, dichotomously forked distally. One or two gradate series of crossveins discernible in proximal part of fragment; other crossveins in distal part are possible.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFAFFFA8FF4CFF2CFEDC1ED6.taxon	discussion	Remarks. The family assignment of this wing fragment is unclear. It may theoretically belong to Hemerobiidae, Berothidae or Dilaridae. The narrow costal space and simple closely-spaced costal veinlets in the distal part of the wing show that this is likely a hind wing. In this case, a dilarid affinity is most probable as the hind wings are heavily maculated only in Dilaridae, especially in the American genus Nallachius Navás, 1909, and some East Asian species of Dilar Rambur, 1838 (see Adams 1970: Figs 1, 11, 12; Martins et al. 2018: Fig. 3 A). The arrangement of the preserved crossveins also shows that a dilarid affinity is most probable. However, the mode of terminal branching of the longitudinal veins in Nallachius strongly differs from that of this specimen. The venation of the extant Dilar is more similar to that of the fragment, but still different. Hind wings of Hemerobiidae and Berothidae are hyaline (not maculated), and their crossveins are arranged otherwise. If this is a forewing, a hemerobiid affinity is most probable, but a berothid affinity is also possible (although very unlikely).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA8FFAEFF4CF984FEED1CE1.taxon	description	urn: lsid: zoobank. org: act: 23 A 6 E 686 - 5 B 20 - 4 BD 9 - B 4 CC-F 211 B 33 B 870 A Fig. 7	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA8FFAEFF4CF984FEED1CE1.taxon	materials_examined	Type material. Holotype USNM PAL 620478, deposited in USNM, a distal part of a forewing. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Lutetian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA8FFAEFF4CF984FEED1CE1.taxon	etymology	Etymology. From the Latin dilutus, pale.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA8FFAEFF4CF984FEED1CE1.taxon	description	Description. Forewing 5.7 mm as preserved (estimated complete length 8.5 mm), 2.4 mm wide. Costal space poorly preserved, no veinlets preserved. Sc terminates on C proximad 2 sc-r. Pterostigma very long (10 times as long as wide), pale, no incorporated veinlets detected within. One crossvein preserved between Sc and RA, 2 sc-r, closing pterostigma proximally. RA long terminating before wing apex, with one distal veinlet closing pterostigma distally. Two crossveins between RA and RP, 2 ra-rp, 3 ra-rp. RP origin not preserved, RP with three branches. RP 1 long forking distad irp, RP 2, RP 3 simple. One crossvein preserved between RP 1 and RP 2, irp. Between stem of RP, RP 1 and MA two crossveins preserved, 2 r-m, 3 r-m. M origin not preserved, MA long, forking distally distad 3 r-m. Two crossveins between MA and MP, 1 im, 2 im, forming two doi. MP partially preserved, pectinately branched, first branch proximad 1 im, second between 1 im, 2 im. MP probably fused with CuA proximally. Cu origin not preserved. CuA partially preserved, distal part of its anterior trace and one branch. CuP, anal veins, basal part of wing not preserved.	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA9FFAEFF4CFD59FEA5191D.taxon	description	Fig. 8	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA9FFAEFF4CFD59FEA5191D.taxon	materials_examined	Type material. Holotype USNM PAL 625900, deposited in USNM. A complete hind wing. Type locality and horizon. Middle Fork of the Flathead River, between Paola and Stanton Creeks approximately 17 miles south of West Glacier; the Coal Creek Member of the Kishenehn Formation, north-western Montana, U. S. A. The Middle Eocene (Luteian).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
CB3687DDFFA9FFAEFF4CFD59FEA5191D.taxon	discussion	Remarks. Paraksenocellia includes two Eocene species, P. borealis Makarkin et al., 2019 from the mid-Ypresian Driftwood Canyon Provincial Park, near Smithers, British Columbia, Canada (type species) and P. australis. This is the oldest genus in Inocelliidae, and possesses some rare character states for the family (see Makarkin et al. 2019 for discussion).	en	Jepson, James E., Makarkin, Vladimir N. (2023): Fossil Neuropterida (Insecta: Neuroptera and Raphidioptera) from the middle Eocene Kishenehn Formation, Montana, USA. Zootaxa 5306 (4): 427-444, DOI: 10.11646/zootaxa.5306.4.2, URL: https://doi.org/10.11646/zootaxa.5306.4.2
